List of bacterial orders

This article lists the orders of the Bacteria. The currently accepted taxonomy is based on the List of Prokaryotic names with Standing in Nomenclature (LPSN)^[1] and National Center for Biotechnology Information (NCBI)^[2] and the phylogeny is based on 16S rRNA-based LTP release 132 by The All-Species Living Tree Project.^[3]

Phylogeny

National Center for Biotechnology Information (NCBI) taxonomy was initially used to decorate the genome tree via tax2tree. The 16S rRNA-based Greengenes taxonomy is used to supplement the taxonomy particularly in regions of the tree with no cultured representatives. List of Prokaryotic names with Standing in Nomenclature (LPSN) is used as the primary taxonomic authority for establishing naming priorities. Taxonomic ranks are normalised using phylorank and the taxonomy manually curated to remove polyphyletic groups.^{[4][5][6][7][8]}

Cladogram was taken from GTDB release 09-RS220 (24th April 2024).^[9][10][11]

Bacteria Chloroflexae Chloroflexota Bacillati "Sysuimicrobiota" Armatimonadota Eremiobacterota Bacillota G Bacillota E Bacillota s.s. "Desulfotomaculota" (Bacillota B) "Selenobacteria" (Bacillota C) Bacillota D "Halanaerobiaeota" (Bacillota F) Bacillota A "Actinobacteriobiontes" Actinomycetota "Bipolaricaulota" Deinococcota Thermotogati "Synergistetes" Atribacterota Synergistota "Thermocalda" Dictyoglomota "Thermodesulfobiota" Coprothermobacterota "Lithacetigenota" Caldisericota Thermotogota Cyanoprokaryota "Margulisiibacteriota" Cyanobacteria "CPR" "Patescibacteria" "Pseudomonadati" "Lindowiibacteriota" "Walliibacteriota" "Muiribacteriota" "Rifleibacteriota" "Babelota" Fusobacteriota "Macinerneyibacteriota" Spirochaetota "Poribacteriota" "Goldiibacteriota" "Firestoneibacteriota" "Desantisiibacteriota" "Aerophobota" Elusimicrobiota "Ratteibacteriota" "Omnitrophota" UBA1439 PUNC01 JAKLEM01 "Auribacterota" JACPWU01 Chlamydiota Verrucomicrobiota {"FCB group"} "Hinthialibacterota" [OLB16] "Sumerlaeota" "Abyssubacteria" "Hydrogenedentota" "Coatesiibacteriota" [RBG-13-66-14] Planctomycetota {"Proteobacteria"}

"Terrabacteria"

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Kingdom Bacillati

Phylum Chloroflexota

Phylogeny of Chloroflexota[9][10][11]

"Dormibacteria" "Dormibacterales" "Limnocylindria" "Limnocylindrales" Ktedonobacteria Ktedonobacterales "Dehalococcoidotia" Tepidiformales "Australimonadales" "Lucifugimonadales" Dehalococcoidales Dehalococcoidia "Caldilineotia" Ardenticatenales "Epilineales" "Caldilineidae" Caldilineales "Anaerolineidae" J036 Thermoflexales "Promineofilales" Aggregatilineales Anaerolineales "Caldilineia" "Chloroflexotia" "Thermobaculales" Thermomicrobiales Chloroflexales Chloroflexia

• Class "Bathosphaeria" Mehrshad et al. 2018
  •  ?"Bathosphaerales" Mehrshad et al. 2018 [JG30-KF-CM66]
• Class "Martimicrobia" Williams et al. 2024
  •  ?"Martimicrobiales" Williams et al. 2024
• Class ?"Poriflexia" Kogawa et al. 2022
• Class "Spiritibacteria" Williams et al. 2024
  •  ?"Spiritibacterales" Williams et al. 2024
• Class "Tarhunnaeia" Williams et al. 2024
  •  ?"Tarhunnaeales" Williams et al. 2024
• Class "Uliximicrobia" Williams et al. 2024
  •  ?"Uliximicrobiales" Williams et al. 2024
• Class "Umbricyclopia" Mehrshad et al. 2018
  •  ?"Umbricyclopales" Mehrshad et al. 2018 (TK10)
• Class "Dormiibacteria" (sic) Montgomery et al. 2021 (AD3)
  •  ?"Aeolococcales" Montgomery et al. 2021
  • "Dormibacterales" (sic) Montgomery et al. 2021
• Class "Limnocylindria" Mehrshad et al. 2018
  • "Limnocylindrales" Mehrshad et al. 2018 (SL56)
• Class Ktedobacteria corrig. Cavaletti et al. 2007
  • Ktedobacterales corrig. Cavaletti et al. 2007 (Thermogemmatisporales)
• Class Dehalococcoidia Löffler et al. 2013
  • "Australimonadales" Prabhu et al. 2024 [UBA2963]
  • Dehalococcoidales Löffler et al. 2013
  • "Lucifugimonadales" Lim et al. 2023
  • "Monstramariales" Landry et al. 2017 [UBA11650]
  • Tepidiformales Kochetkova et al. 2020
• Class "Caldilineia" Oren, Parte & Garrity 2016 ex Cavalier-Smith 2020
  • Aggregatilineales Nakahara et al. 2019 ("Phototrophicales")
  • Anaerolineales Yamada et al. 2006
  • Ardenticatenales Kawaichi et al. 2013
  • Caldilineales Yamada et al. 2006
  • "Epilineales" Petriglieri et al. 2023
  • "Roseilineaceae" Ward et al. 2021 {J036}
  • "Promineifilales" corrig. Chuvochina et al. 2023
  • Thermoflexales Dodsworth et al. 2014
• Class Chloroflexia Gupta et al. 2013
  • Chloroflexales Gupta et al. 2013 (Herpetosiphonales, Kallotenuales)
  • "Thermobaculales" Chuvochina et al. 2023
  • Thermomicrobiales Garrity & Holt 2002 (Sphaerobacterales)

Phylum "Sysuimicrobiota"

• Class "Sysuimicrobiia" Liu et al. 2024 [CSP1-3]
  • "Sysuimicrobiales" Liu et al. 2024

Phylum Armatimonadota

Phylogeny of Armatimonadota[9][10][11]

"Heboniibacteriia" "Heboniibacterales" "Zipacnadia" "Zipacnadales" Abditibacteriia Abditibacteriales HRBIN17 HRBIN17 UBA5829 UBA5829 Fimbriimonadia Fimbriimonadales Chthonomonadetes Chthonomonadales Armatimonadia Armatimonadales

• Class "Heboniibacteriia" corrig. Carlton et al. 2023 (CAIYQO01)
  • "Heboniibacterales" corrig. Carlton et al. 2023
• Class "Zipacnadia" Carlton et al. 2023 (UBA5377)
  • "Zipacnadales" Carlton et al. 2023
• Class Abditibacteriia Tahon et al. 2018 (FBP)
  • Abditibacteriales Tahon et al. 2018
• Class HRBIN17
  • "Candidatus Fervidibacter" Rinke et al. 2013 {HRBIN17: HRBIN17} (OctSpa1-106)
• Class UBA5829
  • "Candidatus Hippobium" Gilroy et al. 2022 {UBA5829: UBA5829}
• Class Fimbriimonadia Im et al. 2012
  • Fimbriimonadales Im et al. 2012
• Class Chthonomonadia Oren, Parte & Garrity 2016
  • Chthonomonadales Lee et al. 2011
• Class Armatimonadia Tamaki et al. 2011 (OP10)
  • Armatimonadales Tamaki et al. 2011 (Capsulimonadales)

Phylum Vulcanimicrobiota

Phylogeny of Vulcanimicrobiota[9][10][11]

"Xenobia" "Xenobiales" Vulcanimicrobiia "Eremiobacterales" Vulcanimicrobiales

• Class "Eudoremicrobiia" Paoli et al. 2022
  •  ?"Eudoremicrobiales" Paoli et al. 2022
• Class "Xenobia" Ji et al. 2021 (UBP9)
  • "Xenobiales" Ji et al. 2021
• Class "Eremiobacteriia" corrig. Ward, Cardona & Holland-Moritz 2019 (WPS2)
  • "Eremiobacterales" Ward, Cardona & Holland-Moritz 2019
  • "Vulcanimicrobiales Yabe et al. 2024 ("Baltobacterales"; "Palusbacterales"; "Rubrimentiphilales")

Clade Firmicutes

Firmicutes incertae sedis

• Class Desulfitisporia Sorokin & Merkel 2023
  • Desulfitisporales Sorokin & Merkel 2023
• Class Fermentithermobacillia Kadnikov et al. 2023
  • Fermentithermobacillales Kadnikov et al. 2023
• Class Thermolithobacteria Sokolova et al. 2007
  • Thermolithobacterales Sokolova et al. 2007

Phylum Bacillota G

• Class Limnochordia Watanabe, Kojima & Fukui 2015
  • Limnochordales Watanabe, Kojima & Fukui 2015
• Class "Hydrogenisporia" Pallen, Rodriguez-R & Alikhan 2022
  • "Capillibacteriales" Pallen, Rodriguez-R & Alikhan 2022 [UBA10575]
  • "Hydrogenisporales" Pallen, Rodriguez-R & Alikhan 2022 [UBA8346]

Phylum Bacillota E

Phylogeny of Bacillota E[9][10][11]

UBA3575 UBA3575 Thermaerobacteria Thermaerobacterales "Symbiobacteriia Symbiobacteriales Sulfobacillia Sulfobacillales

