孔腸科

孔腸科（學名： Opecoelidae）是斜睪目之下一個吸蟲綱複殖亞綱寄生蟲的科^[2]，是複殖亞綱之下物種最豐盛的一個科，包含有近100個屬、接近900個物種。這上千種物種全在淡水和鹹水生活的真骨類支序魚類身上寄生^[3]。

孔腸科
Hamacreadium cribbi
科學分類
界：	動物界 Animalia
門：	扁形動物門 Platyhelminthes
綱：	吸蟲綱 Trematoda
亞綱：	複殖亞綱 Digenea
目：	斜睪目 Plagiorchiida
科：	孔腸科 Opecoelidae Ozaki, 1925[1]

分類

總科

本科在WoRMS屬於Allocreadioidea總科，但在Bray et al.被劃為獨立一個總科 Opecoeloidea Ozaki, 1925 ，又或屬於 Brachycladioidea Odhner, 1905^[3]。

亞科與屬

本科物種尚可分為多個亞科，分別如下：

Helicometrinae亞科

Helicometrinae Bray, Cribb, Littlewood & Waeschenbach, 2016^[3]

• Helicometra Odhner, 1902^[4]
• Helicometrina Linton, 1910^[5]
• Neohelicometra Siddiqi & Cable, 1960^[6]

孔腸亞科

Opecoelinae Ozaki, 1925^[1]

• Anisoporus Ozaki, 1928^[7]
• Anomalotrema Zhukov, 1957^[8]
  • Anomalotrema koiae Gibson & Bray, 1984^[9]
  • Anomalotrema putjatini Zhukov, 1957
• Apertile Overstreet, 1969^[10]
  • Apertile holocentri (Manter, 1947) Overstreet, 1969
  • Apertile overstreeti Toman, 1992
• Coitocaecum Nicoll, 1915^[11]
• Dactylomyza Aken'Ova, 2003^[12]
• Dactylostomum Woolcock, 1935^[13]
• Dimerosaccus Shimazu, 1980^[14]
• Discoverytrema Gibson, 1976^[15]
• Genitocotyle Park, 1937^[16]
• Labracetabulum Reimer, 1987^[17]
• Manteriella Yamaguti, 1958^[18]
• Margolisia Bray, 1987^[19]
• Neodactylostomum Toman, 1996^[20]
• Notoporus Yamaguti, 1938^[21]
• 類孔屬 Opecoeloides Odhner, 1928^[22]
• 孔腸屬 Opecoelus Ozaki, 1925^[1]
• Opegaster Ozaki, 1928^[7]
• Paropecoelus Pritchard, 1966^[23]
• Parvacreadium Manter, 1940^[24]
• Poracanthium Dollfus, 1948^[25]
• Pseudopecoeloides Yamaguti, 1940^[26]
• Pseudopecoelus von Wicklen, 1946^[27]

Opecoelininae亞科

Opecoelininae Gibson & Bray, 1984^[28]

• Bartoliella Aken'Ova, 2003^[12]
• Buticulotrema Blend, Dronen & McEachran, 1993^[29]
• Opecoelina Manter, 1934^[30]

Plagioporinae亞科

Plagioporinae Manter, 1947^[31]

