扇羽鰓屬

扇羽鰓屬（學名：Flabellina），又名扇鰓海蛞蝓屬^[4]，是裸鰓目枝鰓亞目之下的一個海蛞蝓的屬、海洋腹足綱軟體動物扇羽鰓科的成員^[3]。

扇羽鰓屬
Flabellina affinis
科學分類
界：	動物界 Animalia
門：	軟體動物門 Mollusca
綱：	腹足綱 Gastropoda
亞群：	裸側亞群 Nudipleura
目：	裸鰓目 Nudibranchia
亞目：	枝鰓亞目 Cladobranchia
總科：	扇羽鰓總科 Flabellinoidea
科：	扇羽鰓科 Flabellinidae
屬：	扇羽鰓屬 Flabellina Gray [in Griffith & Pidgeon], 1833[1]
模式種
Doris affinis Gmelin, 1791[2]

分類歷史

扇羽鰓屬由Gray於1850年建立，模式種為Flabellina affinis and characterised by the cerata（英語：cerata） being arranged on peduncles and the rhinophore（英語：rhinophore）s being annulate. Many species were added by subsequent authors. In 1981 the genus Coryphella（英語：Coryphella） was merged under the older name Flabellina as despite a large range of morphological characters in the 64 species of Flabellinidae and Coryphellidae（英語：Coryphellidae） known at that time, no clear distinction could be found to separate the species into the two genera.^[5] In 2017 an integrative study of the 71 species then considered to be included in Flabellina redistributed the species into seven families and 26 genera.^[6]

物種

根據WoRMS，本屬包括下列各物種^[3]：

• Flabellina affinis (Gmelin, 1791)

Species retained in Flabellina until new investigations place them correctly, currently Family Flabellinidae incertae sedis:

• Flabellina alternata Ortea & Espinosa, 1998
• Flabellina bertschi Gosliner & Kuzirian, 1990
• Flabellina bulbosa Ortea & Espinosa, 1998
• Flabellina dana Millen & Hamann, 2006
• Flabellina dushia (Marcus Ev. & Er., 1963)
• Flabellina engeli Ev. Marcus & Er. Marcus, 1968
• Flabellina engeli lucianae Dacosta, Cunha, Simone & Schrödl, 2007
• Flabellina evelinae Edmunds, 1989
• Flabellina ilidioi Calado, Ortea & Caballer, 2005
• Flabellina llerae Ortea, 1989

