2019 in paleomalacology

This list 2019 in paleomalacology is a list of new taxa of ammonites and other fossil cephalopods, as well as fossil gastropods, bivalves and other molluscs that are scheduled to be described during the year 2019, as well as other significant discoveries and events related to molluscan paleontology that are scheduled to occur in the year 2019.

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General research

• Description of an assemblage of conchiferan mollusks from the Cambrian Shackleton Limestone (Antarctica) is published by Claybourn et al. (2019).^[1]
• A study on Early Triassic recovery of ammonites and gastropods after the Permian–Triassic extinction event is published by Pietsch et al. (2019).^[2]
• A study on the impact of the Cenomanian-Turonian boundary event and other Cretaceous oceanic anoxic events on generic- and species-level molluscan diversity, extinction rates and ecological turnover is published by Freymueller, Moore & Myers (2019).^[3]
• A juvenile ammonite specimen assigned to the genus Puzosia and four marine gastropods are reported from the Cretaceous Burmese amber of Myanmar by Yu et al. (2019).^[4]
• A study on the timing and nature of recovery of benthic marine ecosystems of Antarctica after the Cretaceous–Paleogene extinction event, as indicated by data from fossils of benthic molluscs, is published by Whittle et al. (2019).^[5]

Ammonites

Research

A Puzosia (Bhimaites) species juvenile shell in Burmese amber

• Dimorphic Indonesian ammonite Macrocephalites keeuwensis is reported from the core of Jara Dome (Kutch district, India) by Jain (2019).^[6]
• Description of new fossils of members of the genus Lytohoplites from the Neuquén Basin (Argentina), taxonomic revision of the Lytohoplites species occurring in this basin, and a study on the ontogeny, probable sexual dimorphism and spectrum of intraspecific variability of members of this genus, is published by Vennari & Aguirre-Urreta (2019).^[7]
• A study on the intraspecific variation through the ontogeny of the Late Cretaceous ammonite Scaphites whitfieldi from the Western Interior of the United States is published by Klein & Landman (2019).^[8]
• A lower jaw of Spathites puercoensis is described from the Turonian Carlile Member of the Mancos Shale (New Mexico, United States) by Landman et al. (2019).^[9]
• A study on periodic variations in carbon and oxygen stable isotope profiles from Campanian and Maastrichtian ammonites belonging to the genus Baculites, and on their implications for inferring the life histories of these ammonites, is published by Ellis & Tobin (2019).^[10]
• A study on the hydrostatic properties of the orthoconic morphotype of Baculites compressus and on their implications for the knowledge of the mode of life of this animal is published by Peterman et al. (2019).^[11]
• A study on the ecology and growth of baculitid and scaphitid ammonites from the Maastrichtian Owl Creek Formation (Mississippi, United States), evaluating whether these taxa may have migrated, is published by Ferguson et al. (2019).^[12]
• A study on a sutural anomaly of ammonite specimens known as sutural pseudoinversion is published by Rogov (2019).^[13]
• A study on the systematics of Late Cretaceous ammonites from the southeastern San Juan Basin (Sandoval County, New Mexico) and on the stratigraphic framework of their fossils is published by Sealey & Lucas (2019).^[14]
• A study on the ontogenetic trajectories of septal spacing between succeeding chambers in Hypophylloceras subramosum and Phyllopachyceras ezoense is published online by Iwasaki, Iwasaki & Wani (2019).^[15]
• A study on the intra- and interspecific variation and ontogenetic trajectories of the kossmaticeratid species Maorites densicostatus and Maorites seymourianus is published by Morón-Alfonso (2019).^[16]
• A pathological ammonite specimen characterized by a pronounced left-right asymmetry in both ornamentation and suture lines is described from the Toarcian of southern France by Jattiot et al. (2019).^[17]
• A study on the morphology and function of hook-like structures found in the body chambers of six specimens of Rhaeboceras halli is published online by Kruta et al. (2019).^[18]