• Allopodocotyle Pritchard, 1966^[32]
• Anthochoanocotyle Kamegai, 1972^[33]
• Bathycreadium Kabata, 1961^[34]
• Bentholebouria Andres, Pulis & Overstreet, 2014^[35]
• Cainocreadium Nicoll, 1909^[36]
• Choanostoma Yamaguti, 1934^[37]
• Choerodonicola Cribb, 2005^[38]
• Decemtestis Yamaguti, 1934^[37]
• Diplobulbus Yamaguti, 1934^[37]
• Eucreadium Dayal, 1950^[39]
• Fairfaxia Cribb, 1989^[40]
• Gaevskajatrema Gibson & Bray, 1982^[41]
• Hamacreadium Linton, 1910^[5]
• Heterochoanostoma Machida, 2014^[42]
• Hysterogonia Hanson, 1955^[43]
• Jerguillicola Bray, 2002^[44]
• Macrourimegatrema Blend, Dronen & Armstrong, 2004^[45]
• Macvicaria Gibson & Bray, 1982^[41]
• Mesocreadium Reimer, 1987^[17]
• Neochoanostoma Bray & Cribb, 1989^[46]
• Neolebouria Gibson, 1976^[15]
• Neoplagioporus Shimazu, 1990^[47]
• Nicolla Wisniewski, 1933^[48]
• Orthodena Durio & Manter, 1968^[49]
• Pachycreadium Manter, 1954^[50]
• Pacificreadium Durio & Manter, 1968^[49]
• Paramanteriella Li, Qiu & Zhang, 1988^[51]
• Paraplagioporus Yamaguti, 1939^[52]
• Pellamyzon Montgomery, 1957^[53]
• Peracreadium Nicoll, 1909^[36]
• Phyllotrema Yamaguti, 1934^[37]
• Pinguitrema Siddiqi & Cable, 1960^[6]
• Plagioporus Stafford, 1904^[54]
• 肢盤吸蟲屬 Podocotyle Dujardin, 1845^[55]
  • Podocotyle angulata Dujardin, 1845
  • Podocotyle apodichthysi Park, 1937
  • Podocotyle araii Gibson, 1986^[56]
  • Podocotyle atherinae Nicoll, 1914
  • Podocotyle atomon (Rudolphi, 1802) Odhner, 1905
  • Podocotyle ayu Takahashi, 1928
  • Podocotyle bathyhelminthos Blend & Dronen, 2015
  • Podocotyle bongosi Nagaty & Abdel Aal, 1962
  • Podocotyle caithnessi Manter, 1954
  • 加州肢盤吸蟲 Podocotyle californica Park, 1937
  • Podocotyle congeri (Yamaguti, 1970) Bartoli, Bray & Gibson, 2003
  • Podocotyle enophrysi Park, 1937
  • Podocotyle gibbonsia Johnston, 1949
  • Podocotyle harrisae Bray & Campbell, 1996
  • 印度肢盤吸蟲 Podocotyle indica Pandey & Tiwari, 1989
  • Podocotyle koshari Nagaty, 1973
  • Podocotyle levinseni Issaitschikov, 1928
  • 梭肢盤吸蟲 Podocotyle lizae Qiu & Liang, 1995
  • Podocotyle longiforme Wang, Wang & Zhang, 1992
  • Podocotyle mehsena (Nagaty, 1941) Martin, Cutmore, Ward & Cribb, 2017
  • Podocotyle nibeae Qiu & Li, 1995
  • Podocotyle nimoyi Blend, Dronen & Armstrong, 2016
  • Podocotyle novella (Maillard & Lambert, 1978) Bartoli, Bray & Gibson, 2003
  • Podocotyle pearsei Manter, 1934
  • Podocotyle radifistuli (Acena, 1941) Gibson & Bray, 1984
  • Podocotyle reflexa (Creplin, 1825) Odhner, 1905
  • Podocotyle schistotesticulata Bray & Campbell, 1996
  • Podocotyle scorpaenae (Rudolphi, 1919) Bartoli & Gibson, 1991
  • Podocotyle simplex (Rudolphi, 1809) Srivastava, 1966
  • Podocotyle sinusacca Ching, 1960
  • Podocotyle synechogobii (Qiu & Li, 1985) Martin, Cutmore, Ward & Cribb, 2017
  • Podocotyle syngnathi Nicoll, 1913
  • Podocotyle tamane Yamaguti, 1942
  • Podocotyle temensis Fischthal & Thomas, 1970
  • Podocotyle tetrastyla Tang, 1990
  • Podocotyle theragrae (Lloyd, 1938) Gibson & Bray, 1984
  • Podocotyle unica (Molin, 1859) Stossich, 1886
• Podocotyloides Yamaguti, 1934^[37]
• Proneohelicometra Hassanine, 2006^[57]
• Propycnadenoides Fischthal & Kuntz, 1964^[58]
• Pseudoplagioporus Yamaguti, 1938^[21]
  • Pseudoplagioporus interruptus Durio & Manter, 1968^[59]
  • Pseudoplagioporus labiatus Martin, Cutmore & Cribb, 2019
  • Pseudoplagioporus lethrini Yamaguti, 1938
  • Pseudoplagioporus mediocris Martin, Cutmore & Cribb, 2019
  • Pseudoplagioporus microrchis Yamaguti, 1942
  • Pseudoplagioporus nebulosae Nagaty & Adbel Aal, 1969
  • Pseudoplagioporus roseovatulus Martin, Cutmore & Cribb, 2019
  • Pseudoplagioporus roseovulatus Martin, Cutmore & Cribb, 2019,2020
  • Pseudoplagioporus tropicus (Durio & Manter, 1968) Martin, Cutmore & Cribb, 2019
• Pseudopycnadena Saad-Fares & Maillard, 1986^[60]
• Pycnadena Linton, 1910^[5]
• Pycnadenoides Yamaguti, 1938^[21]
• Shimazuia Cribb, 2005^[38]
• Sphaerostoma Rudolphi, 1809^[61]
• Tellervotrema Gibson & Bray, 1982^[41]
• Thynstenopera Bilqees & Khatoon, 2004^[62]
• Trilobovarium Martin, Cutmore & Cribb, 2017^[63]
• Urorchis Ozaki, 1927^[64]
• Vesicocoelium Tang, Hsu, Huang & Lu, 1975^[65]
• Villarrealina Bolaños & Salas, 1982^[66]

