钻头螺亚科

钻头螺科（学名：Subulinidae），又名锥蜗牛科，原是柄眼目玛瑙螺亚目非洲大蜗牛总科之下的一个小型热带会呼吸空气的蜗牛的科级生物分类单元，是一类陆上（英语：terrestrial molluscs）肺螺类腹足纲软体动物。

提示：此条目的主题不是钻螺亚科。

钻头螺科 化石时期：Lower Paleocene至Recent[1] PreЄ Є O S D C P T J K Pg N
断顶刍螺Rumina decollata
科学分类
界：	动物界 Animalia
门：	软体动物门 Mollusca
纲：	腹足纲 Gastropoda
目：	柄眼目 Stylommatophora
亚目：	玛瑙螺亚目 Achatinina
总科：	玛瑙螺总科 Achatinoidea
科：	钻头螺科 Subulinidae P. Fischer & Crosse, 1877[2]
Genera
See text

2017年12月，本科与非洲大蜗牛科（玛瑙螺科）合并，原属本科的亚科都成为了玛瑙螺科的亚科；就只馀下原来的钻头螺亚科（学名：Subulininae）的物种^[3]。

分布

本科物种分布于全世界^[1]。

解剖

本科物种的单倍体染色体数量从26到35不等，详见Barker (2001)^[4]。

分类

2005年分类

根据2005年《布歇特和洛克罗伊的腹足类分类_》的弯尿道类非正式群组)^[5] (The following unreferenced unchecked genera are probably from Schileyko (1999))，本科包括以下各个亚科：

• 钻头螺亚科 Subulininae P. Fischer & Crosse, 1877
• Coeliaxinae Pilsbry, 1907^[6]，异名：Cryptelasminae Germain, 1916; Pyrgininae Germain, 1916
• Glessulinae Godwin-Austen, 1920
• 钻螺亚科 Opeatinae Thiele, 1931
• Petriolinae Schileyko, 1999
• Rishetiinae Schileyko, 1999
• Rumininae Wenz, 1923
• Stenogyrinae P. Fischer & Crosse, 1877 - synonym: Obeliscinae Thiele, 1931

属

各亚科所包括的属如下：

钻头螺亚科 Subulininae P. Fischer & Crosse, 1877

• Abaconia Clench, 1938^[7]
• 锥蜗牛属 Allopeas H. B. Baker, 1935^[8]
• Beckianum Baker, 1961^[9]
• Curvella Chaper, 1885^[10]
• Dysopeas Baker, 1927^[11]
• Euonyma Melvill & Ponsonby, 1896^[12]
• Hypolysia Melvill & Ponsonby, 1901^[13]
• Lamellaxis Strebel & Pfeffer, 1882^[9]
• Leptopeas Baker, 1927^[11]
• Leptinaria Beck, 1837^[9]
• Luntia Smith, 1898^{[来源请求]}
• Micropeas Connolly, 1923^{[来源请求]}
• Neoglessula Pilsbry, 1909^{[来源请求]}
• Paropeas Pilsbry, 1906^[14][15]
• Pelatrinia Pilsbry, 1907^{[来源请求]}
• Prosopeas Mörch, 1876^{[来源请求]}
• Pseudoglessula Boettger, 1892^{[来源请求]}
• Pseudopeas Putzeys, 1899^{[来源请求]}
• Striosubulina Thiele, 1933^{[来源请求]}
• Vegrandinia Salvador, Cunha & Simone, 2013^[16]
• Subulina Beck, 1837^[9] - type genus of the family Subulinidae^[5]
• Zootecus Westerlund, 1887^[17]

Coeliaxinae

• Coeliaxis H. Adams & Angas, 1865 - type genus of the subfamily Coeliaxinae^[5]
• Cryptelasmus Pilsbry, 1907^[5][9]
• Ischnocion Pilsbry, 1907^{[来源请求]}
• Nannobeliscus Weyrauch, 1967^{[来源请求]}
• Neosubulina Smith, 1898^{[来源请求]}
• Pseudobalea Shuttleworth, 1854^{[来源请求]}
• Pyrgina Greef, 1882^[5]
• Thomea Girard, 1893^{[来源请求]}

Glessulinae

• Glessula Martens, 1860 - type genus of the subfamily Glessulinae^[5]