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Albanesiceras[19]	Gen. et sp. nov	Valid	Galácz	Middle Jurassic (Bathonian)		Italy	A member of the family Perisphinctidae. The type species is A. damianoi.
Albanites americanus[20]	Sp. nov	Valid	Brayard et al.	Early Triassic		United States ( Idaho)
Allocrioceras irregulare[21]	Sp. nov	Valid	Kennedy & Kaplan	Late Cretaceous (Turonian)		Germany
Anagaudryceras calabozoi[22]	Sp. nov	Valid	Raffi & Olivero in Raffi, Olivero & Milanese	Late Cretaceous (Campanian)	Cerro Toro Formation Rabot Formation Santa Marta Formation	Antarctica (James Ross Island)  Chile
Anagaudryceras sanctuarium[22]	Sp. nov	Valid	Raffi & Olivero in Raffi, Olivero & Milanese	Late Cretaceous (Campanian/Maastrichtian boundary)	Snow Hill Island Formation	Antarctica (James Ross Island)
Anagaudryceras subcompressum[22]	Sp. nov	Valid	Raffi & Olivero in Raffi, Olivero & Milanese	Late Cretaceous (Maastrichtian)	Snow Hill Island Formation	Antarctica (James Ross Island)
Araxoceltites[23]	Gen. et 3 sp. nov	Valid	Korn, Ghaderi & Tabrizi	Permian (Changhsingian)		Iran	Genus includes new species A. sanestapanus, A. laterocostatus and A. cristatus.
Araxoceras insolens[24]	Sp. nov	Valid	Korn & Ghaderi	Permian (Wuchiapingian)		Iran
Araxoceras truncatum[24]	Sp. nov	Valid	Korn & Ghaderi	Permian (Wuchiapingian)		Iran
Balearites berryae[25]	Sp. nov	Valid	Vermeulen et al.	Early Cretaceous		Spain
Beckeroceras[26]	Gen. et comb. nov	Valid	Naglik, De Baets & Klug	Devonian (Emsian)	Dzhaus Beds	Uzbekistan	A member of Agoniatitida belonging to the family Mimosphinctidae. The type species is "Erbenoceras" khanakasuensis Yatskov (1990).
Beulia[27]	Gen. et sp. nov	Valid	Korn & Price	Devonian (Famennian)		Germany	Genus includes new species B. wunderlichi.
Borealiceras[28]	Gen. et comb. nov	Valid	Vermeulen et al.	Early Cretaceous		Germany	Genus includes "Crioceras" stadtlaenderi Müller (1892).
Caribouceras[20]	Gen. et sp. nov	Valid	Brayard et al.	Early Triassic		United States ( Idaho)	Genus includes new species C. slugense.
Carpathiceras[29]	Gen. et comb. et sp. nov	Valid	Meister & Schlögl	Early Jurassic (Sinemurian)		Slovakia	A member of Asteroceratinae. The type species is "Coroniceras (Paracoroniceras)" bordoti Guérin-Franiatte (1966); genus also includes new species C. misiki.
Carthaginites multituberculatus[30]	Sp. nov	Valid	Kennedy in Gale, Kennedy & Walaszczyk	Late Cretaceous (Cenomanian)	Cauvery Basin	India	A member of the family Turrilitidae.
Chieseiceras dolomiticum[31]	Sp. nov	Valid	Brack & Rieber	Middle Triassic (Late Anisian)		Italy
Chtelnicaceras[29]	Gen. et sp. nov	Valid	Meister & Schlögl	Early Jurassic (Sinemurian)		Slovakia	An ammonite with an intermediate morphology between Arietitidae and Oxynoticeratidae. Genus includes new species C. janus.
Damesites africanus[32]	Sp. nov	Valid	Kennedy & Lobitzer	Late Cretaceous (Cenomanian)	Odukpani Formation	Nigeria	A member of the family Desmoceratidae.
Docidoceras higginsi[33]	Sp. nov	Valid	Chandler	Middle Jurassic (Bajocian)		United Kingdom
Dzhaprakoceras brodheadense[34]	Sp. nov	Valid	Work & Mason	Carboniferous (Tournaisian)	Borden Formation	United States ( Kentucky)
Eogunnarites? elegans[30]	Sp. nov	Valid	Kennedy in Gale, Kennedy & Walaszczyk	Late Cretaceous (Cenomanian)	Cauvery Basin	India	A member of the family Kossmaticeratidae.
Epophioceras nodosus[29]	Sp. nov	Valid	Meister & Schlögl	Early Jurassic (Sinemurian)		Slovakia
Euomphaloceras varicostatum[30]	Sp. nov	Valid	Kennedy in Gale, Kennedy & Walaszczyk	Late Cretaceous (Cenomanian)	Cauvery Basin	India
Falciclymenia cabrierensis[27]	Sp. nov	Valid	Korn & Price	Late Devonian		France
Finiclymenia czarnockii[35]	Sp. nov	Valid	Korn & Price	Devonian (Famennian)		Central Europe	A gonioclymeniid ammonite.
Finiclymenia langei[35]	Sp. nov	Valid	Korn & Price	Devonian (Famennian)		Central Europe	A gonioclymeniid ammonite.
Flickia bullata[36]	Sp. nov	Valid	Kennedy & Lobitzer	Late Cretaceous (Cenomanian)	Odukpani Formation	Nigeria
Follotites bighillensis[34]	Sp. nov	Valid	Work & Mason	Carboniferous (Tournaisian)	Borden Formation	United States ( Kentucky)
Gaudryceras submurdochi[22]	Sp. nov	Valid	Raffi & Olivero in Raffi, Olivero & Milanese	Late Cretaceous (Campanian)	Rabot Formation Santa Marta Formation	Antarctica (James Ross Island)
Glaphyrites taimyricus[37]	Sp. nov	Valid	Konovalova & Borissenkov in Borissenkov, Konovalova & Kutygin	Carboniferous	Makarovskaya Formation	Russia ( Krasnoyarsk Krai)
Glaucoceras[38]	Gen. et comb. nov	Valid	Matamales-Andreu et al.	Early Cretaceous (Hauterivian)		Spain   Switzerland	A new genus for "Baculites" renevieri Ooster (1860).
Glyptoxoceras (?) tenuicostatum[21]	Sp. nov	Valid	Kennedy & Kaplan	Late Cretaceous (Turonian)		Germany	A member of the family Diplomoceratidae.
Hoevelia[27]	Gen. et sp. nov	Valid	Korn & Price	Devonian (Famennian)		Germany	Genus includes new species H. schindewolfi.
Homoeoplanulites todaginensis[39]	Sp. nov	Valid	Poulton & Hall	Middle Jurassic (Callovian)		Canada ( British Columbia)
Hoploscaphites macer[40]	Sp. nov	Valid	Landman et al.	Late Cretaceous (Maastrichtian)	Bearpaw Shale Pierre Shale	United States ( Colorado  Montana  Wyoming)
Ivoites meshchankinae[26]	Sp. nov	Valid	Naglik, De Baets & Klug	Devonian (Emsian)		Czech Republic  Uzbekistan	A member of Agoniatitida belonging to the family Mimosphinctidae.
Kamerunoceras multinodosum[30]	Sp. nov	Valid	Kennedy in Gale, Kennedy & Walaszczyk	Late Cretaceous (Turonian)	Karai Formation	India
Kayutoceras pogrebitskyi[37]	Sp. nov	Valid	Borissenkov & Kutygin in Borissenkov, Konovalova & Kutygin	Carboniferous	Olen'ynskaya Formation	Russia ( Krasnoyarsk Krai)
Kirsoceras arteseptatum[41]	Sp. nov	Valid	Bockwinkel, Korn & Herd	Devonian (Famennian)		Germany
Kirsoceras gelamenum[41]	Sp. nov	Valid	Bockwinkel, Korn & Herd	Devonian (Famennian)		Germany
Kitabobactrites[26]	Gen. et sp. nov	Valid	Naglik, De Baets & Klug	Devonian (Emsian)		Czech Republic  Uzbekistan	A member of Bactritida belonging to the family Bactritidae. The type species is K. salimovae.
Kumetaceras[19]	Gen. et sp. nov	Valid	Galácz	Middle Jurassic (Bathonian)		Italy	A possible member of the family Spiroceratidae. The type species is K. enigmaticum.
Kunnamiceras[30]	Gen. et comb. nov	Valid	Kennedy in Gale, Kennedy & Walaszczyk	Late Cretaceous (Cenomanian)	Cauvery Basin	India  Madagascar	A member of the subfamily Acanthoceratinae. The type species is "Ammonites" tropicus Stoliczka (1865); genus also includes "Protacanthoceras" imperatoris Wright & Kennedy (1980).
Latiwitchellia atacamensis[42]	Sp. nov	Valid	Dietze & von Hillebrandt	Middle Triassic (Bajocian)		Chile
Lissoceras maerteni[43]	Sp. nov	Valid	Pavia & Fernandez-Lopez	Middle Jurassic (Bajocian)		France	A member of Haploceratoidea belonging to the family Lissoceratidae.
Lissoceras maizetense[43]	Sp. nov	Valid	Pavia & Fernandez-Lopez	Middle Jurassic (Bajocian)		France	A member of Haploceratoidea belonging to the family Lissoceratidae.
Lissoceras ovale[43]	Sp. nov	Valid	Pavia & Fernandez-Lopez	Middle Jurassic (Bajocian)	El Pedregal Formation	France  Germany  Hungary  Spain	A member of Haploceratoidea belonging to the family Lissoceratidae.
Lissoceras sturanii[43]	Sp. nov	Valid	Pavia & Fernandez-Lopez	Middle Jurassic (Bajocian)		France  Italy  Russia ( Dagestan)  Turkmenistan	A member of Haploceratoidea belonging to the family Lissoceratidae.
Lissoceras submediterraneum[43]	Sp. nov	Valid	Pavia & Fernandez-Lopez	Middle Jurassic (Bajocian)	El Pedregal Formation	France  Spain	A member of Haploceratoidea belonging to the family Lissoceratidae.
Loyezia roberti[25]	Sp. nov	Valid	Vermeulen et al.	Early Cretaceous (Hauterivian)		Spain	A member of the family Crioceratitidae.
Maeneceras tabasense[44]	Sp. nov	Valid	Korn et al.	Probably late early Famennian		Iran	A member of the family Sporadoceratidae.
Medioclymenia media[35]	Sp. nov	Valid	Korn & Price	Devonian (Famennian)		Central Europe	A gonioclymeniid ammonite.
Megatirolites[45]	Gen. et comb. nov	Valid	Balini et al.	Early Triassic (Olenekian)	Sefid-Kuh Limestone Tartaly Formation	Iran  Kazakhstan	A member of Ceratitida belonging to the superfamily Dinaritoidea and to the family Tirolitidae. The type species is "Tirolites" rossicus Kiparisova (1947); genus also includes "Tirolites" armatus Shevyrev (1968).
Menuites armatus[46]	Sp. nov	Valid	Shigeta in Shigeta, Izukura & Nishimura	Late Cretaceous (Campanian)	Chinomigawa Formation	Japan	A member of the family Pachydiscidae.
Mesodistichites[47]	Gen. et comb. et sp. nov	Valid	Krystyn et al.	Late Triassic (Norian)	Nayband Formation	Iran	A member of Ceratitida belonging to the family Distichitidae. The type species is "Distichites" tozeri Seyed-Emami (1975); genus also includes new species M. evolutus.
Metabactrites rakhmonovi[26]	Sp. nov	Valid	Naglik, De Baets & Klug	Devonian (Emsian)		Uzbekistan	A member of Agoniatitida belonging to the family Mimosphinctidae.
Microderoceras crassiornatum[29]	Sp. nov	Valid	Meister & Schlögl	Early Jurassic (Sinemurian)		Slovakia
Microderoceras serratum[29]	Sp. nov	Valid	Meister & Schlögl	Early Jurassic (Sinemurian)		Slovakia
Nelomites[41]	Nom. nov	Valid	Bockwinkel, Korn & Herd	Devonian (Famennian)		Kazakhstan	A replacement name for Melonites Bogoslovskii (1971).
Neophylloceras nodosum[46]	Sp. nov	Valid	Shigeta in Shigeta, Izukura & Nishimura	Late Cretaceous (Campanian)	Chinomigawa Formation	Japan	A member of the family Phylloceratidae.
Noridiscites nodosus[47]	Sp. nov	Valid	Krystyn et al.	Late Triassic (Norian)	Nayband Formation	Iran	A member of Ceratitida belonging to the family Noridiscitidae.
Oraniceras calvum[48]	Sp. nov	Valid	Gulyaev	Middle Jurassic (Bathonian)		Russia	A member of the family Parkinsoniidae.
Oraniceras issae[48]	Sp. nov	Valid	Gulyaev	Middle Jurassic (Bathonian)		Russia	A member of the family Parkinsoniidae.
Pachydiscus wrighti[49]	Sp. nov	Valid	Kennedy	Late Cretaceous		United Kingdom
Paracrioceras damaisi[28]	Sp. nov	Valid	Vermeulen et al.	Early Cretaceous
Parasolenoceras ribiraense[46]	Sp. nov	Valid	Shigeta in Shigeta, Izukura & Nishimura	Late Cretaceous (Campanian)	Chinomigawa Formation	Japan	A member of the family Diplomoceratidae.
Paroecotraustes minor[19]	Sp. nov	Valid	Galácz	Middle Jurassic (Bathonian)		Italy	A member of the family Oppeliidae.
Pernaceras[28]	Gen. et comb. nov	Valid	Vermeulen et al.	Early Cretaceous		France	Genus includes "Ammonites" assimilis Matheron (1880).
Phlycticeras tenue[19]	Sp. nov	Valid	Galácz	Middle Jurassic (Bathonian)		Italy	A member of the family Strigoceratidae.
Praechebbites[50]	Gen. et sp. et comb. nov	Valid	Becker et al.	Devonian (Emsian)		Morocco  China?  France?	A member of the family Mimosphinctidae belonging to the new subfamily Chebbitinae. The type species is P. debaetsi; genus might also include "Anetoceras (Erbenoceras)" mattei Feist (1970) and "Teicherticeras" ilanense Shen (1975).
Prolecanites akiyoshiensis[51]	Sp. nov	Valid	Fujikawa	Carboniferous	Akiyoshi Limestone	Japan
Protacanthoceras parva[30]	Sp. nov	Valid	Kennedy in Gale, Kennedy & Walaszczyk	Late Cretaceous (Cenomanian)	Cauvery Basin	India	A member of the subfamily Acanthoceratinae.
Pseudocrassiceras[52]	Gen. et comb et sp. nov	Valid	Rulleau & Jattiot	Early Jurassic (Toarcian)		France  Spain	A member of the family Hildoceratidae. The type species is "Ammonites" bayani Dumortier (1874); genus also includes "Ammonites" frantzi Reynès (1868), "Hildoceras" comense var. praegruneri Monestier (1931) (raised to the rank of a separate species Pseudocrassiceras praegruneri) and a new species P. arbaulti.
Pseudosonninia[53]	Gen. et sp. nov	Valid	Parent et al.	Middle Jurassic (Callovian)	Los Molles Formation	Argentina	A member of the family Oppeliidae. The type species is P. chacaymelehuensis.
Puzosia (Bhimaites) falx[30]	Sp. nov	Valid	Kennedy in Gale, Kennedy & Walaszczyk	Late Cretaceous (Cenomanian)	Cauvery Basin	India
Schlueterella stinnesbecki[54]	Sp. nov	Valid	Ifrim	Late Cretaceous (Turonian-Coniacian)		Mexico	A member of the family Diplomoceratidae.
Semilissoceras[43]	Gen. et 3 sp. et comb. nov	Valid	Pavia & Fernandez-Lopez	Middle Jurassic (Bajocian)		Algeria  France  Portugal  Spain  United Kingdom	A member of Haploceratoidea belonging to the family Lissoceratidae. The type species is S. ellipticum; genus also includes new species S. turgidulum and S. costellatum, as well as "Oppelia" depereti Flamand (1911) and "Lissoceras" semicostulatum Buckman (1923).
Sicilisphinctes[19]	Gen. et sp. nov	Valid	Galácz	Middle Jurassic (Bathonian)		Italy	A member of the family Perisphinctidae. The type species is S. incertus.
Siemiradzkia mangoldi[55]	Sp. nov	Valid	Metodiev	Middle Jurassic (Bathonian)	Bov Formation	Bulgaria  France	A member of the family Perisphinctidae.
Sikeliceras[19]	Gen. et sp. nov	Valid	Galácz	Middle Jurassic (Bathonian)		Italy	A possible member of the family Spiroceratidae. The type species is S. costatum.
Sokurella elshankae[48]	Sp. nov	Valid	Gulyaev	Middle Jurassic (Bathonian)		Russia	A member of the family Parkinsoniidae.
Sokurella pletniovkae[48]	Sp. nov	Valid	Gulyaev	Middle Jurassic (Bathonian)		Russia	A member of the family Parkinsoniidae.
Stephanoceras (Skirroceras) englandi[56]	Sp. nov	Valid	Chandler	Middle Jurassic (Lower Bajocian)	Inferior Oolite Formation	United Kingdom
Stephanoceras (Riccardiceras) eoeteosum[56]	Sp. nov	Valid	Chandler	Middle Jurassic (Middle or Upper Aalenian)	Inferior Oolite Formation	United Kingdom
Uptonia schlatteri[57]	Sp. nov	Valid	Fauré & Téodori	Early Jurassic (Pliensbachian)		France
Uzbekisphinctes[26]	Gen. et comb. nov	Valid	Naglik, De Baets & Klug	Early Devonian	Daleje Shale Dzhaus Beds Middle Kaub Formation	China  Czech Republic  Germany  Uzbekistan	A member of Agoniatitida belonging to the family Mimosphinctidae. The type species is "Teicherticeras" rudicostatum Bogoslovsky (1980); genus also includes "Convoluticeras" discordans Erben (1965) and "Teicherticeras" primigenitum Erben (1965).
Vedioceras fusiforme[24]	Sp. nov	Valid	Korn & Ghaderi	Permian (Wuchiapingian)		Iran	A member of the family Araxoceratidae.
Watinoceras elegans[30]	Sp. nov	Valid	Kennedy in Gale, Kennedy & Walaszczyk	Late Cretaceous (Cenomanian)	Cauvery Basin	India
Zelandites pujatoi[22]	Sp. nov	Valid	Raffi, Olivero & Milanese	Late Cretaceous (Campanian)	Rabot Formation Santa Marta Formation	Antarctica (James Ross Island)

Other cephalopods

Research

• A study on hydrostatic properties of early Paleozoic nautiloid and endoceratoid cephalopods, and on its implications for the knowledge of ecology of these cephalopods, is published by Peterman, Barton & Yacobucci (2019).^[58]
• A study on the distribution and diversity of cephalopods during the Cambrian–Ordovician transition is published by Fang et al. (2019).^[59]
• A study on cephalopod occurrences in South China and adjacent areas during the Middle to Late Ordovician, aiming to define biogeographic provinces, is published by Fang et al. (2019).^[60]
• A study on changes of geographic range size and rates of speciation and extinction in nautiloid and ammonoid cephalopods living in the North American Midcontinent Sea from the Carboniferous (Pennsylvanian)-early Permian is published by Kolis & Lieberman (2019).^[61]
• Ascocerid fossils are described from the Hirnantian?–Llandovery strata of the Eusebio Ayala and Vargas Peña formations (southeastern Paraguay) by Cichowolski et al. (2019), representing the first ascocerid record from high paleolatitudes of Gondwana.^[62]
• A juvenile specimen of Dolorthoceras, representing the first Devonian cephalopod specimen preserving nacreous structures and the first ectocochleate cephalopod preserving fibrous structures within its conch, is described from the Frasnian Domanic Formation (Russia) by Doguzhaeva (2019).^[63]
• A study on the response of cephalopod assemblages from the area of the present-day European Russia to environmental changes during the early Aptian Oceanic Anoxic Event is published by Rogov et al. (2019).^[64]
• A study on the paleobiology of cephalopods from the Albian Mahajanga Basin of Madagascar will be published by Hoffmann et al. (2019).^[65]
• A proposal for revision of the classification of nautiloid cephalopods presented in Treatise Part K is presented by King & Evans (2019), who name a new subclass Tarphyceratia, new order Rioceratida and new family Bactroceratidae.^[66]
• A study on embryonic and early juvenile development of shell in Peismoceras is published by Manda & Turek (2019).^[67]
• A study on changes of the morphological diversity of Late Permian coiled nautiloids is published online by Korn et al. (2019).^[68]
• A study on the paleobiology of Angulithes mermeti, based on new material from the upper Cenomanian of Egypt, and on its implications for the knowledge of the evolution of Cenomanian lineage of Angulithes, is published by Wilmsen & Nagm (2019).^[69]
• A study on the anatomy of Gordoniconus beargulchensis and on its implications for the knowledge of evolution of coleoids is published by Klug et al. (2019).^[70]
• A study on a putative Sinemurian octocoral Mesosceptron from Montagna del Casale (Sicily, Italy) is published by Pignatti et al. (2019), who reinterpret the fossils of this taxon as incompletely preserved coleoid rostra, and consider Mesosceptron to be a subjective junior synonym of the genus Atractites.^[71]
• A study on four specimens of Clarkeiteuthis conocauda from the Toarcian Posidonia Shale (Germany) preserved with small bony fish in their arm crowns, on the feeding behaviour of members of this species, and on the differences in the mode of life of C. conocauda, Passaloteuthis bisulcata and Hibolithes semisulcatus, is published by Jenny et al. (2019).^[72]
• A study on belemnite body-size dynamics across the Pliensbachian–Toarcian boundary in the Peniche section (Lusitanian Basin, Portugal), evaluating its implications for the knowledge of the impact of the Pliensbachian–Toarcian boundary warming event on belemnite body size, is published by Rita et al. (2019).^[73]
• New belemnite fossils are described from the Berriasian–Hauterivian of Arctic Canada and North-East Greenland by Mutterlose et al. (2019), who also evaluate the implications of these fossils for the knowledge of spatial distribution patterns of belemnites at northern high latitudes.^[74]