Stenakrinae亞科

Stenakrinae Yamaguti, 1970^[67]

• Biospeedotrema Bray, Waeschenbach, Dyal, Littlewood & Morand, 2014^[68]
• Caudotestis Issaitschikov, 1928^[69]
• Hexagrammia Baeva, 1965^[70]
• Neonotoporus Srivastava, 1942^[71]
• Pseudopecoelina Yamaguti, 1942^[72]
• Scorpidotrema Aken'Ova & Cribb, 2003^[73]
• Stenakron Stafford, 1904^[54]

亞科未定

• Polypipapiliotrema Martin, Cutmore & Cribb, 2018
  • Polypipapiliotrema citerovarium Martin, Cutmore & Cribb, 2018
  • Polypipapiliotrema hadrometra Martin, Cutmore & Cribb, 2018
  • Polypipapiliotrema heniochi Martin, Cutmore & Cribb, 2018
  • Polypipapiliotrema ovatheculum Martin, Cutmore & Cribb, 2018
  • Polypipapiliotrema stenometra (Pritchard, 1966) Martin, Cutmore & Cribb, 2018^[74]

參考文獻

1. Ozaki, Y. Preliminary notes on a trematode with anus. Journal of Parasitology. 1925, 12: 51–53 （英語）.
2. Gibson, David; Cribb, Thomas. Opecoelidae Ozaki, 1925. WoRMS. 2013-03-29 [2018-01-31].
3. Bray, Rodney A.; Cribb, Thomas H.; Littlewood, D. Timothy J.; Waeschenbach, Andrea. The molecular phylogeny of the digenean family Opecoelidae Ozaki, 1925 and the value of morphological characters, with the erection of a new subfamily. Folia Parasitologica. 2010, 63: 13 （英語）.
4. Odhner, Theodor. II. 3. Helicometra n. g. Allocreadiinarum.. Mitteilungen zur Kenntnis der Distomem 31. 1902: 160–162 [2018-01-31]. ISSN 0323-6056. （原始內容存檔於2018-10-11） （德語）. |journal=被忽略 (幫助)
5. Linton, E. (1910). Helminth fauna of the Dry Tortugas. II. Trematodes. Carnegie Institution of Washington Publication No. 133, 11–98.
6. Siddiqi, Ather H. & Cable, Raymond M. (1960). Digenetic Trematodes of Marine Fishes of Puerto Rico. Scientific Survey of Porto Rico and the Virgin Islands 17(3).
7. Ozaki, Y. (1928). On some trematodes with anus. Japanese Journal of Zoology 2, 5–33.
8. Zhukov, E. V. (1957). New genera and species of trematodes parasites of fish in the Far-Eastern seas. Zoologicheskii Zhurnal 36, 840–846.
9. David I. Gibson & Rodney A. Bray (1984) On Anomalotrema Zhukov, 1957, Pellamyzon Montgomery, 1957, and Opecoelina Manter, 1934 (Digenea: Opecoelidae), with a description of Anomalotrema koiae sp. nov. from North Atlantic waters, Journal of Natural History, 18:6, 949-964, DOI: 10.1080/00222938400770831
10. Digenetic trematodes of marine teleost fishes from Biscayne Bay, Florida. Tulane Studies in Zoology and Botany 15, 119–176.
11. Nicoll, W. (1915). The trematode parasites of north Queensland. III. Parasites of fishes. Parasitology 8, 24–41.
12. Aken'Ova (2003). Dactylomyza gibsoni n. g., n. sp. (Digenea: Opecoelidae) from Schuettea woodwardi (Waite) (Monodactylidae) from off Western Australia. Systematic Parasitology 55(1), 61–65
13. Woolcock, V. (1935). Digenetic Trematodes from Some Australian Fishes. Parasitology 27(3), 309–331.
14. Shimazu, T. (1980). Dimerosaccus gen. nov. (Digenea: Opecoelidae), with a redescription of its type species, Dimerosaccus oncorhynchi (Eguchi, 1931) comb. nov. Japanese Journal of Parasitology 29(3), 163–168.
15. Gibson, D.I. (1976). Monogenea and Digenea from fishes. Discovery Reports 36, 179–266.
16. Park, J. T. (1937). A new trematode, Genitocotyle acirrus gen. nov., sp. nov. (Allocreadiidae), from Holconotus rhodoterus. Transactions of the American Microscopical Society 56(1), 67.