钻螺亚科 Opeatinae Thiele, 1931

• Eremopeas Pilsbry, 1906^{[来源请求]}
• 钻螺属 Opeas Albers, 1850^[9] - type genus of the subfamily Opeatinae^[5]

Petriolinae

• Bocageia Girard, 1893^{[来源请求]}
• Ceras Dupuis & Putzeys, 1901^{[来源请求]}
• Chilonopsis Fischer de Waldheim, 1848^{[来源请求]}
• Cleostyla Dall, 1896^{[来源请求]}
• Comoropeas Pilsbry, 1906^{[来源请求]}
• Dictyoglessula Pilsbry, 1919^{[来源请求]}
• Homorus Albers, 1850^{[来源请求]}
• Ischnoglessula Pilsbry, 1919^{[来源请求]}
• Itiopiana Preston, 1910^{[来源请求]}
• Kempioconcha Preston, 1913^{[来源请求]}
• Liobocageia Pilsbry, 1919^{[来源请求]}
• Mabilliella Ancey, 1886^{[来源请求]}
• Notapalinus Connolly, 1923^{[来源请求]}
• Nothapalus von Martens, 1897^{[来源请求]}
• Oleata Ortiz de Zarate, 1959^{[来源请求]}
• Oreohomorus Pilsbry, 1919^[18]
• Petriola Dall, 1905 - type genus of the subfamily Petriolinae^[5]
• Subuliniscus Pilsbry, 1919^{[来源请求]}
• Subulona Martens, 1889^{[来源请求]}

Rishetiinae

• Bacillum Theobald, 1840^{[来源请求]}
• Eutomopeas Pilsbry, 1946^{[来源请求]}
• Rishetia Godwin-Austen, 1920 - type genus of the subfamily Rishetiinae^[5]
• 卷轴螺属 Tortaxis Pilsbry, 1906^{[来源请求]}

Rumininae

• Balfouria Crosse, 1884^{[来源请求]}
• Krapfiella Preston, 1911^{[来源请求]}
• Lubricella Haas, 1928^{[来源请求]}
• Namibiella Zilch, 1954^{[来源请求]}
• Obeliscella Jousseaume, 1899^{[来源请求]}
• Riebeckia von Martens, 1883^{[来源请求]}
• Rumina Risso, 1826^[9][19] - type genus of the subfamily Rumininae^[5]
• Xerocerastus Kobelt & Möllendorff, 1902^{[来源请求]}

Stenogyrinae

• Chryserpes Pilsbry, 1906^{[来源请求]}
• Cupulella Aguayo & Jaume, 1948^{[9][来源请求]}
• Dolicholestes Pilsbry, 1906^{[来源请求]}
• Lyobasis Pilsbry, 1903^{[来源请求]}
• Neobeliscus Pilsbry, 1896^{[来源请求]}
• Obeliscus Beck, 1837^[5][9][20]
• Ochroderma Ancey, 1885^{[来源请求]}
• Ochrodermatina Thiele, 1931^{[来源请求]}
• Ochrodermella Pilsbry, 1907^{[来源请求]}
• Plicaxis Sykes, 1903^{[来源请求]}
• Promoussonius Pilsbry, 1906^{[来源请求]}
• Protobeliscus Pilsbry, 1906^{[来源请求]}
• Pseudocolumna Wenz, 1920^{[来源请求]}
• Rectobelus Baker, 1927^{[来源请求]}
• Rhodea Adams, 1855^{[21][来源请求]}
• Stenogyra Shuttleworth, 1854 - type genus of the subfamily Stenogyrinae^[5]
• Synapterpes Pilsbry, 1896^{[来源请求]}
• Zoniferella Pilsbry, 1906^{[来源请求]}