• Class UBA3575
  • "Candidatus Acetocimmeria" Smith et al. 2021 {UBA3575: UBA3575}
• Class Thermaerobacteria Chuvochina et al. 2024
  • Thermaerobacterales Chuvochina et al. 2024
• Class Symbiobacteriia Chuvochina et al. 2024
  • Symbiobacteriales Chuvochina et al. 2024
• Class Sulfobacillia Chuvochina et al. 2024
  • Sulfobacillales Chuvochina et al. 2024

Phylum "Selenobacteria"

Phylogeny of Selenobacteria[9][10][11]

Veillonellales Acidaminococcales "Sporomusales" A UPPP01 "Sporomusales" C "Sporomusales" UMGS1260 DSM-13327 Propionisporales Anaeromusales "Dendrosporobacterales" Selenomonadales

• Class Selenomonadia Oren, Parte & Garrity 2016
  • Acidaminococcales Campbell, Adeolu & Gupta 2015
  • Anaeromusales Chuvochina et al. 2024
  • "Dendrosporobacterales" Pallen, Rodriguez-R & Alikhan 2022
  • "Pelosinaceae" {DSM-13327}
  • Propionisporales Chuvochina et al. 2024
  • Selenomonadales Marchandin et al. 2010
  • "Acetonemaceae" {Sporomusales_A}
  • "Sporomusales"_C
  • "Sporomusales" Cavalier-Smith 2006
  • "Pelorhabdus" {UMGS1260: UMGS1260}
  • Lucifera Sanchez-Andrea et al. 2018 {UPPP01: UPPP01}
  • Veillonellales Campbell, Adeolu & Gupta 2015

Phylum "Desulfotomaculota"

Phylogeny of Desulfotomaculota[9][10][11]

Peptococcia "Thermanaerosceptrales" Peptococcales "Dehalobacteriia" "Dehalobacteriales" "Avidehalobacterales" "Cryptoclostridiales" "Desulfitobacteriia" "Heliobacteriales" Desulfitobacteriales "Moorellia" DULZ01 Desulfitibacterales Calderihabitantales Moorellales DSM‑12270 Thermacetogeniales Syntrophomonadia Syntrophomonadales "Carboxydocellia" Carboxydocellales Thermincolia Thermincolales "Carboxydothermia" Carboxydothermales Desulfotomaculia Ammonificales Desulfotomaculales

• Class Syntrophomonadia Chuvochina et al. 2024
  • Syntrophomonadales Chuvochina et al. 2024
• Class Peptococcia Chuvochina et al. 2024
  • Peptococcales Chuvochina et al. 2024
  • "Thermanaerosceptrales" Pallen, Rodriguez-R & Alikhan 2022 [DRI-13]
• Class "Dehalobacteriia"
  • "Avidehalobacterales" Pallen, Rodriguez-R & Alikhan 2022 [UBA4068]
  • "Cryptoclostridiales" Pallen, Rodriguez-R & Alikhan 2022 [UBA7702]
  • "Dehalobacteriales"
• Class Desulfitobacteriia Chuvochina et al. 2024
  • Desulfitobacteriales Chuvochina et al. 2024
  • "Heliobacteriales" Kinman 1994 ex Cavalier-Smith 2020
• Class DSM-12270
  • Thermacetogeniales Chuvochina et al. 2024
• Class "Moorellia"
  • Calderihabitantales Chuvochina et al. 2024
  • Desulfitibacterales Chuvochina et al. 2024
  • Zhaonellaceae Lv et al. 2020 {DULZ01}
  • Moorellales Lv et al. 2020
• Class "Carboxydocellia" Pallen, Rodriguez-R & Alikhan 2022
  • "[Carboxydocellales]] Chuvochina et al. 2024
• Class Thermincolia Chuvochina et al. 2024
  • "[Thermincolales]] Chuvochina et al. 2024
• Class "Carboxydothermia" Pallen, Rodriguez-R & Alikhan 2022
  • Carboxydothermales Chuvochina et al. 2024
• Class Desulfotomaculia Chuvochina et al. 2024
  • Ammonificales corrig. Chuvochina et al. 2024
  • Desulfotomaculales Chuvochina et al. 2024

Phylum Bacillota D

Phylogeny of Bacillota D[9][10][11]

"Proteinivoracia" "Proteinivoracales" Natranaerobiia Natranaerobiales "Dethiobacteria" SKNC01 "Dethiobacterales"

• Class "Proteinivoracia"
  • "Proteinivoracales"
• Class "Dethiobacteria" Sorokin & Merkel 2022
  • "Candidatus Contubernalis" corrig. Zhilina et al. 2005 {SKNC01: SKNC01}
  • "Dethiobacterales" Sorokin & Merkel 2022
• Class Natranaerobiia Chuvochina et al. 2024
  • Natranaerobiales Mesbah et al. 2007

Phylum "Halanaerobiaeota"

Phylogeny of Halanaerobiia[9][10][11]

"Anoxybacterales" Halanaerobiales Halobacteroidales

• Class "Halanaerobiia" Cavalier-Smith 2020
  • "Anoxybacterales" Pallen, Rodriguez-R & Alikhan 2022 {DY22613}
  • Halanaerobiales corrig. Rainey & Zhilina 1995
  • Halobacteroidales Chuvochina et al. 2024

Phylum Bacillota A

Phylogeny of Bacillota A[9][10][11]

Thermosediminibacteria Koleobacterales Thermosediminibacterales "Thermoanaerobacteria" Caldicellulosiruptorales Thermoanaerobacterales Clostridiia Mahellales Caldicoprobacterales Lutisporales Clostridiales Eubacteriales Tissierellales Peptostreptococcales Christensenellales "Merdicolales" "Egerieisomatales" "Saccharofermentanales" "Acetivibrionales A Acetivibrionales "Petroclostridiales" Monoglobales "Avimonoglobales" "Oscillospirales" Lachnospirales

• Class Thermosediminibacteria Chuvochina et al. 2024
  • Koleobacterales Sakamoto et al. 2021
  • Thermosediminibacterales Zhang et al. 2019 ("Thermovenabulales")
• Class "Thermoanaerobacteria"
  • Caldicellulosiruptorales Chuvochina et al. 2024
  • Thermoanaerobacterales Wiegel 2010 ["Caldanaerobiales"]
• Class "Clostridiia" Oren, Parte & Garrity 2016
  • "Thermoclostridiaceae" Pallen, Rodriguez-R & Alikhan 2022 {"Acetivibrionales" A}
  • Acetivibrionales Chuvochina et al. 2024
  • "Avimonoglobales" Pallen, Rodriguez-R & Alikhan 2022 [UBA1381]
  • Caldicoprobacterales Chuvochina et al. 2024
  • Christensenellales Parks et al. 2018 ex Chuvochina et al. 2024 ("Borkfalkiales"; "Comantemales")
  • Clostridiales Prevot 1953 s.s. ("Plectridiales")
  •  ?"Darwinibacteriales" Puchol-Royo et al. 2023 [MBA03]
  • "Egerieisomatales" Pallen, Rodriguez-R & Alikhan 2022 [UBA1212]
  • Eubacteriales Buchanan 1917
  • Lachnospirales Chuvochina et al. 2024
  • Lutisporales Chuvochina et al. 2024
  • Mahellales Chuvochina et al. 2024
  • "Merdicolales" Pallen, Rodriguez-R & Alikhan 2022 [TANB77]
  • Monoglobales Chuvochina et al. 2024
  • "Oscillospirales" Glendinning et al. 2020
  • "Petroclostridiales" Pallen, Rodriguez-R & Alikhan 2022 [SK-Y3]
  • Peptostreptococcales Chuvochina et al. 2024
  • "Saccharofermentanales" corrig. Parks et al. 2018
  • Tissierellales Alauzet et al. 2020

Phylum "Bacillota"

Phylogeny of Bacillota[9][10][11]

"Bacillia" "Pasteuriales" Kyrpidiales Tumebacillales Alicyclobacillales Desulfuribacillales Calditerricolales Bacillales F Thermicanales "Brockiales" Thermoactinomycetales DSM-22679 Tepidibacillales Paenibacillales Aneurinibacillales Brevibacillales "Rubeoparvulales" Caldalkalibacillales Bacillales_K "Bacillidae" Bacillales E Bacillales G Bacillales H Bacillales I Bacillales D Bacillales "Harrysmithimonadales" Exiguobacteriales Staphylococcales "Erysipelotrichidae" Culicoidibacterales Haloplasmatales "Bathyoplasmales" "Izemoplasmatales" Acholeplasmatales "Aphodocolales" RFN20 Erysipelotrichales Mycoplasmatales Lactobacillales