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Andamanoceras[75]	Gen. et sp. nov	Valid	Niko, Sone & Leman	Ordovician	Lower Setul Limestone	Malaysia	A member of Pseudorthocerida. Genus includes new species A. densiseptum.
Basiloceras[76]	Gen. et sp. nov	Valid	Pohle, Klug & Haas	Devonian (Eifelian and Givetian)		Morocco	A member of Oncocerida belonging to the family Acleistoceratidae. Genus includes new species B. goliath and B. david.
Beloitoceras thorslundi[77]	Sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden	A member of the family Oncoceratidae.
Cabaneroceras[78]	Gen. et sp. nov	Valid	Kröger & Gutiérrez-Marco	Ordovician (Darriwilian)		Spain	A member of Intejocerida. Genus includes new species C. aznari.
Cameroceras motsognir[77]	Sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden
Cimomia kurkurensis[79]	Sp. nov	Valid	Hewaidy, Qot & Moneer	Early Paleocene	Kurkur Formation	Egypt	A member of the family Hercoglossidae.
Clothoceras[77]	Gen. et sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden	A member of the family Proteoceratidae. Genus includes new species C. thornquisti.
Danoceras skalbergensis[77]	Sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden
Deltoidonautilus hassani[79]	Sp. nov	Valid	Hewaidy, Qot & Moneer	Late Paleocene	Tarawan Formation	Egypt	A member of the family Hercoglossidae.
Dicoelites aprilis[80]	Sp. nov	Valid	Dzyuba in Dzyuba, Mitta & Sherstyukov	Middle Jurassic (Bajocian)	Djangura Formation	Russia ( Karachay-Cherkessia)	A belemnite.
Dicoelites octobris[80]	Sp. nov	Valid	Dzyuba in Dzyuba, Mitta & Sherstyukov	Middle Jurassic (Bajocian)	Djangura Formation	Russia ( Karachay-Cherkessia)	A belemnite.
Discoceras amtjaernense[77]	Sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden
Discoceras nilssoni[77]	Sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden
Endoceras naekki[77]	Sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden
Engeseriteuthis[81]	Gen. et sp. nov	Valid	Fuchs et al.	Late Jurassic (late Kimmeridgian to ?early Tithonian)	Geisental Formation	Germany	A member of Muensterelloidea. Genus includes new species E. arcuatus.
Eromangateuthis[82]	Gen. et comb. nov	Valid	Fuchs	Early Cretaceous (Albian)	Allaru Formation	Australia	A member of Octobrachia belonging to the family Plesioteuthidae; a new genus for "Boreopeltis" soniae Wade (1993).
Etchesia[83]	Gen. et sp. nov	Valid	Fuchs	Late Jurassic (Tithonian)	Kimmeridge Clay Formation	United Kingdom	A member of Coleoidea belonging to the family Muensterellidae. Genus includes new species E. martilli.
Furudaloceras[77]	Gen. et sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden	A member of the family Cyrtogomphoceratidae. Genus includes new species F. tomtei.
Grafordites[84]	Gen. et sp. nov	Valid	Niko & Mapes	Carboniferous (Kasimovian)	Graford Formation	United States ( Texas)	A member of the family Dentoceratidae. The type species is G. mcleodi.
Gujiaonautilus[85]	Gen. et sp. nov	Valid	Miao et al.	Permian (Changhsingian)	Dalong Formation	China	A nautiloid. Genus includes new species G. longliensis.
Isbergoceras[77]	Gen. et 2 sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden	A member of the family Graciloceratidae. The type species is I. niger; genus also includes I. consobrinum.
Isorthoceras nikwis[77]	Sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden	A member of the family Proteoceratidae.
Isorthoceras sylphide[77]	Sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden	A member of the family Proteoceratidae.
Isorthoceras urdr[77]	Sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden	A member of the family Proteoceratidae.
Kullsbergoceras[77]	Gen. et sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden	A member of the family Cyrtogomphoceratidae. Genus includes new species K. nissei.
Lambeoceras (Lambeoceras) balashovi[86]	Sp. nov	Valid	Barskov & Marek	Ordovician (Katian)		Russia
Lambeoceras (Lambeoceras) hanae[86]	Sp. nov	Valid	Barskov & Marek	Ordovician (Katian)		Russia
Lambeoceras (Lambeoceras) sibiricum[86]	Sp. nov	Valid	Barskov & Marek	Ordovician (Katian)		Russia
Langgunites[75]	Gen. et sp. nov	Valid	Niko, Sone & Leman	Ordovician	Lower Setul Limestone	Malaysia	A member of Pseudorthocerida. Genus includes new species L. mucronulatus.
Malayorthoceras[75]	Gen. et sp. nov	Valid	Niko, Sone & Leman	Ordovician	Lower Setul Limestone	Malaysia	A member of Orthocerida. Genus includes new species M. gracilentum.
Muensterella jillae[81]	Sp. nov	Valid	Fuchs et al.	Late Cretaceous (Turonian)	South Bosque Formation	United States ( Texas)
Muensterella spinosa[81]	Sp. nov	Valid	Fuchs et al.	Late Jurassic (Tithonian)	Mörnsheim Formation	Germany
Muensterellina[81]	Gen. et sp. nov	Valid	Fuchs et al.	Middle Jurassic (Callovian)	Oxford Clay Formation	United Kingdom	A member of Muensterelloidea. Genus includes new species M. johnjagti.
Neotainoceras zhaoi[85]	Sp. nov	Valid	Miao et al.	Permian (Changhsingian)	Dalong Formation	China	A nautiloid.
Nodopleuroceras gujiaoense[85]	Sp. nov	Valid	Miao et al.	Permian (Changhsingian)	Dalong Formation	China	A nautiloid.
Ordogeisonoceras uppsalaensis[77]	Sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden	A member of the family Geisonoceratidae.
Sepia vandervoorti[87]	Sp. nov	Valid	Košťák, Jagt & Schlögl	Miocene (late Langhian–earliest Serravallian)	Köselerli Formation	Turkey	A species of Sepia.
Shanthaiceras[75]	Gen. et sp. nov	Valid	Niko, Sone & Leman	Ordovician	Lower Setul Limestone	Malaysia	A member of Pseudorthocerida. Genus includes new species S. amplum.
Sibumasuoceras kilimense[75]	Sp. nov	Valid	Niko, Sone & Leman	Ordovician	Lower Setul Limestone	Malaysia	A member of Pseudorthocerida.
Sibumasuoceras scrivenori[75]	Sp. nov	Valid	Niko, Sone & Leman	Ordovician	Lower Setul Limestone	Malaysia	A member of Pseudorthocerida.
Tafadnatoceras[88]	Gen. et sp. nov	In press	Ebbestad et al.	Late Ordovician	Upper Tiouririne Formation	Morocco	A member of Orthocerida belonging to the family Stereoplasmoceratidae. The type species is T. tiouririnense. Announced in 2019; the final version of the article naming it is not published yet.
Tofangoceras kedahense[75]	Sp. nov	Valid	Niko, Sone & Leman	Ordovician	Lower Setul Limestone	Malaysia	A member of Orthocerida.
Tofangoceras rayense[75]	Sp. nov	Valid	Niko, Sone & Leman	Ordovician	Lower Setul Limestone	Malaysia	A member of Orthocerida.
Tofangoceras staiti[75]	Sp. nov	Valid	Niko, Sone & Leman	Ordovician	Lower Setul Limestone	Malaysia	A member of Orthocerida.
Tyrionella[81]	Gen. et sp. nov	Valid	Fuchs et al.	Late Jurassic (Tithonian)	Altmühltal Formation	Germany	A member of Muensterelloidea. Genus includes new species T. fauseri.
Valkyrioceras[77]	Gen. et sp. nov	Valid	Kröger & Aubrechtová	Late Ordovician	Kullsberg Limestone Formation	Sweden	A member of the family Valcouroceratidae. Genus includes new species V. dalecarlia.
Wadema tattai[88]	Sp. nov	In press	Ebbestad et al.	Late Ordovician	Upper Ktaoua Formation	Morocco	A member of Actinocerida belonging to the family Wademidae. Announced in 2019; the final version of the article naming it is not published yet.

Gastropods

Research

• A study on the diversity of gastropod larval conch fossil assemblages of ages ranging from the Ordovician to Carboniferous is published online by Dzik (2019).^[89]
• A study on the recovery of gastropods in the aftermath of the Cretaceous–Paleogene extinction event, based on data from fossils from the former Ankerpoort-Curfs quarry (Geulhem, the Netherlands), is published online by Vellekoop et al. (2019).^[90]
• A probable cyclophoroidean gastropod, possibly a member of the family Cyclophoridae, is described from the Cretaceous amber from Myanmar by Xing et al. (2019), constituting the first confirmed and oldest record of soft-bodied preservation of a snail in Cretaceous amber.^[91]
• A study on the nature of the size dimorphism in the Miocene gastropod Persististrombus deperditus, and on the causes of its evolution, is published by Halder & Paira (2019).^[92]
• New specimens of Spinucella reimersi are described from the Miocene Gram Clay of Denmark by Schnetler (2019), who also provides an emended description of this species.^[93]
• A revision of extant and Neogene members of the genus Antistreptus from the southwestern Atlantic Ocean is published by Pastorino & Griffin (2019).^[94]

