Panxiosteidae

Panxiosteidae is an extinct family of arthrodire placoderms that lived during the Devonian period.

Panxiosteidae Temporal range: Devonian: Givetian to Early Frasnian, ~387.7–382.7 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
Various species of Plourdosteus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	†Placodermi
Order:	†Arthrodira
Suborder:	†Brachythoraci
Clade:	†Eubrachythoraci
Clade:	†Pachyosteomorphi
Superfamily:	†Dunkleosteoidea
Family:	†Panxiosteidae Wang, 1979
Genera
Janiosteus Kimberleyichthys? Panxiosteus Plourdosteus

Phylogeny

The family Panxiosteidae was erected by Wang in 1979. Members of the family are noted for showing morphologically intermediate traits between coccosteids and dunkleosteids.^[1] In the 2010 Carr & Hlavin phylogenetic study, Panxiosteidae was recognized as the sister taxon to the family Dunkleosteidae, which together comprised the superfamily Dunkleosteoidea (one of the three major clades of Eubrachythoraci).^[2]

The phylogeny of Panxiosteidae from the 2013 Zhu & Zhu study is shown in the cladogram below:^[3]

Eubrachythoraci	Coccosteomorphi Coccosteus cuspidatus Harrytoombsia elegans Mcnamaraspis kaprios Incisoscutoidea Compagopiscis croucheri Incisoscutum ritchiei Incisoscutum sarahae Camuropiscidae Latocamurus coulthardi Camuropiscis laidlawi Rolfosteus canningensis Tubonasus lennardensis Fallacosteus turneri Aspinothoracidi Dinichthys herzeri Hadrosteus rapax Gorgonichthys clarki Heintzichthys gouldii Selenosteidae Stenosteus angustopectus Gymnotrachelus hydei Rhinosteus parvulus Pachyosteus bulla Dunkleosteoidea Westralichthys uwagedensis Protitanichthys rockportensis Panxiosteidae Panxiosteus ocullus Janiosteus timanicus Plourdosteus canadensis Dunkleosteidae Eastmanosteus calliaspis Xiangshuiosteus wui Eastmanosteus pustulosus Kiangyousteus yohii Golshanichthys asiatica Dunkleosteus amblyodoratus Dunkleosteus terrelli Dunkleosteus raveri	Pachyosteomorphi

However, the subsequent 2016 Zhu et al. study using a larger morphological dataset recovered Panxiosteidae well outside of Dunkleosteoidea, instead within Coccosteomorphi and then Coccosteoidea as the sister group of Coccosteidae, as shown in the cladogram below:^[1]

Eubrachythoraci

Coccosteomorphi Coccosteoidea Coccosteidae Millerosteus minor Coccosteus cuspidatus Dickosteus threiplandi Watsonosteus fletti Protitanichthys rockportensis Panxiosteidae Plourdosteus canadensis Panxiosteus ocullus Janiosteus timanicus Incisoscutoidea Harrytoombsia elegans Torosteus tuberculatus Torosteus pulchellus Mcnamaraspis kaprios Compagopiscis croucheri Trematosteus fontanellus Camuropiscidae Incisoscutum ritchiei Incisoscutum sarahae Rolfosteus canningensis Tubonasus lennardensis Fallacosteus turneri Camuropiscis laidlawi Latocamurus coulthardi Pachyosteomorphi Rhachiosteus pterygiatus Dunkleosteoidea Eastmanosteus calliaspis Eastmanosteus pustulosus Kiangyousteus yohii Golshanichthys asiatica Westralichthys uwagedensis Dunkleosteus raveri Dunkleosteus terrelli Dunkleosteus amblyodoratus Heterostiidae Heterosteus ingens Yinosteus major Aspinothoracidi Tapinosteus heintzi Bullerichthys fascidens Kendrickichthys cavernosus Bruntonichthys multidens Dinichthys herzeri Hadrosteus rapax Gorgonichthys clarki Selenosteidae Heintzichthys gouldii Pachyosteus bulla Gymnotrachelus hydei Stenosteus angustopectus Brachyosteus dietrichi Melanosteus occitanus Rhinosteus parvulus

References

1. Zhu, You-An; Zhu, Min; Wang, Jun-Qing (1 April 2016). "Redescription of Yinostius major (Arthrodira: Heterostiidae) from the Lower Devonian of China, and the interrelationships of Brachythoraci". Zoological Journal of the Linnean Society. 176 (4): 806–834. doi:10.1111/zoj.12356. ISSN 0024-4082.
2. Carr, R.K.; Hlavin, W.J. (2010). "Two new species of Dunkleosteus Lehman, 1956, from the Ohio Shale Formation (USA, Famennian) and the Kettle Point Formation (Canada, Upper Devonian), and a cladistic analysis of the Eubrachythoraci (Placodermi, Arthrodira)". Zoological Journal of the Linnean Society. 159 (1): 195–222. doi:10.1111/j.1096-3642.2009.00578.x.
3. You-An Zhu; Min Zhu (2013). "A redescription of Kiangyousteus yohii (Arthrodira: Eubrachythoraci) from the Middle Devonian of China, with remarks on the systematics of the Eubrachythoraci". Zoological Journal of the Linnean Society. 169 (4): 798–819. doi:10.1111/zoj12089.