• Class "Bacillia" Cavalier-Smith 2020
  • Alicyclobacillales Chuvochina et al. 2024
  • Aneurinibacillales Chuvochina et al. 2024
  • "Bacillidae" Cavalier-Smith 2020 non Enderlein 1917
    • Bacillales D
    • Bacillales E
    • Bacillales G
    • Bacillales H
    • Bacillales I
    • Bacillales Prevot 1953 s.s. (Caryophanales, "Caldibacillales")
  • "Bacillus thermozeamaize" Mak 2003 {Bacillales F: Bacillaceae_M}
  • Sporolactobacillaceae Ludwig et al. 2010 {Bacillales_K}
  • Brevibacillales Chuvochina et al. 2024
  • "Brockiales" Pallen, Rodriguez-R & Alikhan 2022
  • Caldalkalibacillales Chuvochina et al. 2024
  • Calditerricolales Chuvochina et al. 2024
  • Microaerobacter Khelifi et al. 2011 {DSM-22679: DSM-22679}
  • Desulfuribacillales Sorokin et al. 2021
  • "Erysipelotrichidae" corrig. Cavalier-Smith 2020
    • Acholeplasmatales Freundt et al. 1984 (Anaeroplasmatales)
    • "Aphodocolales" Pallen, Rodriguez-R & Alikhan 2022
    • "Bathyoplasmales" corrig. Zhu, Lian & He 2020 [MAG-NZ]
    • Culicoidibacterales Neupane et al. 2020
    • Erysipelotrichales Ludwig et al. 2010
    • Haloplasmatales Rainey et al. 2008 non Zhou et al. 2022
    • "Izemoplasmatales" corrig. Zheng et al. 2021
    • Mycoplasmatales Freundt 1955 ("Borrelomycetales"; "Anulomycetales"; "Paramycetales"; "Mollicutales"; "Pleuropneumoniales"; Entomoplasmatales, Mycoplasmoidales)
    • RFN20
  • Exiguobacteriales Chuvochina et al. 2024
  • "Harrysmithimonadales" Pallen, Rodriguez-R & Alikhan 2022 [CAJFEE01]
  • Kyrpidiales Chuvochina et al. 2024
  • Lactobacillales Ludwig, Schleifer & Whitman 2010 ("Plocamobacteriales"; "Coccales")
  • Paenibacillales Chuvochina et al. 2024
  • "Pasteuriales" Pallen, Rodriguez-R & Alikhan 2022 [RES148]
  • "Rubeoparvulales"
  • Staphylococcales Chuvochina et al. 2024
  • Tepidibacillales Chuvochina et al. 2024
  • Thermicanales Chuvochina et al. 2024
  • Thermoactinomycetales Chuvochina et al. 2024
  • Tumebacillales Chuvochina et al. 2024

Phylum Actinomycetota

Phylogeny of Actinomycetota[9][10][11]

"Humimicrobiia" "Hakubellales" "Humimicrobiales" "Aquicultoria" "Aquicultorales" "Geothermocultorales" "Oleimmundimicrobiales" "Subteraquimicrobiales" Coriobacteriia Coriobacteriales "Geothermincolia" "Geohydrothermomicrobiales" "Geothermincolales" Rubrobacteria Rubrobacterales Thermoleophilia Gaiellales Miltoncostaeales Thermoleophilales Acidimicrobiia "Actinomarinales" Acidimicrobiales Actinomycetia Nitriliruptoridae Euzebyales Nitriliruptorales Actinobacteridae Actinomycetales "Nanopelagicales" Jiangellales Propionibacteriales Motilibacterales Streptosporangiales Acidothermales Mycobacteriales Sporichthyales Streptomycetales

• Class ?"Syntrophaliphaticia" corrig. Liu et al. 2020
• Class "Humimicrobiia" Jiao et al. 2021
  • "Hakubellales" corrig. Merino et al. 2020
  • "Humimicrobiales" Jiao et al. 2021
• Class "Aquicultoria" Jiao et al. 2021
  • "Aquicultorales" Jiao et al. 2021
  • "Geothermocultorales" Jiao et al. 2021
  • "Oleimmundimicrobiales" Pallen, Rodriguez-R & Alikhan 2022 [UBA3085]
  • "Subteraquimicrobiales" Jiao et al. 2021
• Class Coriobacteriia König 2013
  •  ?"Anaerosomatales" Khomyakova et al. 2022
  • Coriobacteriales Stackebrandt et al. 1997 (Eggerthellales)
• Class "Geothermincolia" Jiao et al. 2021
  • "Geohydrothermomicrobiales" Pallen, Rodriguez-R & Alikhan 2022 [Fen-727]
  • "Geothermincolales" Jiao et al. 2021
• Class Rubrobacteria Suzuki 2013
  • Rubrobacterales Rainey et al. 1997
• Class Thermoleophilia Suzuki & Whitman 2013
  • Gaiellales Albuquerque et al. 2012
  • "Miltoncostaeales" Li et al. 2021
  • Thermoleophilales Reddy & Garcia-Pichel 2009 (Solirubrobacterales)
• Class Acidimicrobiia Norris 2013
  •  ?"Spongiisociales" corrig. Nguyen 2022
  • Acidimicrobiales Stackebrandt, Rainey & Ward-Rainey 1997
  • "Actinomarinales" Ghai et al. 2013
• Class "Actinomycetia" Salam et al. 2020
  • Acidothermales Sen et al. 2014
  • Actinomycetales Buchanan 1917 (Aquipuribacterales; Beutenbergiales; Bifidobacteriales; Bogoriellales; Brevibacteriales; Cellulomonadales; Demequinales; Dermabacterales; Dermatophilales; Microbacteriales; Micrococcales; Kineosporiales; Ruaniales)
  • Euzebyales Kurahashi et al. 2010 (Egibacterales)
  • Jiangellales Tang et al. 2015
  • Motilibacterales Salam et al. 2020
  • Mycobacteriales Janke 1924 (Corynebacteriales; Actinocatenisporales; Actinoplanetales; Actinopolysporales; Antricoccales; Cryptosporangiales; Frankiales; Geodermatophilales; Glycomycetales; Jatrophihabitantales; Micromonosporales; Nakamurellales; Phytomonosporales; Pseudonocardiales)
  • "Nanopelagicales" Neuenschwander et al. 2018
  • Nitriliruptorales Sorokin et al. 2009 (Egicoccales, Salsipaludibacterales)
  • Propionibacteriales Patrick & McDowell 2015
  • Sporichthyales Nouioui et al. 2018
  • Streptomycetales Kampfer 2015 (Catenulisporales, Kitasatosporales)
  • Streptosporangiales Goodfellow 2015

Clade Cyanoprokaryota

Phylum "Margulisiibacteriota"

Phylogeny of Margulisiibacteriota[9][10][11]

"Saganbacteria" "Riflemargulisbacteria" "Marinamargulisbacteria" "Termititenacia"

• Class "Saganbacteria" Carnevali et al. 2018 (WOR1)
• Class "Marinamargulisbacteria" Carnevali et al. 2019
• Class "Riflemargulisbacteria" Carnevali et al. 2019 (GWF2_35_9)
• Class "Termititenacia" Utami et al. 2019
  • "Termititenacales" Utami et al. 2019

Phylum "Cyanobacteriota"

Phylogeny of Cyanobacteriota[9][10][11]

"Sericytochromatia" UBA7694 S15B-MN24 "Melainabacteria" "Caenarcanales" "Obscuribacterales" "Vampirovibrionales" "Gastranaerophilales" Cyanobacteria "Gloeobacteria" Gloeobacterales "Phycobacteria" Thermostichales Prochlorotrichales Gloeoemargaritales Pseudanabenales "Synechococcophycidae" "Limnotrichales" Prochlorotrichales Synechococcales "Acaryochloridales" Neosynechococcales B231 Leptolyngbyales Nodosilineales Oculatellales "Nostocophycidae" Cyanobacteriales

• Class "Sericytochromatia" Soo et al. 2017
  • S15B-MN24 ["Sericytochromatia"; "Tanganyikabacteria"]
  • UBA7694 ["Blackallbacteria"]
• Class "Melainabacteria" Soo et al. 2014
  • "Caenarcaniphilales" Soo et al. 2014
  • "Gastranaerophilales" Di Rienzi et al. 2013
  • "Obscuribacterales" Soo et al. 2014
  • "Vampirovibrionales" Soo et al. 2015 ex Chuvochina et al. 2023
• Class Gloeobacteria Cavalier-Smith 2002
  • Gloeobacterales Cavalier-Smith 2002
• Class Phycobacteria Cavalier-Smith 2002
  • Acaryochloridales Miyashita et al. 2003 ex Strunecký & Mareš 2022 ("Thermosynechococcales")
  •  ?Aegeococcales Strunecký & Mareš 2022
  • Trichothermofontia Daroch et al. 2023 {B231}
  • Cyanobacteriales Rippka & Cohen-Bazire 1983 (Chamaesiphonales, Chroococcales, Chroococcidiopsidales, Coleofasciculales, Desertifilales, Nostocales, Oscillatoriales, Pleurocapsales, Spirulinales, Stigonematales)
  •  ?"Eurycoccales"
  •  ?Geitlerinematales Strunecký & Mareš 2022
  • Gloeoemargaritales Moreira et al. 2016
  • Leptolyngbyales Strunecký & Mareš 2022
  • "Limnotrichales"
  • Neosynechococcales Chuvochina et al. 2024
  • Nodosilineales Strunecký & Mareš 2022 (Phormidesmidales)
  • Oculatellales Strunecký & Mareš 2022 (Elainellales)
  • Prochlorococcaceae Komárek & Strunecky 2020 {"PCC-6307"}
  • Prochlorotrichales Strunecký & Mareš 2022 [PCC-9006]
  • Pseudanabenales Hoffmann, Komárek & Kastovsky 2005
  • Synechococcales Hoffmann, Komárek & Kastovsky 2005
  • Thermostichales Komárek & Strunecký 2020

Kingdom Thermotogati

Phylum Atribacterota

• Class Atribacteria Katayama et al. 2021 (OP9)
  • Atribacterales Katayama et al. 2021 ("Caldatribacteriales")
• Class "Phoenicimicrobiia" Jiao et al. 2024 (JS1)
  • "Phoenicimicrobiales" Jiao et al. 2024 (SB-45)

Phylum Synergistota

• Class Synergistia Jumas-Bilak et al. 2009
  • Synergistales Jumas-Bilak et al. 2009