有異名的物種：

• Flabellina albomaculata Pola, Carmona, Calado & Cervera, 2014，為Edmundsella albomaculata]] (Pola, Carmona, Calado & Cervera, 2014)的異名^[7]
• Flabellina albomarginata (Miller, 1971)，為Coryphellina albomarginata]] (M. C. Miller, 1971)的異名
• Flabellina alisonae Gosliner, 1980:，為Flabellina bicolor的異名
• Flabellina arveloi Ortea & Espinosa, 1998，為Coryphellina arveloi (Ortea & Espinosa, 1998)的異名
• Flabellina athadona (Bergh, 1875)，為Occidenthella athadona (Bergh, 1875)的異名
• Flabellina babai Schmekel, 1972，為Luisella babai (Schmekel, 1972)的異名
• Flabellina baetica Garcia-Gomez, 1984，為Baenopsis baetica (Garcia-Gomez, 1984)的異名
• Flabellina bicolor (Kelaart, 1858)，為Samla bicolor (Kelaart, 1858)的異名
• Flabellina bilas Gosliner & Willan, 1991^[8]，為Samla bilas (Gosliner and Willan, 1991)的異名
• Flabellina californica (Bergh, 1904)，為Coryphella californica Bergh, 1904的異名
• Flabellina cerverai Fischer, van der Velde & Roubos, 2007，為Coryphellina cerverai (Fischer, van der Velde & Roubos, 2007)的異名
• Flabellina confusa Gonzalez-Duarte, Cervera & Poddubetskaia, 2008，為Calmella gaditana (Cervera, García-Gomez & García, 1987)的異名
• Flabellina cooperi (Cockerell, 1901)，為Orienthella cooperi (Cockerell, 1901)的異名
• Flabellina cynara (Marcus & Marcus, 1967)，為Kynaria cynara (Marcus and Marcus, 1967)的異名
• Flabellina delicata Gosliner & Willan, 1991，為Coryphellina delicata (Gosliner & Willan, 1991)的異名
• Flabellina exoptata Gosliner & Willan, 1991，為Coryphellina exoptata (Gosliner & Willan, 1991)的異名(圖片 （頁面存檔備份，存於網際網路檔案館）)
• Flabellina falklandica (Eliot, 1907)，為Itaxia falklandica (Eliot, 1907)的異名
• Flabellina funeka Gosliner & Griffiths, 1981，為Paraflabellina funeka (Gosliner & Griffiths, 1981)的異名
• Flabellina fusca (O'Donoghue, 1921)，為Flabellina trophina (Bergh, 1890)的異名
• Flabellina goddardi Gosliner, 2010，為Pacifia goddardi (Gosliner, 2010)的異名
• Flabellina hamanni Gosliner, 1994，為Coryphellina hamanni (Gosliner, 1994)的異名
• Flabellina ianthina Angas, 1864，為Pteraeolidia ianthina (Angas, 1864)的異名
• Flabellina inornata A. Costa, 1866，為Spurilla neapolitana的異名
• Flabellina insolita Garcia-Gomez & Cervera, 1989，為Fjordia insolita (Garcia-Gomez & Cervera, 1989)的異名
• Flabellina iodinea (J. G. Cooper, 1863)，為Flabellinopsis iodinea (J. G. Cooper, 1863)的異名
• Flabellina ischitana Hirano & Thompson, 1990，為Paraflabellina ischitana (Hirano & Thompson, 1990)的異名
• Flabellina islandica (Odhner, 1937)，為Paracoryphella islandica (Odhner, 1937)的異名
• Flabellina japonica (Volodchenko, 1937)，為Ziminella japonica (Volodchenko, 1937)的異名
• Flabellina macassarana Bergh, 1905，為Samla macassarana (Bergh, 1905)的異名
• Flabellina marcusorum Gosliner & Kuzirian, 1990，為Coryphellina marcusorum (Gosliner & Kuzirian, 1990)的異名
• Flabellina newcombi Angas, 1864：為Facelina newcombi (Angas, 1864)的異名
• Flabellina ornata Angas, 1864：為Austraeolis ornata(Angas, 1864)的異名
• Flabellina pallida (A. E. Verrill, 1900)，為Coryphella pallida A. E. Verrill, 1900的異名
• Flabellina parva (Hadfield, 1963)，為Paracoryphella parva (Hadfield, 1963)的異名
• Flabellina pedata (Montagu, 1815)，為Edmundsella pedata (Montagu, 1816)的異名
• Flabellina pellucida (Alder & Hancock, 1843)，為Carronella pellucida (Alder & Hancock, 1843)的異名
• Flabellina poenicia (Burn, 1957)，為Coryphellina poenicia (Burn, 1957)的異名
• Flabellina polaris Volodchenko, 1946，為Polaria polaris (Volodchenko, 1946)的異名
• Flabellina pricei (MacFarland, 1966)，為Apata pricei (MacFarland, 1966)的異名
• Flabellina riwo Gosliner & Willan, 1991，為Samla riwo]] Gosliner & Willan, 1991的異名
• 紅扇鰓海蛞蝓 Flabellina rubrolineata (O'Donoghue, 1929)，為Coryphellina rubrolineata O'Donoghue, 1929的異名
• Flabellina rubromaxilla Edmunds, 2015 ，為Paraflabellina rubromaxilla (Edmunds, 2015)的異名
• Flabellina rubropurpurata Gosliner & Willan, 1991，為Samla rubropurpurata (Gosliner and Willan 1991)的異名
• Flabellina salmonacea (Couthouy, 1838)，為Ziminella salmonacea (Couthouy, 1838)的異名
• Flabellina scolopendrella Risbec, 1928，為Pteraeolidia ianthina (Angas, 1864)的異名
• Flabellina semperi Bergh, 1870，為Pteraeolidia ianthina (Angas, 1864)的異名
• Flabellina stohleri Bertsch & Ferreira, 1974，為Flabellina telja Marcus & Marcus, 1967的異名
• Flabellina telja Marcus & Marcus, 1967，為Samla telja (Marcus and Marcus, 1967)的異名
• Flabellina trilineata (O'Donoghue, 1921)，為Orienthella trilineata (O'Donoghue, 1921)的異名
• Flabellina trophina (Bergh, 1890)，為Himatina trophina (Bergh, 1890)的異名
• Flabellina vansyoci Gosliner, 1994，為Edmundsella vansyoci (Gosliner, 1994)的異名
• Flabellina verrucicornis A. Costa, 1867:，為Berghia verrucicornis (A. Costa, 1867)的異名
• Flabellina versicolor Costa A., 1866:，為Favorinus branchialis (Rathke, 1806)的異名
• Flabellina verta (Ev. Marcus, 1970)，為Coryphella verta Ev. Marcus, 1970的異名