17. Reimer, L. W. (1987). Opecoelidae (Trematoda) von Meeresfischen vor Moçambique. Angewandte Parasitologie 28, 143–158.
18. Yamaguti, S. (1958). Systema helminthum. Vol. 1. The digenetic trematodes of vertebrates. Interscience Publications, New York. 1–1575.
19. Bray, R. A. (1987). Some helminth parasites of marine fishes of south Africa: family Opecoelidae (Digenea). Journal of Natural History 21(4), 1049–1075.
20. Toman, G. (1992). Digenetic trematodes of marine teleost fishes from the Seychelles, Indian Ocean. III. Acta Parasitologica 37(3), 119–126.
21. Yamaguti, S. (1938). Studies on the Helminth Fauna of Japan. Part 21. Trematodes of Fishes, IV. Kyöto: Yamaguti S. 139.
22. Odhner, T. (1928). Weitere Trematoden mit Anus. Arkiv för Zoologi 20, 1–6.
23. Pritchard, M. H. (1966). Studies on digenetic trematodes of Hawaiian fishes: family Opecoelidae Ozaki. Zoologische Jahrbucher, Abteilung für Systematik 93, 173–202.
24. Manter, H. W. (1940). Digenetic trematodes of fishes from the Galapagos Islands and the neighboring Pacific. Allan Hancock Pacific Expeditions 2, 364–473.
25. Dollfus, R. P. (1948). L'enigme de Distoma furcatum Bremser enfin expliquée. Contribution à la connaissance des trématodes des poissons du genre Mullus en Méditerranée. Bulletin de l'Institut Océanographique, Monaco 45(928), 1–23.
26. Yamaguti, S. (1940). Studies on the helminth fauna of Japan Part 31. Trematodes of fishes, VII. Japanese Journal of Zoology 9, 35–108.
27. von Wicklen, J. H. (1946). The trematode genus Opecoeloides and related genera, with a description of Opecoeloides polynemi n. sp. The Journal of Parasitology 32(2), 156–163.
28. Gibson, D. I. & Bray, R. A. 1984. On Anomalotrema Zhukov, 1957, Pellamyzon Montgomery, 1957 and Opecoelina Manter, 1934 (Digenea, Opecoelidae), with a description of Anomalotrema koiae n. sp. from North Atlantic waters. Journal of Natural History 18, 949–964.
29. Blend, C. K., Dronen, N. O. & McEachran, J. D. (1993). Buticulotrema stenauchenus n. gen., n. sp. (Digenea: Opecoelidae) from Malacocephalus occidentalis and Nezumia aequalis (Macrouridae) from the Gulf of Mexico. The Journal of Parasitology 79(5), 674–676.
30. Manter, H. W. (1934). Some digenetic trematodes from deep-water fish of Tortugas, Florida. Papers from Tortugas Laboratory 28, 257–345.
31. Manter, H.W. (1947). The digenetic trematodes of marine fishes of Tortugas. American Midland Naturalist 38(2), 257–416.
32. Pritchard, M. H. (1966). A revision of the genus Podocotyle (Trematoda: Opecoelidae). Zoologische Jahrbucher, Abteilung für Systematik 93, 158–172.
33. Kamegai, S. (1972). Anthochoanocotyle kihobo n. g., n. sp. (Trematoda: Opecoelidae, Plagioporinae) from deep sea fish. Japanese Journal of Parasitology 21, 333–335.
34. Kabata, Z. (1961). A new genus and species of trematode parasitic in Macrurus fabricii (Sundeval), a deep-sea fish. Proceedings of the Zoological Society of London 136, 285–292.
35. Andres, M. J., Pulis, E. E. & Overstreet, R. M. (2014). New genus of opecoelid trematode from Pristipomoides aquilonaris (Perciformes: Lutjanidae) and its phylogenetic affinity within the family Opecoelidae. Folia Parasitologica 61(3), 223–230.