Phylum "Zhurongbacterota"

Phylum Dictyoglomota

• Class Dictyoglomeria corrig. Patel 2012
  • Dictyoglomerales corrig. Patel 2012

Phylum "Thermodesulfobiota"

• Class "Thermodesulfobiia" Frolov et al. 2022
  • "Thermodesulfobiales" Cavalier-Smith 2020

Phylum Coprothermobacterota

• Class Coprothermobacteria Pavan et al. 2018
  • Coprothermobacterales Pavan et al. 2018

Phylum "Lithacetigenota"

• Class "HKB111"
  • "HKB111"

Phylum "Caldisericota"

• Class Caldisericia Mori et al. 2009 (OP5)
  • Caldisericales Mori et al. 2009
  • "Cryosericales" Martinez et al. 2019

Phylum Thermotogota

Phylogeny of Thermotogota[9][10][11]

Mesoaciditogales Petrotogales Thermotogales

• Class "Thermotogia" Oren, Parte & Garrity 2016 ex Cavalier-Smith 2020
  • Mesoaciditogales Itoh et al. 2016
  • Petrotogales Bhandari & Gupta 2014 (Kosmotogales)
  • Thermotogales Reysenbach 2002 emend. Bhandari & Gupta 2014

Clade Candidate Phyla Radiation

Phylum "Elulota"

• Class "Elulimicrobiia" Rodriguez-R et al. 2020
  •  ?"Elulimicrobiales" Rodriguez-R et al. 2020

Phylogeny of Patescibacteria[9][10][11]

"Wirthbacteria" (CG2-30-54-11) "Microgenomates" "Dojkabacteria" "Katanobacteria" "Microgenomatia" ⊞ ⊟ "Woykebacterales" "Curtissbacterales" "Daviesbacterales" "Roizmanbacterales" (UBA1406) "Gottesmanbacterales" (UBA10105) "Levybacterales" GWA2-44-7 "Chazhemtobacteriales" "Shapirobacterales" (UBA12405) cluster "Saccharibacteria" "Berkelbacteria" "Kazanbacteria" (Kazan) "Howlettbacteria" "Saccharimonadia" cluster "Gracilibacteria" "Absconditabacteria" BD1-5 "Absconditabacterales" "Gracilibacteria" ⊞ ⊟ "Abawacabacterales" (RBG-16-42-10) "Peregrinibacterales" (UBA1369) "Fallacibacteriales" (DOLZORAL124_38_8; UBA4473) "Peribacterales" cluster "Parcubacteria" "Doudnabacteria" "Andersenbacteria" "Torokbacteria" (GCA-2792135) ABY1 ⊞ ⊟ "Kerfeldbacterales" (SBBC01) "Jacksonbacterales" (UBA9629) "Komeilibacterales" (UBA1558) "Kuenenbacterales" (UBA2196) "Veblenbacterales" "Uhrbacterales" (GWA2-46-9) "Magasanikbacterales" "Buchananbacterales" "Falkowbacterales" "Moisslbacterales" (UBA2591) "Paceibacteria" ⊞ ⊟ "Moranbacterales" UBA6257 "Spechtbacterales" "Terrybacterales" "Parcunitrobacterales" (GWA2-38-13b) "Portnoybacterales" "Paceibacterales" "Azambacterales" (UBA10092) "Yanofskybacterales" (2-02-FULL-40-12) "Sungbacterales" "Ryanbacterales" "Giovannonibacterales" "Niyogibacterales" "Tagabacterales" UBA9973 cluster

Phylum "Patescibacteria"

• Class "Wirthbacteria"
• Class "Dojkabacteria" (WS6)
• Class "Katanobacteria" (WWE3)
• Class "Microgenomatia"
  • Clade GWA2-44-7
  • "Chazhemtobacteriales" Pallen, Rodriguez-R & Alikhan 2022 [UBA1400]
  • "Curtissbacterales"
  • "Daviesbacterales"
  • "Gottesmanbacterales"
  • "Levybacterales"
  • "Roizmanbacterales"
  • "Shapirobacterales"
  • "Woykebacterales" (RIF34)
• Class "Absconditabacteria"
  • "Absconditabacterales" (sic)Yakimov et al. 2022 (SR1)
  • "Candidatus Altimarinus" Rinke et al. 2013 {BD1-5: UBA6164} (GN02)
• Class "Gracilibacteria"
  • "Abawacabacterales" (RIF46)
  • "Fertabacterales" (DOLZORAL124_38_8; "Fallacibacteriota")
  • "Peregrinibacterales" (PER)
  • "Peribacterales" Anantharaman et al. 2016
• Class "Berkelbacteria" (ACD58)
• Class "Kazanbacteria" (Kazan)
• Class "Howlettbacteria" (CPR2)
• Class "Saccharimonadia" corrig. Lemos et al. 2019 (TM7)
  • "Saccharimonadales" corrig. Lemos et al. 2019 ex Chuvochina et al. 2023
• Class "Andersenbacteria" (RIF9)
• Class "Doudnabacteria" (SM2F11)
• Class "Torokbacteria" (GCA-2792135)
• Class "ABY1"
  •  ?"Brownbacterales"
  • "Buchananbacterales" (RIF37)
  • "Falkowbacterales"
  • "Jacksonbacterales" (RIF38)
  • "Kerfeldbacterales" (RIF4)
  • "Komeilibacterales" (RIF6)
  • "Kuenenbacterales"
  • "Magasanikbacterales"
  • "Moisslbacterales" [UBA2591]
  • "Uhrbacterales"
  • "Veblenbacterales" (RIF39)
• Class "Paceibacteria" Chuvochina et al. 2023
  • "Azambacterales"
  • Clade UBA6257
  • "Giovannonibacterales" (UBA11713)
  •  ?"Montesolbacterales"
  • "Moranbacterales" (OD1-i)
  • "Niyogibacterales" (RIF11; HO2-45-28)
  • "Paceibacterales" Chuvochina et al. 2023
  • "Parcunitrobacterales" Castelle et al. 2017
  • "Portnoybacterales" (RIF22)
  • "Ryanbacterales" (RIF10)
  • "Spechtbacterales" (RIF19)
  • "Sungbacterales" (RIF17)
  • "Tagabacterales" (RIF12)
  • "Terrybacterales" (RIF13)
  • "Yanofskybacterales" (2-02-FULL-40-12)

Clade

Clade "Fusobacterida"

Phylum "Muiribacteriota"

• Class "Muiribacteriia" corrig. Chuvochina et al. 2023
  • "Muiribacteriales" corrig. Chuvochina et al. 2023

Phylum "Rifleibacteriota"

• Class "Ozemibacteria" corrig. Kadnikov et al. 2018 (RIF32/ACD39)
  • "Ozemibacterales" corrig. Kadnikov et al. 2018

Phylum "Macinerneyibacteriota"

• Class "Macinerneyibacteriia" corrig. Yadav et al. 2020
  • "Macinerneyibacteriales" corrig. Yadav et al. 2020

Phylum Fusobacteriota

• Class Fusobacteriia corrig. Staley & Whitman 2012
  • Fusobacterales Staley & Whitman 2012 ("Bactepneumoniales")

Clade

Phylum "Bipolaricaulota"

• Class "Bipolaricaulia" Chuvochina et al. 2023
  • "Bipolaricaulales" Chuvochina et al. 2023 (KB1 group, OP1)
  • "Fraserbacterales" [GTDB tag: RIF31; RBG-16-55-9]

Phylum Deinococcota

• "Deinococcia" Cavalier-Smith 2020
  • Deinococcales Rainey et al. 1997 (Thermales; Trueperales) (radio-resistant micrococci)

Kingdom Pseudomonadati

Clade

Phylum "Goldiibacteriota"

• "Candidatus Cellulosimonas" Doud et al. 2020

Phylum Elusimicrobiota

• Class "Elusimicrobiia" Oren, Parte & Garrity 2016 ex Cavalier-Smith 2020 (TG1)
  • Elusimicrobiales Geissinger et al. 2010
  • "Obscuribacteriales" Uzun et al. 2023
• Class "Endomicrobiia" Stingl et al. 2018
  • "Endomicrobiales" Zheng et al. 2018

Phylum "Aerophobota"

• Class "Aerophobia" Chuvochina et al. 2023 (CD12)
  • "Aerophobales" Chuvochina et al. 2023

Phylum "Auribacterota"

Phylogeny of "Auribacterota"[9][10][11]

"Tritonobacteria" "Tritonobacterales" "Erginobacteria" "Erginobacterales" "Auribacteria" "Auribacterales" "Ancaeobacteria" "Ancaeobacterales" "Theseobacterales"

• Class "Ancaeobacteria" Williams et al. 2022 [UBA6266]
  • "Ancaeobacterales" Williams et al. 2022
  • "Theseobacterales" Williams et al. 2022
• Class "Auribacteria" Williams et al. 2022 [SURF-26]
  • "Auribacterales" Williams et al. 2022
• Class "Erginobacteria" Williams et al. 2022 [PUNC01]
  • "Erginobacterales" Williams et al. 2022
• Class "Tritonobacteria" Williams et al. 2022 [UBA1439]
  • "Tritonobacterales" Williams et al. 2022

Phylum "Omnitrophota"

Phylogeny of Omnitrophota[9][10][11]