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Acirsa torensis[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark	A species of Acirsa.
Acmaea selandica[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark	A species of Acmaea.
Actinella fecundaerrata[96]	Nom. nov	Valid	Teixeira et al.	Quaternary		Madeira	A species of Actinella.
Agaronia almaghribensis[97]	Sp. nov	Valid	Pacaud & Lebrun	Eocene (Lutetian)		Morocco	A species of Agaronia.
Ageria laxa[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark
Ageria skeldervigensis[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark
Allmonia[98]	Gen. et comb. nov	Valid	Harzhauser & Landau	Oligocene (Rupelian) to Miocene (Tortonian)		Algeria  Austria  Belgium  Bosnia and Herzegovina  Bulgaria  Czech Republic  France  Germany  Greece  Hungary  India  Iran  Italy  Netherlands  Pakistan  Portugal  Slovakia  Slovenia   Switzerland  Turkey	A member of the family Turritellidae. The type species is "Turritella" cathedralis Brongniart (1823); genus also includes several European Cenozoic species previously placed in Protoma.
Allossospira[88]	Gen. et comb. nov	In press	Ebbestad et al.	Late Ordovician	Upper Ktaoua Formation	Morocco	A member of the family Gosseletinidae belonging to the subfamily Euryzoninae. The type species is "Lophospira" debganensis Horný (1997). Announced in 2019; the final version of the article naming it is not published yet.
Alvania amphitrite[99]	Sp. nov	Valid	Thivaiou, Harzhauser & Koskeridou	Miocene (Aquitanian)	Pentalofos Formation	Greece	A species of Alvania.
Amaea alicae[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark
Amaea stevniensis[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark
Amaurellina ravni[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark
Amaurotoma? multispirata[100]	Sp. nov	Valid	Ketwetsuriya, Cook & Nützel	Permian (Wordian)	Ratburi Group	Thailand	A member of the family Trochonematidae.
Ananias belaensis[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Eotomariidae.
Ananias bashkiriensis[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Eotomariidae.
Anematina indiscreta[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Elasmonematidae.
Aneudaronia[102]	Gen. et sp. nov	Valid	Guzhov	Jurassic		Russia	A member of the family Stuoraxidae. Genus includes new species A. elegans.
Anomphalus glaucus[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia
Apachella rugosa[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Eotomariidae.
Aphera chattica[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Aphera.
Aquitanobursa[104]	Gen. et comb. nov	Valid	Sanders, Merle & Puillandre	Oligocene (Rupelian) to early Pliocene		Dominican Republic  France  Italy  Pakistan  Venezuela  United States?	A member of the family Bursidae. The type species is "Ranella" grateloupi d'Orbigny (1852); genus also includes "Bursa" amphitrites Maury (1917), "Apollon" inaequicrenatus Cossmann & Peyrot (1924), "Ranella" morrisi d'Archiac & Haime (1853), "Ranella" tuberosa Grateloup (1833) and possibly also "Bursa (Bufonariella)" chipolana Schmelz (1997).
Archaeocyclotus[105]	Gen. et sp. nov	Valid	Asato & Hirano in Hirano et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Cyclophoridae. The type species is A. plicatula.	Archaeocyclotus plicatula
Archimediella carpathica[98]	Sp. nov	Valid	Harzhauser & Landau	Middle Miocene		Austria  Bosnia and Herzegovina  Hungary  Poland  Romania  Slovakia	A member of the family Turritellidae.
Arene jakobseni[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark	A species of Arene.
Argentisioliella[106]	Gen. et sp. nov	Valid	Miquel	Early Pliocene	Irene Formation	Argentina	A member of the family Cochliopidae. Genus includes new species A. pardignasi.
Arribazona seleukensis[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Orthonematidae.
Ascheria salina[107]	Sp. nov	Valid	Kiel et al.	Early Oligocene	Talara Basin	Peru	A member of the family Hokkaidoconchidae.
Ataphrus knoesensis[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark
Athleta benoisti[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Athleta.
Athleta confusus[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Athleta.
Athleta maculosus[103]	Sp. nov	Valid	Lozouet	Miocene		France	A species of Athleta.
Athleta memoirae[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Athleta.
Athleta peyreirensis[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Athleta.
Attiliosa gallica[108]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Attiliosa.
Attiliosa pouweri[108]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Attiliosa.
Auritoma[109]	Gen. et sp. nov	Valid	Szabó et al.	Jurassic		Italy	Genus includes new species A. lenticula.
Bandelzyga[101]	Gen. et sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Pseudozygopleuridae. Genus includes new species B. elegans.
Bathrotomaria jakobseni[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark
Bayania falconneti[110]	Sp. nov	Valid	Pacaud	Paleogene		France
Baylea longispira[101]	Sp. nov	Valid	Mazaev	Permian		China  Russia	A member of the family Lophospiridae.
Bellazona acuta[111]	Sp. nov	Valid	Mazaev	Early Permian		Russia
Bellerophon shakhtauensis[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia
Birgella burchi[112]	Sp. nov	Valid	Naranjo-García & Aguillón in Vega et al.	Late Cretaceous (late Campanian)	Cerro del Pueblo Formation	Mexico	A member of the family Hydrobiidae.
Bleytonella saalensis[113]	Sp. nov	Valid	Gründel, Keupp & Lang	Late Jurassic (Kimmeridgian)		Germany	A member of the family Pickworthiidae.
Bobellis[114]	Gen. et sp. nov	Valid	Peel	Early Silurian	Samuelsen Høj Formation	Greenland	A member of Euomphaloidea belonging to the family Pycnomphalidae. The type species is B. oliveri.
Bocourtia (Kuschelenia?) bonariensis[106]	Sp. nov	Valid	Miquel	Early Pliocene	Irene Formation	Argentina	A member of the family Bulimulidae.
Bolicharewia[101]	Gen. et sp. et comb. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Anomphalidae. Genus includes new species B. edita, as well as "Anomphalus" straparoliformis Mazaev (1997).
Bonellitia cryptica[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Bonellitia.
Bonellitia peyroti[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Bonellitia.
Boreomica costaspiralis[113]	Sp. nov	Valid	Gründel, Keupp & Lang	Late Jurassic (Kimmeridgian)		Germany	A member of Rissooidea belonging to the family Palaeorissoinidae.
Busambrella[109]	Gen. et sp. nov	Valid	Szabó et al.	Jurassic		Italy	Genus includes new species B. fasciata.
Buvignieria convexa[113]	Sp. nov	Valid	Gründel, Keupp & Lang	Late Jurassic (Kimmeridgian)		Germany	A member of Rissooidea belonging to the family Palaeorissoinidae.
Buvignieria racitana[113]	Sp. nov	Valid	Gründel, Keupp & Lang	Late Jurassic (Kimmeridgian)		Germany	A member of Rissooidea belonging to the family Palaeorissoinidae.
Callispira taosiformis[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Sinuspiridae.
Calyptraea baldii[115]	Sp. nov	Valid	Vicián, Kovács & Stein	Late Oligocene – early Miocene		Hungary	A species of Calyptraea.
Ceratia rodvigensis[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark	A species of Ceratia.
Cerion uva gouldi[116]	Subsp. nov	Valid	Harasewych	Pleistocene		Aruba	A subspecies of Cerion uva.
Cerithiella selandica[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark	A species of Cerithiella.
Cerithiopsis restemlongis[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark	A species of Cerithiopsis.
Chartronella belaensis[117]	Sp. nov	Valid	Szabó & Jaitly	Middle Jurassic (Callovian)	Chari Formation	India	A member of the family Paraturbinidae.
Chelotia pervicina[118]	Sp. nov	Valid	Berezovsky & Pacaud	Middle Eocene		Ukraine	A member of the family Pleurotomariidae.
Chlorozyga asiatica[100]	Sp. nov	Valid	Ketwetsuriya, Cook & Nützel	Permian (Wordian)	Ratburi Group	Thailand	A member of the family Imoglobidae.
Chriskingia[119]	Gen. et comb. nov	Valid	Janssen & Wilson	Miocene		Dominican Republic  Trinidad and Tobago	A member of the family Atlantidae; a new genus for "Atlanta" rotundata Gabb (1873).
Clavatula danuvii[115]	Sp. nov	Valid	Vicián, Kovács & Stein	Late Oligocene – early Miocene		Hungary	A species of Clavatula.
Clavilithes tamdakhtensis[97]	Nom. nov	Valid	Pacaud & Lebrun	Eocene (Lutetian)		Morocco	A species of Clavilithes.
Claviscala minor[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark
Cleopatra adami[120]	Nom. nov	Valid	Neiber & Glaubrecht	Early Pleistocene	Kaiso Series	Uganda	A species of Cleopatra; a replacement name for Viviparus cylindricus Adam (1957).
Clithon (Pictoneritina) coeuvrensis[121]	Sp. nov	Valid	Vrinat	Eocene (Ypresian)		France	A species of Clithon.
Clithon (Pictoneritina) occultatus[121]	Sp. nov	Valid	Vrinat	Eocene (Ypresian)		France	A species of Clithon.
Clithon (Pictoneritina) tigrinus[121]	Sp. nov	Valid	Vrinat	Eocene (Ypresian)		France	A species of Clithon.
Cochlis odovychenorum[122]	Sp. nov	Valid	Pedriali, Sosso & Dell'Angelo	Miocene		Ukraine	A member of the family Naticidae.
Cochlis ukrainensis[122]	Sp. nov	Valid	Pedriali, Sosso & Dell'Angelo	Miocene		Ukraine	A member of the family Naticidae.
Coninoda strekwera[113]	Sp. nov	Valid	Gründel, Keupp & Lang	Late Jurassic (Kimmeridgian)		Germany	A member of the family Eustomatidae.
Coniscala umbilica[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark
Contortia grateloupi[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A member of the family Cancellariidae.
Contortia intermedia[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A member of the family Cancellariidae.
Contortia zitae[115]	Sp. nov	Valid	Vicián, Kovács & Stein	Late Oligocene – early Miocene		Hungary	A member of the family Cancellariidae.
Coptostylus harzhauseri[110]	Nom. nov	Valid	Pacaud	Paleogene		France
Coroniopsis mortoni[123]	Sp. nov	Valid	Tracey & Craig in Tracey, Craig & Gain	Eocene (Lutetian)	Selsey Formation	United Kingdom	A member of the family Turridae.
Cosmocerithium veliger[124]	Sp. nov	Valid	Guzhov	Late Jurassic		Russia	A member of the family Cerithiopsidae.
Cossmannina alfischeri[125]	Sp. nov	Valid	Foster et al.	Early Triassic		China
Cranopsis cretacea[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark	A species of Cranopsis.
Cristalloella (Wonwalica) permica[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Tofanellidae.
Cryptaulax? parvum[113]	Sp. nov	Valid	Gründel, Keupp & Lang	Late Jurassic (Kimmeridgian)		Germany	A member of Cerithioidea belonging to the family Cryptaulacidae.
Cryptaulax? triangulare[113]	Sp. nov	Valid	Gründel, Keupp & Lang	Late Jurassic (Kimmeridgian)		Germany	A member of Cerithioidea belonging to the family Cryptaulacidae.
Cryptoptyxis rarenodosa[113]	Sp. nov	Valid	Gründel, Keupp & Lang	Late Jurassic (Kimmeridgian)		Germany	A member of the family Maoraxidae.
Cylichna stevniensis[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark	A species of Cylichna.
Cyllene satoi[126]	Sp. nov	Valid	Amano	Late Pliocene	Tentokuji Formation	Japan	A species of Cyllene.
Cypraeorbis nadaii[115]	Sp. nov	Valid	Vicián, Kovács & Stein	Late Oligocene – early Miocene		Hungary	A cowry.
Cystiscus pseudobrevis[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Cystiscus.
Dalliella elisae[110]	Sp. nov	Valid	Pacaud	Paleogene		France
Denticulofusus[127]	Gen. et comb. nov	Valid	Vermeij & Pacaud	Eocene (Lutetian and Priabonian)		France	A member of the family Columbariidae. The type species is "Fusus" gothicus Deshayes (1834); genus also includes "Coluzea" cavelieri Gain & Le Renard (2017) and "Coluzea" monicae Belliard, Gain & Le Renard (2017).
Dentimargo salvani[97]	Sp. nov	Valid	Pacaud & Lebrun	Eocene (Lutetian)		Morocco	A species of Dentimargo.
Dictyotomaria balabanovi[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Phymatopleuridae.
Discula cameroni[96]	Sp. nov	Valid	Teixeira et al.	Quaternary		Madeira	A species of Discula.
Discula polymorpha docaensis[96]	Subsp. nov	Valid	Teixeira et al.	Quaternary		Madeira
Doggerostra riedeli affinis[102]	Subsp. nov	Valid	Guzhov	Late Jurassic		Russia	A member of Heterobranchia of uncertain phylogenetic placement.
Dolicholatirus lorenzi[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark	A species of Dolicholatirus.
Donaldina erwini[125]	Sp. nov	Valid	Foster et al.	Early Triassic		China
Dragonia[124]	Gen. et 2 sp. nov	Valid	Guzhov	Late Jurassic		Russia	A member of the family Cerithiopsidae. Genus includes new species D. minuta and D. longa.
Eirlysia reticulatiformis[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Phymatopleuridae.
Elymicyclus[109]	Gen. et 4 sp. nov	Valid	Szabó et al.	Jurassic		Italy	Genus includes new species E. alternatus, E. ietumensis, E. martae and E. garibaldii.
Emarginula acutidens[109]	Sp. nov	Valid	Szabó et al.	Jurassic		Italy	A species of Emarginula.
Emarginula burgioi[109]	Sp. nov	Valid	Szabó et al.	Jurassic		Italy	A species of Emarginula.
Engina brunettii[128]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Engina.
Eoplatyzona ratchaburiensis[100]	Sp. nov	Valid	Ketwetsuriya, Cook & Nützel	Permian (Wordian)	Ratburi Group	Thailand	A member of the family Gosseletinidae.
Eopleurotoma bella[123]	Sp. nov	Valid	Tracey & Craig in Tracey, Craig & Gain	Eocene (Lutetian)		France	A species of Eopleurotoma.
Eopleurotoma bicincta[123]	Sp. nov	Valid	Tracey & Craig in Tracey, Craig & Gain	Eocene (Lutetian)		France	A species of Eopleurotoma.
Eopleurotoma boxleyi[123]	Sp. nov	Valid	Tracey & Craig in Tracey, Craig & Gain	Eocene (Lutetian)		France	A species of Eopleurotoma.
Eopleurotoma christinae[123]	Sp. nov	Valid	Tracey & Craig in Tracey, Craig & Gain	Eocene (Lutetian)		France	A species of Eopleurotoma.
Eopleurotoma hastula[123]	Sp. nov	Valid	Tracey & Craig in Tracey, Craig & Gain	Eocene (Lutetian)		France	A species of Eopleurotoma.
Eopleurotoma scitula[123]	Sp. nov	Valid	Tracey & Craig in Tracey, Craig & Gain	Eocene (Lutetian)		France	A species of Eopleurotoma.
Eopleurotoma vauvillensis[123]	Sp. nov	Valid	Tracey & Craig in Tracey, Craig & Gain	Eocene (Lutetian)		France	A species of Eopleurotoma.
Eovoluta adegokei[97]	Nom. nov	Valid	Pacaud & Lebrun	Eocene (Lutetian)		Morocco
Epalxis symondsi[123]	Sp. nov	Valid	Tracey & Craig in Tracey, Craig & Gain	Eocene (Lutetian)		France	A species of Epalxis.
Epalxis toddi[123]	Sp. nov	Valid	Tracey & Craig in Tracey, Craig & Gain	Eocene (Lutetian)		France	A species of Epalxis.
Episcomitra silvae[128]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Episcomitra.
Euthria presselierensis[128]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Euthria.
Exelissa indiana[117]	Sp. nov	Valid	Szabó & Jaitly	Middle Jurassic (Callovian)	Gadhada Formation	India	A member of the family Cryptaulacidae.