2017年分类

现时本亚科只馀下下列各属^[22][23]：

• 锥蜗牛属 Allopeas H. B. Baker, 1935
• Beckianum H. B. Baker, 1961^[9]
• Curvella Chaper, 1885
• Dysopeas H. B. Baker, 1927^[11]
• Euonyma Melvill & Ponsonby, 1896
• Fortuna Schlickum & Strauch, 1972
• Hypolysia Melvill & Ponsonby, 1901
• Lamellaxis Strebel & Pfeffer, 1882^[9]
• Leptinaria H. Beck, 1837^[9]
• Leptopeas H. B. Baker, 1927^[11]
• Micropeas Connolly, 1923^{[来源请求]}
• Neoglessula Pilsbry, 1909^{[来源请求]}
• Paropeas Pilsbry, 1906^[14][15]
• Pelatrinia Pilsbry, 1907^{[来源请求]}
• Prosopeas Mörch, 1876^{[来源请求]}
• Pseudoglessula O. Boettger, 1892
• Pseudopeas Putzeys, 1899^{[来源请求]}
• Striosubulina Thiele, 1933^{[来源请求]}
• 钻头螺属 Subulina H. Beck, 1837^[9]：本亚科的模式属^[5]
• Vegrandinia Salvador, Cunha & Simone, 2013^[16]
• Zootecus Westerlund, 1887^{[17][来源请求]}

物种

下列部分本科物种：

• 锥蜗牛属 Allopeas
  • 锥蜗牛 Allopeas clavulinum^[24]
  • 大锥蜗牛 Allopeas gracilis
  • 细锥蜗牛 Allopeas pyrgula
• 钻螺属 Opeas Albers, 1850^[9] - type genus of the subfamily Opeatinae^[5]
  • 细钻螺 Opeas gracile (Hutton, 1834)^[25]
  • 条纹钻螺 Opeas striatissimum (Gredler, 1882)^[26]
  • 小囊钻螺
  • 棒形钻螺
  • 索形钻螺
  • 四川钻螺
• 卷轴螺属
  • 竖卷轴螺
  • 柑卷轴螺
• Subulina属
  • 原锥蜗牛 Subulina octona