"Omnitrophia" "Omnitrophales" "Velamenicoccia" "Aquiviventales" "Gorgyraeales" "Pluralincolimonadales" JABMRG01 "Taenaricolales" "Aquitaenarimonadales" "Tantalellales" "Kappaeales" "Velesiimonadales" "Kaelpiales" "Duberdicusellales" "Ghiorseimicrobiales" "Gygaeellales" "Velamenicoccales" "Zapsychrales"

• Class "Omnitrophia" (OP3)
  • "Omnitrophales"
• Class "Velamenicoccia" Pallen, Rodriguez-R & Alikhan 2022 [Koll11]
  • "Aquitaenarimonadales" Seymour et al. 2022
  • "Aquiviventales" Seymour et al. 2022
  • "Duberdicusellales" Seymour et al. 2022
  • "Ghiorseimicrobiales" Seymour et al. 2022
  • "Gorgyraeales" Seymour et al. 2023 [UBA10183]
  • "Gygaeellales" Seymour et al. 2023 [GIF10]
  • "Candidatus Orphnella" Williams et al. 2021 {JABMRG01: JABMRG01}
  • "Kaelpiales" Seymour et al. 2022 [4484-49]
  • "Kappaeales" Seymour et al. 2022
  • "Pluralincolimonadales" Seymour et al. 2022
  • "Taenaricolales" Seymour et al. 2022
  • "Tantalellales" Seymour et al. 2022 [SKK-01]
  • "Velamenicoccales" Pallen, Rodriguez-R & Alikhan 2022 [UBA1572]
  • "Velesiimonadales" Seymour et al. 2022
  • "Zapsychrales" Seymour et al. 2022

Clade Spirochaetobacteriobiontes

Phylum "Babelota"

• Class "Babeliae" (sic) Yeoh et al. 2016 (TM6)
  • "Babelales" corrig. Yeoh et al. 2016

Phylum Spirochaetota

Phylogeny of Spirochaetota[9][10][11]

"Leptospirae" Turneriellales Leptospirales Leptospiria "Euspirochaetae" Brevinematia Brevinematales Brachyspiria Brachyspirales "Exilispiria" "Exilispirales" Spirochaetia Borreliales "Entomospirales" DSM-17781 Spirochaetales E Spirochaetales A "Salinispirales" "Marispirochaetales" "Sediminispirochaetales" Sphaerochaetales Treponematales

• Class Leptospiria Chuvochina et al. 2024
  • Leptospirales corrig. Gupta et al. 2014
  • Turneriellales Chuvochina et al. 2024
• Class Brevinematia Chuvochina et al. 2024
  • Brevinematales Gupta et al. 2014
• Class Brachyspiria Chuvochina et al. 2024
  • Brachyspirales corrig. Gupta et al. 2014
• Class "Exilispiria" Pallen, Rodriguez-R & Alikhan 2022
  • "Exilispirales" Pallen, Rodriguez-R & Alikhan 2022
• Class Spirochaetia Paster 2020
  •  ?Spirochaetales Buchanan 1917
  • Borreliales Chuvochina et al. 2024
  • Spirochaeta cellobiosiphila Breznak & Warnecke 2008 {DSM-17781: DSM-17781}
  • "Entomospirales" Pallen, Rodriguez-R & Alikhan 2022 [WRBN01]
  • "Marispirochaetales" Pallen, Rodriguez-R & Alikhan 2022 [JC444]
  • "Salinispirales" Pallen, Rodriguez-R & Alikhan 2022 [DSM-27196]
  • "Sediminispirochaetales" Pallen, Rodriguez-R & Alikhan 2022 [DSM-16054]
  • Sphaerochaetales Chuvochina et al. 2024
  • Spirochaeta thermophila Aksenova et al. 1992 {Spirochaetales_A}
  • Spirochaetales_E
  • Treponematales Chuvochina et al. 2024

Clade Planctobacteria

Phylum Planctomycetota

Phylogeny of Planctomycetota[9][10][11]

"Uabimicrobiia" "Uabimicrobiales" "Brocadiia" "Brocadiales" Phycisphaeria Sedimentisphaerales Tepidisphaerales Phycisphaerales Planctomycetia Isosphaerales Gemmatales Planctomycetales Pirellulales

• Class "Uabimicrobiia" Lodha, Narvekar & Karodi 2021
  • "Uabimicrobiales" Lodha, Narvekar & Karodi 2021
• Class "Brocadiia" JLodha, Narvekar & Karodi 2021
  • "Brocadiales" Jetten et al. 2010
  • "Hypogeohydatales" Wu et al. 2023 [YC14-17: JP1]
  • "Wukongales" Wu et al. 2023 [YC18]
• Class Phycisphaeria Oren, Parte & Garrity 2016
  • Phycisphaerales Fukunaga et al. 2010
  • Sedimentisphaerales Spring et al. 2018
  • Tepidisphaerales Kovaleva et al. 2015
• Class Planctomycetia Ward 2020
  • Gemmatales Dedysh et al. 2020
  • Isosphaerales Dedysh et al. 2020
  • Pirellulales Dedysh et al. 2020
  • Planctomycetales Schlesner & Stackebrandt 1987 em. Ward 2011

Phylum Chlamydiota

• Class Chlamydiia Horn 2011
  • "Similichlamydiales" Pallen, Rodriguez-R & Alikhan 2022 [Hat2]
  • Chlamydiales Storz & Page 1971 em. Everett et al. 1999 (Parachlamydiales, "Anoxychlamydiales", "Simkaniales")

Phylum Verrucomicrobiota

Phylogeny of Verrucomicrobiota[9][10][11]

Lentisphaeria Lentisphaerales Oligosphaerales Victivallales Kiritimatiellae "Spyradenecales" Kiritimatiellales "Verrucomicrobiia" Opitutales "Limisphaerales" "Methylacidiphilales" Terrimicrobiales Verrucomicrobiales

• Class Lentisphaeria Cho et al. 2004
  • Lentisphaerales Cho et al. 2004
  • Oligosphaerales Qiu et al. 2013
  • Victivallales Cho et al. 2004
• Class Kiritimatiellae Spring et al. 2017
  • Kiritimatiellales Spring et al. 2017 [incl. Tichowtungiales]
  • "Spyradenecales" Pallen, Rodriguez-R & Alikhan 2022 [RFP12]
• Class "Verrucomicrobiia" Oren, Parte & Garrity 2016 ex Cavalier-Smith 2020
  • Opitutales Choo et al. 2007 (Puniceicoccales)
  • "Limisphaerales" Podosokorskaya et al. 2023
  • "Methylacidiphilales" Op den Camp 2009 ["Pedosphaerales"]
  • Terrimicrobiales García-López et al. 2020 ("Chthoniobacterales")
  • Verrucomicrobiales Ward-Rainey et al. 1996 em. Yoon et al. 2008

Clade

Phylum "Saltatorellota"

• Class "Saltatorellae" Wiegand et al. 2019
  •  ?"Saltatorellales" Wiegand et al. 2019

Phylum "Hinthialibacterota"

• Class "Hinthialibacteria" Williams et al. 2022
  • "Hinthialibacterales" Williams et al. 2022

Phylum "Sumerlaeota"

• Class "Sumerlaeia" Kadnikov et al. 2018
  • "Sumerlaeales" Kadnikov et al. 2018

Phylum "Poribacteriota"

• Class ?"Entoporibacteria" Podell et al. 2019
• Class ?"Pelagiporibacteria" Podell et al. 2019

Phylum "Hydrogenedentota"

• Class "Hydrogenedentia" Chuvochina et al. 2023 (NKB19)
  • "Hydrogenedentales" corrig. Chuvochina et al. 2023

Clade FCB group

Phylum "Heilongiota"

• Class "Heilongiia" corrig. Zhang et al. 2022
  •  ?"Heilongiales" Zhang et al. 2022

Phylum "Neomarinimicrobiota"

•  ?"Candidatus Neomarinimicrobium" Rinke et al. 2013 corrig. Oren et al. 2020 (Candidate phylum SAR406; candidate phylum Marine Group A)

Phylum "Tianyaibacteriota"

•  ?"Candidatus Tianyaibacterium" corrig. Cui et al. 2021

Phylum "Fermentibacterota"

• Class "Fermentibacteria" Kirkegaard 2016 ("Aegiribacteria")
  • "Fermentibacterales" Kirkegaard 2016 (Hyd24-12)

Phylum "Latescibacterota"

• Class "Handelsmanbacteria" (RIF27)
• Class "Latescibacteria" Chuvochina et al. 2023
  • "Latescibacterales" Chuvochina et al. 2023 (WS3)

Phylum "Krumholzibacteriota"

• Class "Krumholzibacteriia" corrig. Youssef et al. 2019
  • "Delphibacterales" (DOLZORAL124_64_63; LZORAL124-64-63)
  • "Krumholzibacteriales" Youssef et al. 2019 (UBP1)

Phylum Gemmatimonadota

• Class "Glassbacteria" (RIF5)
• Class Gemmatimonadetes Zhang et al. 2003
  •  ?"Palaucibacterales" Aldeguer-Riquelme, Antón & Santos 2022 [PAUC43f]
  • Gemmatimonadales Zhang et al. 2003
  • Longimicrobiales Pascual et al. 2016

Phylum "Hydrothermota"

• Class "Stahlbacteria" (WOR-3)
• Class "Hydrothermia" Chuvochina et al. 2019 ex Chuvochina et al. 2023
  • "Hydrothermales" Chuvochina et al. 2019 ex Chuvochina et al. 2023 (EM3)
  • "Caldipriscales" Pallen, Rodriguez-R & Alikhan 2022 [LBFQ01]

Phylum "Cloacimonadota"