Epalxis? rosenkrantzi[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark	Transferred to the genus Fusulculus by Pacaud (2020), who coined a replacement name Fusulculus hanseni.[129]
Epetrium skeldervigensis[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark
Epitonium (Epitonium) zhuoi[130]	Sp. nov	Valid	Yu, Wang & Jarzembowski	Late Cretaceous (Cenomanian	Burmese amber	Myanmar	A gastropod of uncertain phylogenetic placement. Originally described as a species of Epitonium; Neubauer et al. (2019) transferred this species to the pupinid genus Pseudopomatias.[131]
Epitonium (Papyriscala) lyui[130]	Sp. nov	Valid	Yu, Wang & Jarzembowski	Late Cretaceous (Cenomanian	Burmese amber	Myanmar	A gastropod of uncertain phylogenetic placement. Originally described as a species of Epitonium; Neubauer et al. (2019) transferred this species to the pupinid genus Pseudopomatias.[131]
Euconospira skuini[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Phymatopleuridae.
Eucyclomphalus? marenostrum[109]	Sp. nov	Valid	Szabó et al.	Jurassic		Italy
Eucycloscala gracilicostatus[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark
Euspira jhuraensis[132]	Sp. nov	Valid	Das et al.	Uncertain; Late Jurassic[132] or Miocene[133]		India	A species of Euspira.
Euspira lakhaparensis[132]	Sp. nov	Valid	Das et al.	Uncertain; Late Jurassic[132] or Miocene[133]		India	A species of Euspira.
Euspira sirenkoi[122]	Sp. nov	Valid	Pedriali, Sosso & Dell'Angelo	Miocene		Ukraine	A species of Euspira.
Euthema hesoana[105]	Sp. nov	Valid	Asato & Hirano in Hirano et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Diplommatinidae.	Euthema hesoana
Falsobuvigna[113]	Gen. et sp. nov	Valid	Gründel, Keupp & Lang	Late Jurassic (Kimmeridgian)		Germany	A member of Rissooidea belonging to the new family Falsobuvignidae. The type species is F. peregrina.
Fischeriella sicula[109]	Sp. nov	Valid	Szabó et al.	Jurassic		Italy
Fissurella? stantoni[134]	Sp. nov	Valid	Powell & Geiger	Miocene	Topanga Canyon Formation	United States ( California)	A member of the family Fissurellidae.
Fusimorio salbriacensis[103]	Sp. nov	Valid	Lozouet	Miocene		France	A member of the family Cancellariidae.
Fusinus schnetleri[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark	A species of Fusinus.
Garviea craccoi[135]	Sp. nov	Valid	Zamberlan & Checchi	Eocene (late Ypresian/early Lutetian)		Italy	A member of the family Cypraeidae belonging to the subfamily Gisortiinae.
Gemmula garviei[123]	Sp. nov	Valid	Tracey & Craig in Tracey, Craig & Gain	Eocene (Lutetian)		France	A species of Gemmula.
Gemmula tuckeri[123]	Sp. nov	Valid	Tracey & Craig in Tracey, Craig & Gain	Eocene (Lutetian)		France	A species of Gemmula.
Gemmulopsis[123]	Gen. et sp. nov	Valid	Tracey & Craig in Tracey, Craig & Gain	Eocene (Lutetian)		France	A member of the family Turridae. The type species is G. nigellensis.
Gibberula amurakucensis[97]	Sp. nov	Valid	Pacaud & Lebrun	Eocene (Lutetian)		Morocco	A species of Gibberula.
Gibberula bezoyensis[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Gibberula.
Gibberula ligeriana[128]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Gibberula.
Gibberula pseudoaquitanica[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Gibberula.
Gibberula subovulatoides[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Gibberula.
Girtyspira oblonga[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Meekospiridae.
Globodoma magna[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Gosseletinidae. Originally described as a species of Globodoma, but subsequently transferred to the genus Nemaspira.[136]
Goniasma gerasimovi[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Goniasmatidae.
Goniobasis unilirata[112]	Sp. nov	Valid	Naranjo-García & Aguillón in Vega et al.	Late Cretaceous (late Campanian)	Cerro del Pueblo Formation	Mexico	A member of the family Pleuroceridae.
Gymnocerithium? convexoconcavum[113]	Sp. nov	Valid	Gründel, Keupp & Lang	Late Jurassic (Kimmeridgian)		Germany	A member of Campaniloidea belonging to the family Gymnocerithiidae.
Gyrodes mahalanobisi[132]	Sp. nov	Valid	Das et al.	Uncertain; Late Jurassic[132] or Miocene[133]		India	A member of the family Naticidae.
Haitia taylori[112]	Sp. nov	Valid	Naranjo-García & Aguillón in Vega et al.	Late Cretaceous (late Campanian)	Cerro del Pueblo Formation	Mexico	A member of the family Physidae.
Harmatia tokodensis[137]	Sp. nov	Valid	Kovács & Vicián	Eocene		Hungary	A member of the family Muricidae.
Hemicerithium perrettai[110]	Sp. nov	Valid	Pacaud	Eocene		United Kingdom
Hesperiella cyrtocostata[100]	Sp. nov	Valid	Ketwetsuriya, Cook & Nützel	Permian (Wordian)	Ratburi Group	Thailand	A member of the family Porcelliidae.
Heteronatica[102]	Gen. et sp. nov	Valid	Guzhov	Jurassic		Russia	A member of the family Cornirostridae. Genus includes new species H. globosa.
Hinea gaellae[110]	Sp. nov	Valid	Pacaud	Paleogene		France	A species of Hinea.
Hirsuticyclus[138]	Gen. et sp. nov	Valid	Neubauer, Xing & Jochum	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Cyclophoridae. The type species is H. electrum.
Holospira thompsoni[112]	Sp. nov	Valid	Naranjo-García & Aguillón in Vega et al.	Late Cretaceous (late Campanian)	Cerro del Pueblo Formation	Mexico	A species of Holospira.
Homalopoma acaste[99]	Sp. nov	Valid	Thivaiou, Harzhauser & Koskeridou	Miocene (Aquitanian)	Pentalofos Formation	Greece	A species of Homalopoma.
Hydrobia galatoniana[139]	Sp. nov	Valid	Esu & Girotti	Oligocene (Chattian)	Galatone Formation	Italy	A species of Hydrobia sensu lato.
Hydrobia ionica[139]	Sp. nov	Valid	Esu & Girotti	Oligocene (Chattian)	Galatone Formation	Italy	A species of Hydrobia sensu lato.
Hyphantozyga? khaophrikensis[100]	Sp. nov	Valid	Ketwetsuriya, Cook & Nützel	Permian (Wordian)	Ratburi Group	Thailand	A member of the family Pseudozygopleuridae.
Involuta coxi[140]	Sp. nov	Valid	Halder & Das	Eocene		India	A volute.
Jponsia aryanamensis[110]	Sp. nov	Valid	Pacaud			Iran
Jponsia yesimae[110]	Sp. nov	Valid	Pacaud	Paleogene		Turkey
Jumaramaria[117]	Gen. et sp. nov	Valid	Szabó & Jaitly	Middle Jurassic (Bathonian)	Jhurio Formation	India	A possible member of the family Pleurotomariidae. The type species is J. jumaraensis.
Jurassiscala[109]	Gen. et 2 sp. nov	Valid	Szabó et al.	Jurassic		Italy	Genus includes new species J. sturanii and possibly also J? tenuiretis.
Keilostoma nautagarona[110]	Sp. nov	Valid	Pacaud	Paleogene		France
Laevitomaria babalusciae[109]	Sp. nov	Valid	Szabó et al.	Jurassic		Italy
Lagocheilus cretaspira[105]	Sp. nov	Valid	Asato & Hirano in Hirano et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A species of Lagocheilus.	Lagocheilus cretaspira
Lagocheilus electrospira[105]	Sp. nov	Valid	Asato & Hirano in Hirano et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	Originally described as a species of Lagocheilus. Transferred to the genus Eotrichophorus by Balashov (2020).[141]	Lagocheilus electrospira
Leiostyla castanheiraensis[96]	Sp. nov	Valid	Teixeira et al.	Quaternary		Madeira	A species of Leiostyla.
Leiostyla cooki[96]	Sp. nov	Valid	Teixeira et al.	Quaternary		Madeira	A species of Leiostyla.
Leiostyla desertaensis[96]	Sp. nov	Valid	Teixeira et al.	Quaternary		Madeira	A species of Leiostyla.
Leiostyla simulans[96]	Sp. nov	Valid	Teixeira et al.	Quaternary		Madeira	A species of Leiostyla.
Leptaxis groviana antiqua[96]	Subsp. nov	Valid	Teixeira et al.	Quaternary		Madeira	A species of Leptaxis.
Leptaxis isambertoi[96]	Sp. nov	Valid	Teixeira et al.	Quaternary		Madeira	A species of Leptaxis.
Levizygopleura composita[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Pseudozygopleuridae.
Levizygopleura magna[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Pseudozygopleuridae.
Littoraria (Littorinopsis) massicardi[110]	Sp. nov	Valid	Pacaud	Paleogene		France	A species of Littoraria.
Lophospira latilabra[88]	Sp. nov	In press	Ebbestad et al.	Late Ordovician	Upper Ktaoua Formation	Morocco	A member of the family Lophospiridae. Announced in 2019; the final version of the article naming it is not published yet.
Lyria peyrehoradensis[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Lyria.
Maestratia[142]	Gen. et comb. nov	Valid	Dolin & Aguerre	Oligocene (Rupelian)		France	A member of the family Cypraeidae. The type species is "Pustularia (Conocypraea)" rugifera Schilder (1927).
Masaevia[102]	Gen. et sp. nov	Valid	Guzhov	Jurassic		Russia	A member of Heterobranchia of uncertain phylogenetic placement. Genus includes new species M. sinistra.
Mathilda dania[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark	A species of Mathilda.
Melanopsis neubaueri[110]	Sp. nov	Valid	Pacaud	Paleogene		France	A species of Melanopsis.
Melarhaphe perminima[110]	Sp. nov	Valid	Pacaud	Paleogene		France	A species of Melarhaphe.
Melarhaphe vanhyftei[110]	Sp. nov	Valid	Pacaud	Paleogene		France	A species of Melarhaphe.
Merica admirabilis[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Merica.
Merica estotiensis[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Merica.
Merica mutabilis[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Merica.
Mesalia bohnhavasae[98]	Sp. nov	Valid	Harzhauser & Landau	Middle Miocene		Hungary	A species of Mesalia.
Mesalia danica[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark	A species of Mesalia.
Mesalia sovisi[98]	Sp. nov	Valid	Harzhauser & Landau	Middle Miocene		Austria  Czech Republic	A species of Mesalia.
Mesalia stryriaca[98]	Sp. nov	Valid	Harzhauser & Landau	Middle Miocene		Austria	A species of Mesalia.
Mitrella clava[128]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Mitrella.
Mitrella ligeriana[128]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Mitrella.
Mitrella miopicta[128]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Mitrella.
Mitrella pseudoinedita[128]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Mitrella.
Mitrella pseudoblonga[128]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Mitrella.
Mitrella pseudoturgidula[128]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Mitrella.
Monimirus[101]	Gen. et sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Stuoraxidae. Genus includes new species M. bonus.
Monodonta joetsuensis[143]	Sp. nov	Valid	Amano	Late Pliocene	Nadachi Formation	Japan	A species of Monodonta.
Murexsul primigenius[144]	Sp. nov	Valid	Merle & Pacaud	Eocene (Priabonian)		France	A species of Murexsul.
Neverita amerrukensis[97]	Nom. nov	Valid	Pacaud & Lebrun	Eocene (Lutetian)		Morocco	A species of Neverita.
Nipteraxis parilis[97]	Sp. nov	Valid	Pacaud & Lebrun	Eocene (Lutetian)		Morocco
Novlepatella uralensis[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Daminilidae.
Ocinebrina houarti[108]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Ocinebrina.
Olssonia[104]	Gen. et comb. nov	Valid	Sanders, Merle & Puillandre	Eocene to early Miocene	Chira Formation Talara Formation	Peru	A member of the family Bursidae. The type species is "Bursa" chira Olsson (1930); genus also includes Bursa chira var. yasila Olsson (1930), raised to the rank of a separate species Olssonia yasila.
Opaliopsis carlsbergi[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark
Orecopia inopinata[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Omphalotrochidae.
Orthonema conspicuum[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Orthonematidae.
Orthonema nonnullum[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Orthonematidae.
Orthonema perspicuum[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Orthonematidae.
Orthonema repandum[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Orthonematidae.
Orthonychia korotkovae[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Platyceratidae.
Palaeoloxotoma[95]	Nom. nov	Valid	Hansen	Early-Middle Jurassic to Late Cretaceous (Maastrichtian)		Denmark  France  Jamaica  New Zealand  Poland	A member of the family Fissurellidae belonging to the subfamily Emarginulinae; a replacement name for Loxotoma Fischer (1885).
Palaeostylus confragosus[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Palaeostylidae.
Palaeostylus nervosus[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Palaeostylidae.
Payraudeautia ermesi[122]	Sp. nov	Valid	Pedriali, Sosso & Dell'Angelo	Miocene		Ukraine	A species of Payraudeautia.
Payraudeautia sabrinae[122]	Sp. nov	Valid	Pedriali, Sosso & Dell'Angelo	Miocene		Ukraine	A species of Payraudeautia.
Payraudeautia varovtsiana[122]	Sp. nov	Valid	Pedriali, Sosso & Dell'Angelo	Miocene		Ukraine	A species of Payraudeautia.
Paziella gallica[108]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Paziella.
Paziella gracilenta[108]	Sp. nov	Valid	Landau et al.	Miocene (Tortonian)		France	A species of Paziella.
Paziella modesta[115]	Sp. nov	Valid	Vicián, Kovács & Stein	Late Oligocene – early Miocene		Hungary	A species of Paziella.
Paziella parveenae[144]	Sp. nov	Valid	Merle & Pacaud	Eocene (Ypresian)		Pakistan	A species of Paziella.
Paziella zsoldosi[137]	Sp. nov	Valid	Kovács & Vicián	Eocene		Hungary	A species of Paziella.
Perakella[145]	Gen. et 2 sp. nov	Valid	Mazaev	Permian		Malaysia  Russia	Genus includes new species P. shakhtauensis and P. batteni.
Peruvispira uralensis[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Eotomariidae.
Phragmolites lissoni[146]	Sp. nov	Valid	Ebbestad & Gutiérrez-Marco	Ordovician (Sandbian)	Calapuja Formation	Peru
Pictavia lactera[113]	Sp. nov	Valid	Gründel, Keupp & Lang	Late Jurassic (Kimmeridgian)		Germany	A member of the family Ampullinidae.
Pisania redoniensis[128]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Pisania.
Planiturbo lerensis[117]	Sp. nov	Valid	Szabó & Jaitly	Middle Jurassic (Callovian)	Chari Formation	India	A member of the family Metriomphalidae.
Platyceras sagum[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia
Platyzona shikhanensis[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Pithodeidae.
Pleurocera gigantica[112]	Sp. nov	Valid	Naranjo-García & Aguillón in Vega et al.	Paleocene	Las Encinas Formation	Mexico	A species of Pleurocera.
Plocezyga boikoi[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Pseudozygopleuridae.
Plocezyga convexa[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Pseudozygopleuridae.
Plocezyga minuta[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Pseudozygopleuridae.
Plocezyga venusta[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Pseudozygopleuridae.
Polygona substrigosa[128]	Nom. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Polygona; a replacement name for Fusus strigosus Millet (1865).
Ponticypraea[147]	Gen. et sp. nov	Valid	Fehse et al.	Late Eocene		Australia	A member of the family Cypraeidae. Genus includes new species P. egregia.