参考文献

1. Family summary for Subulinidae. AnimalBase. 2006-02-21 [2011-03-15]. （原始内容存档于2015-12-27） （英语）.
2. Fischer P. & Crosse (1877). Mission scientifique au Mexique et dans l'Amérique Centrale. Rechearches zoologiques (7)1(6): 592.
3. Bouchet, P.; Rocroi, J.-P.; Hausdorf, B.; Kaim, A.; Kano, Y.; Nützel, A.; Parkhaev, P.; Schrödl, M.; Strong, E.E. Revised Classification, Nomenclator and Typification of Gastropod and Monoplacophoran Families. Malacologia. 2017, 61 (1–2): 1–526. OCLC 7296100651. S2CID 91051256. doi:10.4002/040.061.0201.
4. Barker, G. M. Gastropods on Land: Phylogeny, Diversity and Adaptive Morphology. Barker G. M. (编). The biology of terrestrial molluscs. Oxon, UK: CABI Publishing. 2001: 1–146, cited pages: 139 and 142. ISBN 0-85199-318-4 （英语）.
5. Bouchet, P.; Rocroi, J.-P.; Frýda, J.; Hausdorf, B.; Ponder, W.; Valdés, Á. & Warén, A. Classification and nomenclator of gastropod families. Malacologia (Hackenheim, Germany: ConchBooks). 2005, 47 (1–2): 1–397. ISBN 3-925919-72-4. OCLC 61351079. OL 42989175M. S2CID 83847814.
6. Pilsbry, H. A. ?. Manual of Conchology. 1907, 2 (18): 72 （英语）.
7. Bouchet, Philippe. Cone with the wind: a pelagie marine speeies deseribed as an endemie land snail From the Bahamas (PDF). THE NAUTILUS. 2002, 116 (1): 32-35 [2021-05-30]. （原始内容 (PDF)存档于2021-06-02）.
8. Baker, H. B. Review of the anatomy of Pupillidae and related groups. HA Pilsbry. 1935.
9. "Mollusca" （页面存档备份，存于互联网档案馆）. Diversidad Biológica Cubana, accessed 23 March 2011.
10. Van Bruggen, Adolf Cornelis. A new land snail, Curvella transvaalensis, from southern Africa (Gastropoda, Pulmonata: Subulinidae). Annals of the Transvaal Museum. 1978 [2021-05-30]. （原始内容存档于2021-06-02）.
11. Baker, Horace B. 1930: The mollusca collected by the University of Michigan – Williamson expedition in Venezuela. Occasional Paper of Museum of zoology. 210:1-51
12. F.L.S, James Cosmo Melvill M. A.; F.Z.S, John Henry Ponsonby. XLVII.—Descriptions of new terrestrial mollusca from South Africa. Journal of Natural History. 2009-10-06 [2021-05-30]. doi:10.1080/00222939608680460. （原始内容存档于2021-06-02） （英语）.
13. James Cosmo Melvill M. A., F. L. S.; F.Z.S, John Henry Ponsonby. XLI.—Descriptions of Fourteen new Species of Terrestrial Mollusca from South Africa. Journal of Natural History. 2009-12-01 [2021-05-30]. doi:10.1080/03745480109442927. （原始内容存档于2021-06-02） （英语）.
14. Pilsbry H. A. (1906). In: Tryon G. W. & Pilsbry H. A. (1906) Manual of Conchology (2)18: 14.
15. Naggs F. (1994). "The reproductive anatomy of Paropeas achatinaceum and a new concept of Paropeas (Pulmonata: Achatinoidea: Subulinidae)". Journal of Molluscan Studies 60(2): 175-191. doi:10.1093/mollus/60.2.175.
16. Salvador R. B., Cunha C. M. & Simone L. R. L. (2013). "Taxonomic revision of the orthalicid land snails (Pulmonata: Stylommatophora) from Trindade Island, Brazil". Journal of Natural History 47(13-14): 949-961. doi: 10.1080/00222933.2012.759290.
17. "Genus summary for Zootecus" （页面存档备份，存于互联网档案馆）. AnimalBase, accessed 11 october 2010.
18. Wronski T. & Hausdorf B. (2009). "Oreohomorus apio new species from Uganda (Gastropoda: Subulinidae)". Journal of Conchology 40(1): 55-60. abstract Archive.is的存档，存档日期2010-12-29.
19. "Genus summary for Rumina" （页面存档备份，存于互联网档案馆）. AnimalBase, accessed 11 october 2010.
20. （葡萄牙文） Salgado N. C. & Coelho A. C. S. (2003). "Moluscos terrestres do Brasil (Gastrópodes operculados ou não, exclusive Veronicellidae, Milacidae e Limacidae)". Revista de Biología Tropical 51(Suppl. 3): 149-189. (with English abstract), PDF （页面存档备份，存于互联网档案馆）.
21. "New paper on Rhodea" （页面存档备份，存于互联网档案馆）. Ashbreure's posterous, published 1 April 2011, accessed 1 April 2011.
22. Bieler R, Bouchet P, Gofas S, Marshall B, Rosenberg G, La Perna R, Neubauer TA, Sartori AF, Schneider S, Vos C, ter Poorten JJ, Taylor J, Dijkstra H, Finn J, Bank R, Neubert E, Moretzsohn F, Faber M, Houart R, Picton B, Garcia-Alvarez O (编). Subulinidae P. Fischer & Crosse, 1877. MolluscaBase –通过WoRMS.
23. Bieler R, Bouchet P, Gofas S, Marshall B, Rosenberg G, La Perna R, Neubauer TA, Sartori AF, Schneider S, Vos C, ter Poorten JJ, Taylor J, Dijkstra H, Finn J, Bank R, Neubert E, Moretzsohn F, Faber M, Houart R, Picton B, Garcia-Alvarez O (编). Subulininae P. Fischer & Crosse, 1877. MolluscaBase –通过WoRMS.
24. 国际贝库：锥蜗牛. 台湾贝类资料库. [2009-08-11]. （原始内容存档于2016-03-04）.
25. 中国科学院动物研究所. 细钻螺. 《中国动物物种编目数据库》. 中国科学院微生物研究所. [2009-04-28]. （原始内容存档于2016-03-05）.
26. 中国科学院动物研究所. 条纹钻螺. 《中国动物物种编目数据库》. 中国科学院微生物研究所. [2009-04-28]. （原始内容存档于2016-03-05）.

延伸阅读

• Schileyko A. A. (1999). "Treatise on Recent Terrestrial Pulmonate Molluscs. Part 4 Draparnaudiidae, Caryodidae, Macrocyclidae, Acavidae, Clavatoridae, Dorcasiidae, Sculptariidae, Corillidae, Plectopylidae, Megalobulimidae, Strophocheilidae, Cerionidae, Achatinidae, Subulinidae, Glessulidae, Micractaeonidae, Ferrussaciidae". In: Ruthenica., Supplement, 2(4): 435-564, Moscow, ISSN 0136-0027.

外部链接