• Class "Cloacimonadia" Williams et al. 2021 ex Chuvochina et al. 2023
  • "Candidatus Celaenobacter" Williams et al. 2021 {JGIOTU-2: JGIOTU-2}
  • "Cloacimonadales" Williams et al. 2021 ex Chuvochina et al. 2023 (WWE1)

Phylum Fibrobacterota

• Class "Raymondbacteria" (RIF7)
• Class Chitinivibrionia Sorokin et al. 2014
  • Chitinivibrionales Sorokin et al. 2014 ["Chitinispirillales"]
• Class Fibrobacteria Spain et al. 2012
  • Fibrobacterales Spain et al. 2012 ["Fibromonadales"]

Phylum "Electryoneota"

• Class "Electryoneia" Williams et al. 2022
  • Electryoneales Williams et al. 2022
  • Hatepunaeales Williams et al. 2022
• Class "Tariuqbacteria" Vigneron, Vincent & Lovejoy 2023
  •  ?Tariuqbacterales Vigneron, Vincent & Lovejoy 2023

Phylum "Marinisomatota"

• Class "Marinisomatia"
  • "Marinisomatales" (SAR406)

Phylum "Cosmopoliota"

• Class "Cosmopolitia" (sic) Zhang et al. 2023
  • "Cosmopoliales" Zhang et al. 2023 (SAR406)

Phylum Calditrichota

• Class Calditrichia Kublanov et al. 2022
  • Calditrichales Kublanov et al. 2020

Phylum Bacteroidota

Phylogeny of Bacteroidota[9][10][11]

"Kapaibacteriia" "Kapaibacteriales" "Kryptonia" "Kryptoniales" "Ignavibacteriia" Ignavibacteriales "Chlorobiia" Chlorobiales Rhodothermia Balneolales Rhodothermales Bacteroidia Cytophagales Chitinophagales Sphingobacteriales Bacteroidales Flavobacteriales

• Class "Kapaibacteriia" corrig. Chuvochina et al. 2023 (OPB56)
  • "Kapaibacteriales" corrig. Chuvochina et al. 2023
• Class "Kryptonia"
  • "Kryptoniales"
• Class "Ignavibacteriia" Oren, Parte & Garrity 2016 ex Cavalier-Smith 2020
  • Ignavibacteriales Iino et al. 2010
• Class "Chlorobiia" Oren, Parte & Garrity 2016 ex Cavalier-Smith 2020
  • Chlorobiales Gibbons & Murray 1978
• Class Rhodothermia Munoz, Rossello-Mora & Amann 2017
  • Balneolales Munoz, Rossello-Mora & Amann 2017
  • Rhodothermales Munoz, Rossello-Mora & Amann 2017
• Class Bacteroidia Krieg 2012
  • Bacteroidales Krieg 2012 [Marinilabiliales; "Sulfidibacteriales"]
  • Chitinophagales Munoz, Rossello-Mora & Amann 2017 [Saprospirales]
  • Cytophagales Leadbetter 1974 [Cyclobacteriales]
  • Flavobacteriales Bernardet 2012
  • Sphingobacteriales Kämpfer 2012

Clade Proteobacteria

Phylum "Canglongiota"

• Class "Canglongiliia" Zhang et al. 2022
  •  ?"Canglongiales" Zhang et al. 2022

Phylum Acidobacteriota

Phylogeny of Acidobacteriota[9][10][11]

"Holophagia" Thermotomaculales Acanthopleuribacterales Holophagales "Fischerbacteria" "Guanabaribacteriia" "Guanabaribacteriales" "Aminicenantia" "Aminicenantales" Thermoanaerobaculia Thermoanaerobaculales "Polarisedimenticolia" "Polarisedimenticolales" Vicinamibacteria Vicinamibacterales Blastocatellia "Chloracidobacteriales" Blastocatellales "Acidobacteriia" "Acidoferrales" Bryobacterales Acidobacteriales

• Class "Holophagia" Oren, Parte & Garrity 2016
  • Acanthopleuribacterales Fukunaga et al. 2008
  • Holophagales Fukunaga et al. 2008
  • Thermotomaculales Dedysh & Yilmaz 2018
• Class "Fischerbacteria" (RIF25)
• Class "Guanabaribacteriia" corrig. Tschoeke et al. 2020
  • "Guanabaribacteriales" corrig. Tschoeke et al. 2020
• Class "Aminicenantia" Chuvochina et al. 2023
  • "Aminicenantales" Kadnikov et al. 2019 ex Chuvochina et al. 2023 (OP8)
• Class Thermoanaerobaculia Dedysh & Yilmaz 2018
  •  ?"Multivorales" Nguyen 2022
  • Thermoanaerobaculales Dedysh & Yilmaz 2018
• Class "Polarisedimenticolia" Flieder et al. 2021
  • "Polarisedimenticolales" Flieder et al. 2021
• Class Vicinamibacteria Dedysh & Yilmaz 2018
  •  ?"Marinacidobacterales" Nguyen 2022
  • Vicinamibacterales Dedysh & Yilmaz 2018
• Class Blastocatellia Pascual et al. 2016
  •  ?"Frugalibacteriales" Ruhl et al. 2022
  • Blastocatellales Pascual et al. 2016
  • "Chloracidobacteriales"
• Class Acidobacteriia Thrash & Coates 2010
  •  ?"Versatilivorales" Nguyen 2022
  • Acidobacteriales Cavalier-Smith 2002 [Terriglobales]
  • "Acidoferrales" Epihov et al. 2021
  • Bryobacterales Dedysh & Yilmaz 2018 ["Solibacterales"]

Phylum SAR324

• Class Lambdaproteobacteria Anantharaman et al. 2016 (RIF24)

Phylum Bdellovibrionota

Phylogeny of Bdellovibrionota[9][10][11]

Oligoflexia Oligoflexales Silvanigrellales Bdellovibrionia Bdellovibrionales Bacteriovoracia Bacteriovoracales

• Class Oligoflexia Nakai et al. 2014
  • Oligoflexales Nakai et al. 2014
  • Silvanigrellales Hahn et al. 2017
• Class Bdellovibrionia Waite et al. 2020
  • Bdellovibrionales Garrity, Bell & Lilburn 2006
• Class Bacteriovoracia Waite et al. 2020
  • Bacteriovoracales Hahn et al. 2017

Phylum Lernaellota

• Class "Lernaellia" Williams et al. 2022 [FEN-1099]
  • "Lernaellales" Williams et al. 2022

Phylum "Binatota"

• Class "Binatia" Chuvochina et al. 2019 ex Chuvochina et al. 2023
  • "Binatales" Chuvochina et al. 2019 ex Chuvochina et al. 2023

Phylum Myxococcota

Phylogeny of Myxococcota[9][10][11]

Myxococcia Myxococcales "Bradymonadia" Bradymonadales Polyangia Haliangiales Nannocystales Polyangiales

• Class Myxococcia Waite et al. 2020
  • Myxococcales Tchan, Pochon & Prévot 1948 [Archangiales; "Angiobacteriales"; "Angiococcales"] (fruiting gliding bacteria)
• Class "Bradymonadia" corrig. Chuvochina et al. 2023
  • Bradymonadales Wang et al. 2015
• Class Polyangiia corrig. Waite et al. 2020
  • Haliangiales Waite et al. 2020
  • Nannocystales (Reichenbach 2007) Waite et al. 2020
  • Polyangiales Tchan, Pochon & Prevot 1948 [Myxobacterales]

Phylum "Deferrisomatota"

• Class "Deferrisomatia" Waite et al. 2020
  • Deferrisomatales Waite et al. 2020

Phylum "Deferrimicrobiota"

• Class "Anaeroferrophilia" corrig. Murphy et al. 2021
  • "Anaeroferrophilales" Murphy et al. 2021
• Class "Deferrimicrobiia" corrig. Begmatov et al. 2022
  • Deferrimicrobiales Begmatov et al. 2022

Phylum "Moduliflexota"

• Class "Moduliflexia" Sekiguchi et al. 2015 (KSB3)
  • "Moduliflexales" Sekiguchi et al. 2015

Phylum "Methylomirabilaeota"

• Class "Methylomirabilia" Chuvochina et al. 2023
  • "Methylomirabilales" Cabrol et al. 2020 ex Chuvochina et al. 2023
  • "Rokubacteriales"

Phylum "Tectimicrobiota"

• Class "Entotheonellia" Chuvochina et al. 2023
  • "Entotheonellales" Schmidt et al. 2000 ex Chuvochina et al. 2023

Phylum Nitrospinota

• Class "Nitrospinia" Lucker et al. 2013
  • "Nitrospinales" Lucker et al. 2013

Phylum Nitrospirota

• Class "Thermodesulfovibrionia" Umezawa et al. 2021
  •  ?"Mariimomonadales" Yoon et al. 2023
  • "Thermodesulfovibrionales" Umezawa et al. 2021 ["Magnetobacteriales"]
• Class "Nitrospiria" Oren et al. 2015
  • "Nitrospirales" Garrity & Holt 2001
  • "Troglogloeales" Yu et al. 2022 ["Manganitrophales"]

Phylum Desulfobacterota F

• Class Desulfuromonadia Waite et al. 2020
  • Desulfuromonadales corrig. Kuever, Rainey & Widdel 2006
  • Geobacterales Waite et al. 2020

Phylum Thermodesulfobacteriota

Phylogeny of Thermodesulfobacteriota[9][10][11]