Procerithium? inaequetuberculata[100]	Sp. nov	Valid	Ketwetsuriya, Cook & Nützel	Permian (Wordian)	Ratburi Group	Thailand	A member of the family Procerithiidae.
Propeucyclus[109]	Gen. et 3 sp. nov	Valid	Szabó et al.	Jurassic		Italy	Genus includes new species P. sicanus, P. obesus and possibly also P? semireticulatus.
Provalvata maior[148]	Sp. nov	Valid	Cataldo et al.	Early Cretaceous (Barremian)	Huitrín Formation	Argentina
Provalvata minor[148]	Sp. nov	Valid	Cataldo et al.	Early Cretaceous (Barremian)	Huitrín Formation	Argentina
Provanna pelada[107]	Sp. nov	Valid	Kiel et al.	Early Oligocene	Talara Basin	Peru	A species of Provanna.
Pseudocochlespira transversa[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark
Pseudolivella inadspecta[97]	Sp. nov	Valid	Pacaud & Lebrun	Eocene (Lutetian)		Morocco
Pseudonebularia sceauxensis[128]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Pseudonebularia.
Pseudotoma danica[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark
Pseudozygopleura applicata[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Pseudozygopleuridae.
Ptychidia austrorotundata[98]	Nom. nov	Valid	Harzhauser & Landau	Early Miocene		Austria  Slovakia	A member of the family Turritellidae; a replacement name for Turritella turris rotundata Schaffer (1912).
Ptychidia erynella[98]	Sp. nov	Valid	Harzhauser & Landau	Early and middle Miocene		Austria  Czech Republic  Hungary	A member of the family Turritellidae.
Pusia brebioni[128]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Pusia.
Pusia pseudoplicatula[128]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Pusia.
Pusia renauleauensis[128]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Pusia.
Pusia sublaevis[128]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Pusia.
Pyramistomia aliakmoni[99]	Sp. nov	Valid	Thivaiou, Harzhauser & Koskeridou	Miocene (Aquitanian)	Pentalofos Formation	Greece	A member of the family Pyramidellidae belonging to the subfamily Odostomiinae.
Pyrazopsis mexcala[110]	Sp. nov	Valid	Pacaud			Mexico
Pyrgotrochus vorosi[109]	Sp. nov	Valid	Szabó et al.	Jurassic		Italy
Pyropelta seca[107]	Sp. nov	Valid	Kiel et al.	Early Oligocene	Talara Basin	Peru	A species of Pyropelta.
Radvanospira[88]	Gen. et comb. et sp. nov	In press	Ebbestad et al.	Late Ordovician		Czech Republic  Morocco  Spain	A member of the family Platyceratidae. The type species is "Naticopsis" antiquata Barrande in Perner (1903); genus also includes new species R. baniensis. Announced in 2019; the final version of the article naming it is not published yet.
Ramusatomaria[109]	Gen. et sp. nov	Valid	Szabó et al.	Jurassic		Italy	Genus includes new species R. nuda.
Retimusina[109]	Gen. et 2 sp. nov	Valid	Szabó et al.	Jurassic		Italy	Genus includes new species R. poseidoni and possibly also R? tritoni.
Retispira khaophrikensis[100]	Sp. nov	Valid	Ketwetsuriya, Cook & Nützel	Permian (Wordian)	Ratburi Group	Thailand	A member of the family Bellerophontidae belonging to the subfamily Knightitinae.
Rimella aldjazairensis[110]	Sp. nov	Valid	Pacaud	Eocene		Algeria	A species of Rimella.
Rissoina albaresensis[110]	Sp. nov	Valid	Pacaud	Paleogene		France	A species of Rissoina.
Romanekia[142]	Gen. et comb. nov	Valid	Dolin & Aguerre	Eocene (Lutetian)		France	A member of the family Cypraeidae. The type species is "Proadusta" francki Gain, Le Renard & Belliard (2012).
Rotfanella gerasimovi[102]	Sp. nov	Valid	Guzhov	Jurassic		Russia	A member of the family Cimidae.
Rotfanella reticulata[102]	Sp. nov	Valid	Guzhov	Jurassic		Russia	A member of the family Cimidae.
Rugosacyclus[113]	Gen. et comb. nov	Valid	Gründel, Keupp & Lang	Late Jurassic (Kimmeridgian)		Germany	A member of Caenogastropoda of uncertain phylogenetic placement. The type species is "Spirocyclina" rugosa Brösamlen (1909).
Saalensia[113]	Gen. et sp. nov	Valid	Gründel, Keupp & Lang	Late Jurassic (Kimmeridgian)		Germany	A member of Sorbeoconcha belonging to the family Brachytrematidae. The type species is S. birugata.
Sallya bimaris[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Pseudophoridae.
Sallya indigena[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Pseudophoridae.
Scelidotoma aldersoni[134]	Sp. nov	Valid	Powell & Geiger	Miocene	Topanga Canyon Formation	United States ( California)	A species of Scelidotoma.
Schistoloma electrothauma[105]	Sp. nov	Valid	Asato & Hirano in Hirano et al.	Late Cretaceous (Cenomanian)	Burmese amber	Myanmar	A member of the family Pupinidae.	Schistoloma electrothauma
Seleukella[101]	Gen. et sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Palaeozygopleuridae. Genus includes new species S. costata.
Semisulcospira (Biwamelania) gamoensis[149]	Sp. nov	Valid	Matsuoka & Miura	Early Pleistocene	Gamo Formation	Japan	A species of Semisulcospira.
Semisulcospira (Biwamelania) nojirina[149]	Sp. nov	Valid	Matsuoka & Miura	Late Pliocene	Ayama Formation Koka Formation	Japan	A species of Semisulcospira.
Semisulcospira (Biwamelania) reticulataformis[149]	Sp. nov	Valid	Matsuoka & Miura	Late Pliocene	Ayama Formation	Japan	A species of Semisulcospira.
Semisulcospira (Biwamelania) tagaensis[149]	Sp. nov	Valid	Matsuoka & Miura	Early Pleistocene	Gamo Formation	Japan	A species of Semisulcospira.
Shurovites robustus[113]	Sp. nov	Valid	Gründel, Keupp & Lang	Late Jurassic (Kimmeridgian)		Germany	A member of Cerithioidea belonging to the family Cryptaulacidae.
Shwedagonia korolukae[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Phymatopleuridae.
Shwedagonia pagoda[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Phymatopleuridae.
Sigmesalia ganntourensis[97]	Sp. nov	Valid	Pacaud & Lebrun	Eocene (Lutetian)		Morocco
Sigmesalia imitatrix[97]	Sp. nov	Valid	Pacaud & Lebrun	Late Cretaceous (Maastrichtian)		Madagascar
Siphonochelus tambacounda[144]	Sp. nov	Valid	Merle & Pacaud	Eocene (Lutetian)		Senegal	A species of Siphonochelus.
Solarioconulus kachchhensis[117]	Sp. nov	Valid	Szabó & Jaitly	Middle Jurassic (Callovian)	Chari Formation	India	A member of the family Ataphridae.
Spiroscala elegans[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Phymatopleuridae.
Stegocoelia abscisa[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Orthonematidae.
Stegocoelia arta[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Orthonematidae.
Stegocoelia centrosinuata[100]	Sp. nov	Valid	Ketwetsuriya, Cook & Nützel	Permian (Wordian)	Ratburi Group	Thailand	A member of the family Goniasmatidae.
Stenozone spiralis[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Eotomariidae.
Stephanozyga seminuda[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Pseudozygopleuridae.
Streptacis nalivkini[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Streptacididae.
Strobeus bashkiriensi[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Soleniscidae.
Strophostylus fortuitus[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Platyceratidae.
Stuoraxis crassa[102]	Sp. nov	Valid	Guzhov	Jurassic		Russia	A member of the family Stuoraxidae.
Sulcoactaeon? haboensis[117]	Sp. nov	Valid	Szabó & Jaitly	Middle Jurassic (Bathonian)	Patcham Formation	India	A member of the family Aplustridae.
Sulcomitrella sceauxensis[128]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Sulcomitrella.
Sveltia castellum[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Sveltia.
Sveltia ruginosa[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Sveltia.
Sveltia varonei[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Sveltia.
Tapinotomaria cancellata[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Portlockiellidae.
Tatara pseudosassia[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark
Tectonatica anistratenkorum[122]	Sp. nov	Valid	Pedriali, Sosso & Dell'Angelo	Miocene		Ukraine	A species of Tectonatica.
Tectonatica pseudoprietoi[122]	Sp. nov	Valid	Pedriali, Sosso & Dell'Angelo	Miocene		Ukraine	A species of Tectonatica.
Tectus extraconicus[150]	Sp. nov	Valid	Gain, Belliard & Le Renard	Eocene		France	A species of Tectus.
Tectus fluctus[150]	Sp. nov	Valid	Gain, Belliard & Le Renard	Eocene		France	A species of Tectus.
Tectus fredevillensis[150]	Sp. nov	Valid	Gain, Belliard & Le Renard	Eocene		France	A species of Tectus.
Tectus gervillii[150]	Sp. nov	Valid	Gain, Belliard & Le Renard	Eocene		France	A species of Tectus.
Tectus? indecorus[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark	Possibly a species of Tectus.
Tectus leae[150]	Sp. nov	Valid	Gain, Belliard & Le Renard	Eocene		France	A species of Tectus.
Tectus planibasis[150]	Sp. nov	Valid	Gain, Belliard & Le Renard	Eocene		France	A species of Tectus.
Tectus richardi[150]	Sp. nov	Valid	Gain, Belliard & Le Renard	Eocene		France	A species of Tectus.
Tectus spinalis[150]	Sp. nov	Valid	Gain, Belliard & Le Renard	Eocene		France	A species of Tectus.
Tectus torticostulus[150]	Sp. nov	Valid	Gain, Belliard & Le Renard	Eocene		France	A species of Tectus.
Temnotropis stevniensis[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark
Termihabena[151]	Gen. et sp. et comb. nov	Valid	Mazaev	Early Permian		Russia	Genus includes new species T. lirata, as well as Т. nikitowkensis (Yakowlew, 1899), T. asiatica (Licharew, 1967), T. millegranosa (Girty, 1934) and T. pinegensis (Mazaev, 2006).
Thereitis carlsbergi[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark
Thereitis cretacea[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark
Thylacodes contendis[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark	A species of Thylacodes.
Toronyella[109]	Gen. et 2 sp. nov	Valid	Szabó et al.	Jurassic		Italy	Genus includes new species T. lineata and T. margaritata.
Torquesia (Ispharina) atax[110]	Nom. nov	Valid	Pacaud	Paleogene		France
Trachydomia shkurkoi[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Trachyspiridae.
Trapanimaria[109]	Gen. et 3 sp. nov	Valid	Szabó et al.	Jurassic		Italy	Genus includes new species T. gattoi, T. nicolosiensis and possibly also T? pallinii.
Tritia albrechti[152]	Sp. nov	Valid	Stein	Miocene		North Sea basin	A species of Tritia.
Tritia girondica kautskyi[152]	Subsp. nov	Valid	Stein	Miocene		North Sea basin
Tritia hemmoorica[152]	Sp. nov	Valid	Stein	Miocene		North Sea basin	A species of Tritia.
Tritia jyllandica[152]	Sp. nov	Valid	Stein	Miocene		North Sea basin	A species of Tritia.
Tritia karinwienrichae[152]	Nom. nov	Valid	Stein	Miocene		Germany	A species of Tritia; a replacement name for Nassarius karinae Wienrich (2001).
Tritia kostejana krocki[152]	Subsp. nov	Valid	Stein	Miocene		North Sea basin
Tritia mostafavii[152]	Nom. nov	Valid	Stein	Miocene		Germany	A species of Tritia; a replacement name for Hinia coronata Mostafavi (1978).
Tritia mothsi[152]	Sp. nov	Valid	Stein	Miocene		North Sea basin	A species of Tritia.
Tritia sorgenfreii[152]	Sp. nov	Valid	Stein	Miocene		North Sea basin	A species of Tritia.
Tritia turtaudierei[128]	Sp. nov	Valid	Landau, Ceulemans & Van Dingenen	Miocene (Tortonian)		France	A species of Tritia.
Tritia westfalica[152]	Sp. nov	Valid	Stein	Miocene		North Sea basin	A species of Tritia.
Tritonophon grandis[88]	Sp. nov	In press	Ebbestad et al.	Late Ordovician	Lower Second Bani Formation	Morocco	A member of the family Bellerophontidae. Announced in 2019; the final version of the article naming it is not published yet.
Trochotomaria conoidea[109]	Sp. nov	Valid	Szabó et al.	Jurassic		Italy
Trochotomaria polymorpha[109]	Sp. nov	Valid	Szabó et al.	Jurassic		Italy
Tropacerithium[113]	Gen. et 2 sp. nov	Valid	Gründel, Keupp & Lang	Late Jurassic (Kimmeridgian)		Germany	A member of Cerithioidea possibly belonging to the family Cryptaulacidae. The type species is T. cumaritum; genus also includes T. danubii.
Tychonia fortis[101]	Sp. nov	Valid	Mazaev	Early Permian		Russia	A member of the family Anomphalidae.
Typhina cryptica[144]	Sp. nov	Valid	Merle & Pacaud	Eocene (Ypresian)		France	A species of Typhina.
Typhina radulfiensis[144]	Sp. nov	Valid	Merle & Pacaud	Eocene (Priabonian)		France	A species of Typhina.
Unitas aequalis[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A member of the family Cancellariidae.
Unitas aquitanica[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A member of the family Cancellariidae.
Unitas arenosa[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A member of the family Cancellariidae.
Unitas decorticata[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A member of the family Cancellariidae.
Unitas gradata[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A member of the family Cancellariidae.
Unitas heinbergi[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark
Unitas sanctistephani[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A member of the family Cancellariidae.
Unitas substephanensis[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A member of the family Cancellariidae.
Urlocella undulata[102]	Sp. nov	Valid	Guzhov	Jurassic		Russia	A member of the family Cimidae.
Unzhispira[102]	Gen. et sp. nov	Valid	Guzhov	Jurassic		Russia	A member of the family Cimidae. Genus includes new species U. minuta.
Vatopsis heinbergi[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark
Vermeijia[126]	Gen. et sp. nov	Valid	Amano	Late Pliocene	Kuwae Formation	Japan	A member of the family Capulidae. Genus includes new species V. japonica.
Vernelia samae[125]	Sp. nov	Valid	Foster et al.	Early Triassic		China
Vexillum? rosenkrantzi[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark	Possibly a species of Vexillum.
Vexillum tenestolidum[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark	A species of Vexillum.
Vicinocerithium vynckei[110]	Nom. nov	Valid	Pacaud	Paleogene		France
Vicnigoria[101]	Gen. et sp. nov	Valid	Mazaev	Early Permian		Russia	Probably a member of the family Trochonematidae. Genus includes new species V. monstrata.
Viennella[98]	Gen. et comb. et sp. nov	Valid	Harzhauser & Landau	Middle Miocene		Austria  Hungary	A member of the family Turritellidae. The type species is "Turritella" incisaeformis Csepreghy-Meznerics (1956); genus also includes new species V. ignorata.
Viviparus riegensis[110]	Nom. nov	Valid	Pacaud	Paleogene		France	A species of Viviparus.
Volvarina sibuzatiana[103]	Sp. nov	Valid	Lozouet	Oligocene (Chattian)		France	A species of Volvarina.
Wateletia custodia[110]	Nom. nov	Valid	Pacaud	Paleogene		France
Worthenia? waterhouse[100]	Sp. nov	Valid	Ketwetsuriya, Cook & Nützel	Permian (Wordian)	Ratburi Group	Thailand	A member of the family Lophospiridae.
Zarnglaffia palermitana[109]	Sp. nov	Valid	Szabó et al.	Jurassic		Italy
Zarnglaffia polygonalis[109]	Sp. nov	Valid	Szabó et al.	Jurassic		Italy
Zebinella selandica[95]	Sp. nov	Valid	Hansen	Cretaceous-Paleogene boundary		Denmark	A species of Zebinella.
Zizipupa[102]	Gen. et sp. nov	Valid	Guzhov	Jurassic		Russia	A member of the family Ampezzanildidae. Genus includes new species Z. costata.