生物學

如同其他蓑海牛裸鰓動物物種那樣，本屬物種在吃掉刺胞動物後，能將牠們的刺絲胞儲存在自身背部的角突裡^[9][10]。

參考文獻

1. Griffith, E & E. Pidgeon. 1833-1834. The animal kingdom arranged in conformity with its organization, by the Baron Cuvier, member of the Institute of France, etc..., with supplementary additions to each order. Volume the twelfith. The Mollusca and Radiata. Whittaker, London. viii + 601 pp., pl 1-41
2. Gmelin, Johann Friedrich. 1791. In: C. Linnaeus. Systema Naturae, per regna tria naturae, : secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. ed. 13, 1(6):3103- 3107, 3147-3148.
3. Flabellina Gray, 1833. WoRMS.
4. Davies, Stephen. 左麗愛 (Catalina Chor) , 編. 海中噚 Fathoming the Sea. 香港海事博物館. ISBN 978-988-18233-2-8 （中文（繁體）及英語）.
5. Gosliner T.M., Griffiths R.J. (1981) Description and revision of some South African aeolidacean Nudibranchia (Mollusca, Gastropoda). Annals of the South African Museum 84: 105–150.
6. Korshunova, T.; Martynov, A.; Bakken, T.; Evertsen, J.; Fletcher, K.; Mudianta, W.; Saito, H.; Lundin, K.; Schrödl, M.; Picton, B. (2017). Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys. 717: 1-139.
7. Marta Pola, Leila Carmona, Gonçalo Calado & Juan Lucas Cervera (2015) A new nudibranch, Flabellina albomaculata sp. nov. (Flabellinidae), from the Cape Verde Archipelago with comparisons among all eastern Atlantic violet Flabellina spp., Marine Biology Research, 11:2, 218-222, DOI: 10.1080/17451000.2014.923102
8. Gosliner T. M. & Willan R. C. (1991). "Review of the Flabellinidae (Nudibranchia: Aeolidacea) from the tropical Indo-Pacific, with the descriptions of five new species". The Veliger 34(2): 97-133. page 109.
9. Frick, K. Predator Suites and Flabellinid Nudibranch Nematocyst Complements in the Gulf of Maine. SF Norton (編). Diving for Science: Proceedings of the American Academy of Underwater Sciences in 22nd Annual Scientific Diving Symposium. 2003 [2008-07-03]. （原始內容存檔於2011-01-06） （英語）.
10. Le Bris, S.; Sutour, J.-M. Flabellina rubrolineata (O'Donoghue, 1929). DORIS. 2014-03-29 [2018-04-26]. （原始內容存檔於2014-10-08） （法語）.

• ICZN. (1966). Opinion 781. Flabellina Voigt, 1834 (Gastropoda): Placed on the Official List of Generic Names. Bulletin of Zoological Nomenclature. 23 (2-3): 104-105 （頁面存檔備份，存於網際網路檔案館）
• Vaught, K.C. (1989). A classification of the living Mollusca. American Malacologists: Melbourne, FL (USA). ISBN 0-915826-22-4. XII, 195 pp
• Petit, R. E. & Coan., E. V. 2008. The molluscan taxa made available in the Griffith & Pidgeon (1833-1834) edition of Cuvier, with notes on the editions of Cuvier and on Wood's Index Testaceologicus. Malacologia 50: 219-264 page(s): 222

外部連結