"Zymogeniia" "Zymogeniales" Desulfomonilia Desulfomonilales Syntrophia Syntrophales Desulfobulbia Desulfobulbales Dissulfuribacteria Dissulfuribacterales "Thermodesulfobacteriia" Thermodesulfobacteriales Desulfofervidia "Desulfofervidales" Desulfovibrionia Desulfovibrionales Desulfobaccia Desulfobaccales Desulfarculia "Adiutricales" Desulfarculales Syntrophobacteria Syntrophobacterales "Desulfatiglandia" Desulfatiglandales Desulfobacteria Desulfobacterales

• Class "Zymogeniia" corrig. Murphy et al. 2021
  • "Zymogeniales" Murphy et al. 2021
• Class Desulfomonilia Waite et al. 2020
  • Desulfomonilales Waite et al. 2020
• Class Syntrophia Waite et al. 2020
  • Syntrophales Waite et al. 2020
• Class Dissulfuribacteria Waite et al. 2020
  • Dissulfuribacterales Waite et al. 2020
• Class Thermodesulfobacteriia corrig. Hatchikian, Ollivier & Garcia 2002
  • Thermodesulfobacteriales Hatchikian, Ollivier & Garcia 2002
• Class Desulfobulbia Waite et al. 2020
  • Desulfobulbales Waite et al. 2020
• Class Desulfofervidia Waite et al. 2020
  • "Desulfofervidales" Waite et al. 2020
• Class Desulfovibrionia Waite et al. 2020
  • Desulfovibrionales Kuever, Rainey & Widdel 2006
• Class Desulfobaccia Waite et al. 2020
  • Desulfobaccales Waite et al. 2020
• Class Desulfarculia Waite et al. 2020
  • "Adiutricales" Waite et al. 2020
  • Desulfarculales corrig. Kuever, Rainey & Widdel 2006
• Class Syntrophobacteria Waite et al. 2020
  • Syntrophobacterales Waite et al. 2020
• Class "Desulfatiglandia" Pallen, Rodriguez-R & Alikhan 2022 [DSM-4660]
  • Desulfatiglandales Waite et al. 2020
• Class Desulfobacteria Waite et al. 2020
  • Desulfobacterales Kuever, Rainey & Widdel 2006

Phylum "Calescibacteriota"

• Class "Calescibacteriia" Chuvochina et al. 2023 (EM19)
  • "Calescibacteriales" Chuvochina et al. 2023

Phylum Desulfobacterota G

• Class Syntrophorhabdia Waite et al. 2020
  • Syntrophorhabdales Waite et al. 2020

Phylum Acidulodesulfobacteriota

• Class "Acidulodesulfobacteriia" (sic) Pallen, Rodriguez-R & Alikhan 2022 [SZUA-79]
  • "Acidulidesulfobacterales" corrig. Tan et al. 2019

Phylum "Thermosulfidibacterota"

• Class "Thermosulfidibacteria" Cavalier-Smith 2020 ex Chuvochina et al. 2023
  • "Thermosulfidibacterales" Cavalier-Smith 2020 ex Chuvochina et al. 2023

Phylum "Chrysiogenota"

• Class Chrysiogenetes Garrity & Holt 2002
  • Chrysiogenales Garrity & Holt 2002

Phylum Deferribacterota

• Class Deferribacteres Huber & Stetter 2002 em. Jumas-Bilak et al. 2009
  • Deferribacterales Huber & Stetter 2002 ["Geovibriales"]

Phylum Aquificota

• Class "Desulfurobacteriia" Chuvochina et al. 2023
  • Desulfurobacteriales Gupta & Lali 2014
• Class "Aquificia" Oren, Parte & Garrity 2016 ex Cavalier-Smith 2020
  • Aquificales Reysenbach 2002 emend. Gupta & Lali 2013
  • "Hydrogenothermales" Chuvochina et al. 2023

Phylum Campylobacterota

• Class Desulfurellia Waite et al. 2020
  • Desulfurellales Kuever, Rainey & Widdel 2006
• Class "Campylobacteria" Oren, Parte & Garrity 2016 ex Waite et al. 2017
  • Campylobacterales Garrity, Bell & Lilburn 2006
  • Nautiliales Miroshnichenko et al. 2004

Phylum Leptospirillaeota

• Class "Leptospirillia" Chuvochina et al. 2023
  • "Leptospirillales" Chuvochina et al. 2023

Phylum Pseudomonadota

Phylogeny of Pseudomonadota[9][10][11]

"Caulobacteria" "Enterousiales" "Scatocolales" "Rickettsiidae" Rhizobiales A Rickettsiales A "Pelagibacterales" Rickettsiales "Caulobacteridae" cont'd "Mariprofundia" Mariprofundales "Magnetococcia" Magnetococcales "Pseudomonadia" "Enterobacteriae" "Comchoanobacterales" "Thalassomicrobiales" "Evansiales" Enterobacterales UBA11654 "Foliamicales" "Porifericomitales" "Porisulfidales" Immundisolibacterales Acidiferrobacterales Arenicellales "Acidithiobacillidae" Acidithiobacillales Thiohalorhabdales "Neisseriidae" Burkholderiales "Pseudomonadidae" cont'd

• Class "Anaeropigmentatia" corrig. Murphy et al. 2021
  •  ?Anaeropigmentatales corrig. Murphy et al. 2021
• Class Caulobacteria" Oren, Parte & Garrity 2016 ex Cavalier-Smith 2020
  •  ?"Futianiales" Liu et al. 2022
  •  ?"Methylospongiales" Podell et al. 2020
  • "Enterousiales" Pallen, Rodriguez-R & Alikhan 2022 {Rs-D84}
  • "Scatocolales" Pallen, Rodriguez-R & Alikhan 2022 {RF32}
  • Sublass "Rickettsiidae" corrig. Ferla et al. 2013
    • Liberibacter Fagen et al. 2014 {Rhizobiales_A: Rhizobiaceae_A}
    • "Pelagibacterales" Grote et al. 2012
    • Rickettsiales_A
    • Rickettsiales Gieszczykiewicz 1939 emend. Dumler et al. 2001

Phylogeny of "Caulobacteridae"[9][10][11]

"Paracaedibacterales" Holosporales "Caedimonadales" "Micavibrionales" "Puniceispirillales" "Thalassobaculales" "Marivibrionales" Rhodospirillales A "Dongiales" "Tagaeales" "Stellales" "Elsterales" "Oceanibaculales" Kiloniellales "Inquilinales" "Azospirillales" Acetobacterales" Rhodospirillales "Reyranellales" "Planktothermales" "Tistrellales" "Geminicoccales" "Ferrovibrionales" Minwuiales Sneathiellales Sphingomonadales "Zavarziniales" Micropepsales Caulobacterales Rhodobacterales "Parvibaculales" Hyphomicrobiales

• • Sublass "Caulobacteridae" Ferla et al. 2013
    • "Acetobacterales" Chuvochina et al. 2023
    • "Caedimonadales"
    • "Azospirillales" Chuvochina et al. 2023
    • Caulobacterales Henrici & Johnson 1935 (Hyphomonadales; Maricaulales; Parvularculales)
    • "Dongiales" Chuvochina et al. 2023
    • "Elsterales" Chuvochina et al. 2023
    • "Ferrovibrionales" Chuvochina et al. 2023
    • "Geminicoccales" Chuvochina et al. 2023
    • Holosporales Szokoli et al. 2016
    • Hyphomicrobiales Douglas 1957 (Rhizobiales)
    • "Inquilinales" Pallen, Rodriguez-R & Alikhan 2022 [DSM-16000]
    • Kiloniellales Wiese et al. 2009
    • "Marivibrionales" Pallen, Rodriguez-R & Alikhan 2022
    • "Methylospongiales" Podell et al. 2020
    • "Micavibrionales"
    • Micropepsales Harbison et al. 2016
    • Minwuiales Sun et al. 2018
    • "Oceanibaculales" Chuvochina et al. 2023
    • "Paracaedibacterales" Chuvochina et al. 2023
    • "Parvibaculales"Chuvochina et al. 2023
    • "Planktothermales" Zhou et al. 2020 [UBA7887]
    • "Puniceispirillales" Chuvochina et al. 2023
    • "Reyranellales" Chuvochina et al. 2023
    • Rhodobacterales Garrity, Bell & Lilburn 2006
    • Rhodospirillales A
    • Rhodospirillales Pfennig & Truper 1971
    • Sphingomonadales Yabuuchi & Kosako 2006 (Emcibacterales; Iodidimonadales; Kordiimonadales; Rhodothalassiales)
    • Sneathiellales Kurahashi et al. 2008
    • Stellales Pallen, Rodriguez-R & Alikhan 2022 [ATCC43930]
    • "Tagaeales" Pallen, Rodriguez-R & Alikhan 2022 [CACIAM-22H2]
    • "Thalassobaculales" Chuvochina et al. 2023
    • "Tistrellales" Chuvochina et al. 2023
    • "Zavarziniales" Chuvochina et al. 2023
• Class "Mariprofundia" Cavalier-Smith 2020 ["Zetaproteobacteria"]
  • Mariprofundales Makita et al. 2017
• Class "Magnetococcia" Parks et al. 2018 ex Chuvochina et al. 2023
  • Magnetococcales Bazylinski et al. 2013
• Class Pseudomonadia" Oren, Parte & Garrity 2016
  •  ?"Ferritrophicales" Weiss et al. 2007
  •  ?"Kopriimonadales" Kwon et al. 2005c
  •  ?Methylohalomonadales Sorokin & Merkel 2023
  •  ?"Rariloculales" Nguyen et al. 2022
  •  ?"Spongiifermentales" Nguyen et al. 2023
  •  ?"Tethybacterales" Taylor et al. 2021
  •  ?"Taraoceanobacteraceae" Zhou et al. 2020 {UBA11654}
  • Acidiferrobacterales Kojima, Shinohara & Fukui 2015
  • Arenicellales Teramoto, Yagyu & Nishijima 2015
  • "Comchoanobacterales" Needham et al. 2022
  • Enterobacterales Adeolu et al. 2016 ("Bacteriales"; Orbales)
  • "Foliamicales" Nguyen et al. 2023
  • Immundisolibacterales Corteselli, Aitken & Singleton 2017
  • "Johnevansiales" Chuvochina et al. 2023
  • "Porifericomitales" Nguyen et al. 2023
  • "Poriferisulfidales" Nguyen et al. 2023 ex Chuvochina et al. 2023
  • "Thalassomicrobiales" Zhou et al. 2020 [SAR86]
  • Subclass "Acidithiobacillidae" Cavalier-Smith 2020
    • Acidithiobacillales Garrity, Bell & Lilburn 2005
    • Thiohalorhabdales Sorokin & Merkel 2023
  • Subclass "Neisseriidae" Cavalier-Smith 2020 (Betaproteobacteria)
    • Burkholderiales Garrity, Bell & Lilburn 2006 ("Ferrovales"; "Gallionellales"; Hydrogenophilales; Methylophilales; Neisseriales; Nitrosomonadales; "Procabacteriales"; Rhodocyclales; Spirillales; Sulfuricellales)