Other molluscs

Research

• A study on shell microstructures of specimens of Anabarella australis from the Cambrian Series 2 Xinji Formation of the North China Block is published by Li et al. (2019).^[153]
• A study on the relative importance of bivalves and brachiopods in the fossil assemblages from the Carboniferous Pennsylvanian Breathitt Formation of Kentucky is published by Hsieh, Bush & Bennington (2019).^[154]
• A study on transitional Permian-Triassic bivalve fauna from five sections of littoral clastic facies in southwestern China is published by Song et al. (2019).^[155]
• A study on body size changes in bivalves in the aftermath of the Permian–Triassic extinction event is published online by Atkinson & Wignall (2019).^[156]
• A study on changes of body size of bivalves belonging to the family Limidae prior to, and in the aftermath of the Triassic–Jurassic extinction event is published by Atkinson et al. (2019).^[157]
• A study on changes in the body size of benthic marine bivalves and brachiopods from the Lusitanian Basin (Portugal) before the Toarcian oceanic anoxic event is published by Piazza et al. (2019).^[158]
• A study on the impact of the early Toarcian extinction event on fossil brachiopods and bivalves known from the Iberian Range (Spain) is published by Danise et al. (2019).^[159]
• A study on the impact of the Early Toarcian Oceanic Anoxic Event on the population of the bivalve species Posidonotis cancellata from the Neuquén Basin (Argentina) is published by Franch et al. (2019).^[160]
• A study on the temporal and spatial distribution of pectinid and ostreid bivalves in Middle and Late Jurassic sequences exposed in the Tanggula Mountains (China) is published by Sha (2019).^[161]
• A study on muscle attachment sites preserved in phosphatized inoceramid Gnesioceramus anglicus, and on their implications for inferring the palaeobiology of inoceramid bivalves, is published by Knight & Morris (2019).^[162]
• A sclerochronological study of shells of the gigantic inoceramids Sphenoceramus schmidti and S. sachalinensis from the middle Campanian Yezo Basin in Hokkaido (Japan) is published by Walliser et al. (2019).^[163]
• A study on the relationship of growth rate in the scallop Carolinapecten eboreus to temperature and primary production, and on its implications for inferring the cause of extinction of this species, is published by Johnson et al. (2019).^[164]

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Acharax svalbardensis[165]	Sp. nov	Valid	Hansen et al.	Late Pleistocene		Fram Strait	A bivalve belonging to the family Solemyidae.
Atomodesma? hautmanni[125]	Sp. nov	Valid	Foster et al.	Early Triassic		China	A bivalve.
Atrina multicostata[166]	Sp. nov	Valid	Friesenbichler et al.	Middle Triassic (Anisian)	Caerace Formation	Romania	A bivalve, a species of Atrina.
Baxoitrigonia[167]	Gen. et sp. nov		Stiller & Chen	Late Jurassic (Tithonian)	Lagongtang Formation	China	A bivalve. Genus includes new species B. baxoiensis.
Bengtsonella[168]	Gen. et comb. nov	Junior homonym	Cooper & Leanza	Early Cretaceous (Albian)	Riachuelo Formation	Brazil	A bivalve belonging to the family Megatrigoniidae and the subfamily Megatrigoniinae. The type species is "Anditrigonia" britoi Hessel (2005). The generic name is preoccupied by Bengtsonella Muller & Hinz (1991) and Bengtsonella Mostler (1996); Ceccolini & Cianferoni (2021) coined a replacement name Bengtsonigonia.[169]
Californigonia[168]	Gen. et comb. nov	Valid	Cooper & Leanza	Jurassic (Callovian–early Oxfordian)	Bicknell Sandstone	Canada ( British Columbia)  United States ( California)	A bivalve belonging to the subfamily Megatrigoniinae. The type species is "Trigonia" plumasensis Hyatt (1892).
Callocardia (Nitidavenus) youssoufia[97]	Sp. nov	Valid	Pacaud & Lebrun	Eocene (Lutetian)		Morocco	A bivalve.
Callucina itaporangensis[170]	Sp. nov	Valid	Ayoub-Hannaa et al.	Cretaceous	Cotinguiba Formation	Brazil  United Kingdom	A bivalve belonging to the family Lucinidae.
Cardiocardita peregrina[97]	Sp. nov	Valid	Pacaud & Lebrun	Eocene (Lutetian)		Morocco	A bivalve.
Carditella pitufina[171]	Sp. nov	Valid	Pérez	Early Pliocene		Argentina	A bivalve belonging to the family Condylocardiidae.
Chlamys (Praechlamys) prima[166]	Sp. nov	Valid	Friesenbichler et al.	Middle Triassic (Anisian)	Caerace Formation	Romania	A bivalve, a species of Chlamys.
Ckaraosippur[172]	Gen. et comb. et sp. nov	Valid	Santelli & del Río	Miocene–Pliocene	Bahía Inglesa Formation Camarones Formation Navidad Formation	Argentina  Chile	A scallop. The type specie is "Pecten" calderensis Möricke (1896); genus also includes new species C. camachoi.
Conchocele yatsuoensis[173]	Sp. nov	Valid	Amano et al.	Early Miocene	Kurosedani Formation	Japan	A bivalve.
Costacallista discrepans[97]	Sp. nov	Valid	Pacaud & Lebrun	Eocene (Lutetian)		Morocco	A bivalve.
Craginella[168]	Gen. et comb. nov	Valid	Cooper & Leanza	Late Jurassic (Kimmeridgian)	Malone Formation	Mexico  United States ( Louisiana  Texas)	A bivalve belonging to the subfamily Megatrigoniinae. The type species is "Trigonia" goodellii Cragin (1897).
Damborenella[168]	Gen. et comb. nov	Valid	Cooper & Leanza	Late Jurassic (Tithonian)		Argentina  Chile  Colombia  Peru	A bivalve belonging to the subfamily Megatrigoniinae. The type species is "Trigonia" eximia Philippi (1899); genus also includes "Trigonia" discors Philippi (1899), "Anditrigonia" lamberti Levy (1967) and "Anditrigonia eximia" tesselicaudata Leanza (1993) (raised to the rank of a separate species Damborenella tessellicaudata).
Dentalium mcganna[174]	Sp. nov	Valid	Powell, Clites & Poust	Miocene	Sobrante Formation	United States ( California)	A species of Dentalium.
Dietotenhosen[172]	Gen. et comb. nov	Valid	Santelli & del Río	Miocene–Pliocene	Bahía Inglesa Formation Coquimbo Formation Horcón Formation Hornillos Formation Huaricangana Formation La Portada Formation Pisco Formation Taime Formation	Chile  Peru	A scallop. Genus includes D. hupeanus (Philippi, 1887) and D. remondi (Philippi, 1887).
Dosinia (Dosinisca) vredenburgi[175]	Sp. nov	Valid	Borkar & Kulkarni	Miocene	Chhasra Formation Khari Nadi Formation Lower Mekran Series	India  Pakistan	A bivalve belonging to the family Veneridae; a species of Dosinia.
Dosinia (Dosinisca) wynnei[175]	Sp. nov	Valid	Borkar & Kulkarni	Miocene	Chhasra Formation Khari Nadi Formation Lower Mekran Series	India  Pakistan	A bivalve belonging to the family Veneridae; a species of Dosinia.
Entolium reticulatum[166]	Sp. nov	Valid	Friesenbichler et al.	Middle Triassic (Anisian)	Caerace Formation	Romania	A bivalve.
Eosphaera[176]	Gen. et comb. nov	Valid	Fürsich, Heinze & Ayoub-Hannaa	Jurassic (Callovian to Kimmeridgian)		Egypt  India  Madagascar  Portugal	A bivalve belonging to the family Lucinidae. The type species is "Sphaera" madagascariensis Newton (1889).
Chaetopleura abbessi[177]	Sp. nov	Valid	Cherns & Schwabe	Oligocene (Chattian) and Miocene (Burdigalian)[178]		France	A chiton. Originally described as a species of Chaetopleura; subsequently transferred to the genus Ischnochiton by Dell'Angelo et al. (2018).[178]
Chaetopleura gaasi[177]	Sp. nov	Valid	Cherns & Schwabe	Oligocene (Rupelian)		France	A chiton, Originally described as a species of Chaetopleura; subsequently transferred to the new genus Spinochiton by Dell'Angelo et al. (2018).[178]
Helcionella lemdadensis[179]	Sp. nov	Valid	Geyer, Valent & Meier	Cambrian	Tannenknock Formation	Germany
Hoernesia? danisa[125]	Sp. nov	Valid	Foster et al.	Early Triassic		China	A bivalve.
Homopleura[180]	Gen. et comb. et 2 sp. nov	Valid	Masse & Fenerci-Masse	Cretaceous		Bulgaria  France	A rudist bivalve. The type species is "Monopleura" affinis Matheron (1878); genus also includes "Monopleura" imbricata Matheron (1842) and "Monopleura" coquandi Matheron (1878), as well as new species H. rustrella and H. balkanica.
Inoceramus chiplonkari[30]	Sp. nov	Valid	Walaszczyk in Gale, Kennedy & Walaszczyk	Late Cretaceous (Cenomanian and Turonian)	Karai Formation	India  Japan
Ischnochiton crovatoi[181]	Sp. nov	Valid	Dell'Angelo, Sosso & Bonfitto	Pleistocene		Italy	A chiton, a species of Ischnochiton.
Ischnochiton fehsei[177]	Sp. nov	Valid	Cherns & Schwabe	Eocene		France	A chiton, a species of Ischnochiton.
Lasaea firma[182]	Sp. nov	Valid	Lesport & Lozouet in Lesport et al.	Miocene (Aquitanian)		France	A bivalve belonging to the family Lasaeidae.
Leptostega frankenwaldensis[179]	Sp. nov	Valid	Geyer, Valent & Meier	Cambrian	Tannenknock Formation	Germany	A helcionelloid.
Neilo altamirano[107]	Sp. nov	Valid	Kiel et al.	Early Oligocene	Talara Basin	Peru	A bivalve belonging to the family Malletiidae.
Neocrassina (Pruvostiella) madagascariensis[176]	Sp. nov	Valid	Fürsich, Heinze & Ayoub-Hannaa	Late Jurassic (Oxfordian)	Morondava Basin	Madagascar	A bivalve belonging to the family Astartidae.
Palaeonucula feruglioi[183]	Sp. nov	Valid	Damborenea & Pagani	Early Jurassic		Argentina	A bivalve.
Parailsanella luonanensis[184]	Sp. nov	Valid	Li et al.	Cambrian Series 2	Xinji Formation	China	A helcionellid.
Peronaea basteroti[182]	Sp. nov	Valid	Lesport, Lozouet & Pacaud in Lesport et al.	Miocene (Aquitanian)		France	A bivalve, a species of Peronaea.
Phaenodesmia? piatnitzkyi[183]	Sp. nov	Valid	Damborenea & Pagani	Early Jurassic		Argentina	A bivalve.
Pinna simionescui[166]	Sp. nov	Valid	Friesenbichler et al.	Middle Triassic (Anisian)	Caerace Formation	Romania	A bivalve, a species of Pinna.
Platymyoidea morrisi[185]	Sp. nov	Valid	Hodges	Early Jurassic		United Kingdom	A bivalve.
Pleurophopsis matsumotoi[186]	Sp. nov	Valid	Amano et al.	Late Oligocene to early Miocene	Nabae Group	Japan	A bivalve belonging to the family Vesicomyidae.
Pleurophopsis talarensis[107]	Sp. nov	Valid	Kiel et al.	Early Oligocene	Talara Basin	Peru	A bivalve belonging to the family Vesicomyidae.
Praedicerocardium[166]	Gen. et sp. nov	Valid	Friesenbichler et al.	Middle Triassic (Anisian)	Caerace Formation	Romania	A bivalve. Genus includes new species P. vetulus.
Praenucula pojetai[88]	Sp. nov	In press	Ebbestad et al.	Late Ordovician		Morocco	A bivalve belonging to the family Praenuculidae. Announced in 2019; the final version of the article naming it is not published yet.
Procardia inouei[187]	Sp. nov	Valid	Amano	Paleocene	Katsuhira Formation	Japan	A bivalve belonging to the group Anomalodesmata and to the family Parilimyidae.
Pseudophopsis[107]	Gen. et comb. nov	Valid	Kiel et al.	Eocene and Early Oligocene	Talara Basin	Cuba  Peru	A bivalve belonging to the group Heterodonta and to the family Kalenteridae. The type species is "Pleurophopsis" peruviana Olsson (1931); genus also includes "Unio" bitumen Cooke (1919).
Quenstedtia madagascariensis[176]	Sp. nov	Valid	Fürsich, Heinze & Ayoub-Hannaa	Late Jurassic (Oxfordian)	Morondava Basin	Madagascar	A bivalve belonging to the family Quenstedtiidae.
Ramentoides xixiangensis[188]	Sp. nov	Valid	Qin et al.	Cambrian		China	A member of Maikhanellidae.
Romaniamya[166]	Gen. et sp. nov	Valid	Friesenbichler et al.	Middle Triassic (Anisian)	Caerace Formation	Romania	A bivalve. Genus includes new species R. mahmudiaensis.
Rutitrigonia cintarojensis[148]	Sp. nov	Valid	Cataldo et al.	Early Cretaceous (Barremian)	Huitrín Formation	Argentina	A bivalve.
Ryderia tehuelchana[183]	Sp. nov	Valid	Damborenea & Pagani	Early Jurassic		Argentina	A bivalve.
Scythentolium anisicum[166]	Sp. nov	Valid	Friesenbichler et al.	Middle Triassic (Anisian)	Caerace Formation	Romania	A bivalve.
Sogdianagonia[189]	Nom. nov	Valid	Schneider & Kelly	Late Cretaceous (Cenomanian to Turonian)		Kyrgyzstan  Uzbekistan	A bivalve belonging to the family Megatrigoniidae; a replacement name for Turkestanella Tashiro (1979). The type species is "Trigonia" turkestanensis Arkhangelsky (1916).
Solemya lucifuga[190]	Sp. nov	Valid	Pérez-Barría & Nielsen	Early Miocene	Ranquil Formation	Chile	A bivalve, a species of Solemya.
Stenoplax monila[177]	Sp. nov	Disputed	Cherns & Schwabe	Oligocene (Rupelian)		France	A chiton. Originally described as a species of Stenoplax; Dell'Angelo et al. (2020) considered this species to be a junior synonym of Lepidochitona oligocaena.[191]
Tindaria kretensis[192]	Sp. nov	Valid	Koskeridou, La Perna & Giamali	Early Pliocene		Greece	A bivalve.
Tonicella lira[177]	Sp. nov	Valid	Cherns & Schwabe	Oligocene (Chattian)		France	A chiton, a species of Tonicella.
Vaccinites alceotarlaoi[193]	Sp. nov	Valid	Pons et al.	Late Cretaceous (early Campanian)		Serbia  Spain	A rudist bivalve.
Vaccinites bilottei[193]	Sp. nov	Valid	Pons et al.	Late Cretaceous (late Campanian)		Bulgaria	A rudist bivalve.
Vaccinites pirenaicus[193]	Sp. nov	Valid	Pons et al.	Late Cretaceous (late Campanian)		Spain	A rudist bivalve.
Wareniconcha mercenarioides[194]	Sp. nov	Valid	Kase et al.	Pliocene		Philippines	A bivalve belonging to the family Vesicomyidae.
Xylophagella littlei[195]	Sp. nov	Valid	Hryniewicz in Hryniewicz et al.	Late Paleocene	Basilika Formation	Norway	A bivalve belonging to the family Pholadidae.
Yoldiella spitsbergensis[195]	Sp. nov	Valid	Amano in Hryniewicz et al.	Late Paleocene	Basilika Formation	Norway	A bivalve belonging to the family Yoldiidae.

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