Phylogeny of "Pseudomonadidae"[9][10][11]

"Chromatiae" "Nitrosococcales" UBA4486 Beggiatoales "Inmiraniales" "Thioalbales" "Tenderiales" "Sulfuriflexales" Thiohalospirales "Thiohalophilales" Thiohalomonadales "Reidiellales" "Thiogranales" Thiohalobacterales "Thiopontiales" Chromatiales Methylococcales Thiotrichales "Ectothiorhodospirales" Halofilales" "Granulosicoccales" "Halothiobacillales" "Competibacterales" "Nitrococcales" "Macondimonadales" Nevskiales SLND01 "Woeseiales" "Steroidobacterales" Xanthomonadales Cardiobacteriales "Thioglobales" "Thiomicrospirales" "Piscirickettsiales" "Berkiellales" "Francisellales" "Aquicellales" "Diplorickettsiales" "Coxiellales" Legionellales "Pelagibaculales" "Vibrionales" Pseudomonadales

• • Subclass "Pseudomonadidae" Cavalier-Smith 2020
    • "Aquicellales" Pallen, Rodriguez-R & Alikhan 2022 [DSM-16500]
    • Beggiatoales Buchanan 1957
    • "Berkiellales" Chuvochina et al. 2023
    • Cardiobacteriales Garrity, Bell & Lilburn 2005
    • Chromatiales Imhoff 2005
    • "Competibacterales" Chuvochina et al. 2023
    • "Coxiellales" Chuvochina et al. 2023
    • "Diplorickettsiales" Chuvochina et al. 2023
    • "Ectothiorhodospirales" Chuvochina et al. 2023
    • "Francisellales" Chuvochina et al. 2023
    • "Granulosicoccales" Chuvochina et al. 2023
    • "Halofilales" Pallen, Rodriguez-R & Alikhan 2022
    • "Halothiobacillales" Chuvochina et al. 2023
    • "Inmiraniales" Pallen, Rodriguez-R & Alikhan 2022 [DSM-100275]
    • Legionellales Garrity, Bell & Lilburn 2005
    • Methylococcales Bowman 2005
    • "Macondimonadales" Pallen, Rodriguez-R & Alikhan 2022 [UBA5335]
    • Natronospira Sorokin, Kublanov & Khijniak 2017 {SLND01: SLND01}
    • Nevskiales Naushad et al. 2015 ("Salinisphaerales")
    • "Nitrococcales" Chuvochina et al. 2023
    • "Nitrosococcales" Chuvochina et al. 2023
    • "Pelagibaculales" Pallen, Rodriguez-R & Alikhan 2022 [HP12]
    • "Piscirickettsiales" Chuvochina et al. 2023
    • Pseudomonadales Orla-Jensen 1921 (Cellvibrionales; Moraxellales; Oceanospirillales)
    • "Reidiellales" Pallen, Rodriguez-R & Alikhan 2022 [GCF-013343005]
    • "Riflewellaceae" Zhou et al. 2020 {UBA4486}
    • "Steroidobacterales" Chuvochina et al. 2023
    • "Sulfuriflexales" Pallen, Rodriguez-R & Alikhan 2022 [AKS1]
    • "Tenderiales" Chuvochina et al. 2023
    • "Thioalbales" Pallen, Rodriguez-R & Alikhan 2022 [DSM-26407]
    • "Thioglobales" Pallen, Rodriguez-R & Alikhan 2022 [PS1]
    • "Thiogranales" Pallen, Rodriguez-R & Alikhan 2022 [DSM-19610]
    • Thiohalobacterales Sorokin & Merkel 2023
    • Thiohalomonadales Sorokin & Merkel 2023
    • "Thiohalophilales" Pallen, Rodriguez-R & Alikhan 2022 {UBA6429}
    • Thiohalospirales Sorokin & Merkel 2023
    • "Thiomicrospirales" Chuvochina et al. 2023
    • "Thiopontiales" Pallen, Rodriguez-R & Alikhan 2022 [GCF-002020875]
    • Thiotrichales Garrity, Bell & Lilburn 2005
    • "Vibrionales" ("Aerobacterales"; Aeromonadales; Alteromonadales; Kangiellales; Pasteurellales)
    • "Woeseiales" Chuvochina et al. 2023
    • Xanthomonadales Saddler & Bradbury 2005 non Chadefaud 1950a (Lysobacterales)

List of Candidate Phyla

• "Qinglongiota" Zhang et al. 2022
• "Salinosulfoleibacteria" Tazi et al. 2006
• "Teskebacteria" Dojka 1998 (WS1)

See also

• Branching order of bacterial phyla (Woese, 1987)
• Branching order of bacterial phyla (Gupta, 2001)
• Branching order of bacterial phyla (Cavalier-Smith, 2002)
• Branching order of bacterial phyla (Rappe and Giovanoni, 2003)
• Branching order of bacterial phyla (Battistuzzi et al.,2004)
• Branching order of bacterial phyla (Ciccarelli et al., 2006)
• Branching order of bacterial phyla after ARB Silva Living Tree
• Branching order of bacterial phyla (Genome Taxonomy Database, 2018)
• Bacterial phyla
• List of Archaea genera
• List of bacteria genera
• LPSN, list of accepted bacterial and archaeal names
• Human microbiome project
• Microorganism

References

1. J.P. Euzéby. "Bacteria". List of Prokaryotic names with Standing in Nomenclature (LPSN). Retrieved 2021-06-27.
2. Sayers; et al. "Bacteria". National Center for Biotechnology Information (NCBI) taxonomy database. Retrieved 2021-03-20.
3. All-Species Living Tree Project."16S rRNA-based LTP release 132". Silva Comprehensive Ribosomal RNA Database. Retrieved 2020-08-20.
4. Parks, DH; Chuvochina, M; Waite, DW; Rinke, C; Skarshewski, A; Chaumeil, PA; Hugenholtz, P (November 2018). "A standardized bacterial taxonomy based on genome phylogeny substantially revises the tree of life". Nature Biotechnology. 36 (10): 996–1004. bioRxiv 10.1101/256800. doi:10.1038/nbt.4229. PMID 30148503. S2CID 52093100.
5. Parks, DH; Chuvochina, M; Chaumeil, PA; Rinke, C; Mussig, AJ; Hugenholtz, P (September 2020). "A complete domain-to-species taxonomy for Bacteria and Archaea". Nature Biotechnology. 38 (9): 1079–1086. bioRxiv 10.1101/771964. doi:10.1038/s41587-020-0501-8. PMID 32341564. S2CID 216560589.
6. Chaumeil, PA; Mussig, AJ; Hugenholtz, P; Parks, DH (15 November 2019). "GTDB-Tk: a toolkit to classify genomes with the Genome Taxonomy Database". Bioinformatics. 36 (6): 1925–1927. doi:10.1093/bioinformatics/btz848. PMC 7703759. PMID 31730192.
7. Almeida, Alexandre; Nayfach, Stephen; Boland, Miguel; Strozzi, Francesco; Beracochea, Martin; Shi, Zhou Jason; Pollard, Katherine S.; Sakharova, Ekaterina; Parks, Donovan H.; Hugenholtz, Philip; Segata, Nicola; Kyrpides, Nikos C.; Finn, Robert D. (20 July 2020). "A unified catalog of 204,938 reference genomes from the human gut microbiome". Nature Biotechnology. 39 (1): 105–114. doi:10.1038/s41587-020-0603-3. PMC 7801254. PMID 32690973.
8. Nayfach, Stephen; et al. (9 November 2020). "A genomic catalog of Earth's microbiomes". Nature Biotechnology. 39 (4): 499–509. doi:10.1038/s41587-020-0718-6. PMC 8041624. PMID 33169036.
9. "GTDB release 09-RS220". Genome Taxonomy Database. Retrieved 10 May 2024.
10. "ar53_r220.sp_label". Genome Taxonomy Database. Retrieved 10 May 2024.
11. "Taxon History". Genome Taxonomy Database. Retrieved 10 May 2024.