36. Nicoll, W. (1909). Studies on the structure and classification of the digenetic trematodes. Quarterly Journal of Microscopical Science, New Series 53(3), 391–487.
37. Yamaguti, S. (1934). Studies on the helminth fauna of Japan. Part 2. Trematodes of fishes. I. Japanese Journal of Zoology 5, 249–541.
38. Cribb, T.H. (2005). Family Opecoelidae Ozaki, 1925. In Bray, R. A., Gibson, D. I. & Jones, A. (Eds.), Keys to the Trematoda. Vol. 3 (pp. 443–531). London, UK: CABI Publishing and The Natural History Museum.
39. Dayal, J. 1950. Trematode parasites of Indian fishes - Part III. Two new trematodes of the family Allocreadidae from fresh water fishes of India. Indian Journal of Helminthology 2 1–10.
40. Cribb, T. (1989). Fairfaxia lethrini, gen. et sp. nov. (Digenea : Opecoelidae), from Lethrinus chrysostomus Richardson from the Southern Great Barrier Reef. Australian Journal of Zoology 37(1), 67–70.
41. Gibson, D. I. & Bray, R. A. (1982). A study and reorganization of Plagioporus Stafford, 1904 (Digenea: Opecoelidae) and related genera, with special reference to forms from European Atlantic waters. Journal of Natural History 16(4), 529–559.
42. Machida, M. (2014). Seven Species of Opecoelid Digeneans (Trematoda) from Fishes of Southern Japan, Palau and the Philippines. Bulletin of the National Museum of Natural Sciences, Ser. A 40(1), 1–3.
43. Hanson, M. L. (1955). Some Digenetic Trematodes of Plectognath Fishes of Hawaii. Proceedings of the Helminthological Society of Washington 22(2), 75–87.
44. Bray, R. A. (2002). Three species of plagioporine opecoelids (Digenea), including a new genus and two new species, from marine fishes from off the coast of Chile. Systematic Parasitology 51(3), 227–236.
45. Blend, C. K., Dronen, N. O. & Armstrong, H. W. (2004). Macrourimegatrema brayi n. gen., n. sp. (Digenea: Opecoelidae) from four species of deep-sea macrourid fishes from the Gulf of Mexico and Caribbean Sea, with a list of endohelminths reported from species of Bathygadus and Gadomus (Macrouridae). Zootaxa 566(1), 1–18.
46. Bray, R. A. & Cribb, T. H. (1989). Digeneans of the family Opecoelidae Ozaki, 1925 from the southern Great Barrier Reef, including a new genus and three new species. Journal of Natural History 23(2), 429–473.
47. Shimazu, T. (1990).Trematodes of a new genus, Neoplagioporus gen. n. (Digenea: Opecoelidae: Plagioporinae), and an unidentified opecoelid from freshwater fishes of Japan. Japanese Journal of Parasitology 39(4), 384–396.
48. Wisniewski, L. W. (1933). Beitrag zur systematik der Coitocaecidae (Trematoda).Nicolla g. n., Ozakia g. n., Coitocaecum proavitum sp. n. Mémoires de l'Académie Polonaise des Sciences et des Lettres, Sér. B, 27–41.
49. Durio, W. O. & Manter, H. W. (1968). Some Digenetic Trematodes of Marine Fishes of New Caledonia. Part II. Opecoelidae and Lepocreadiidae. The Journal of Parasitology 54(4), 747.
50. Manter, H. W. (1954). Some digenetic trematodes from fishes of New Zealand. Transactions of the Royal Society of New Zealand 82, 475–568.
51. Li, Q. K.; Qiu, Z. Z.; Zhang, R. S. Digenetic trematodes of fishes from the Bo-Hai Sea, China. 6. (Trematoda: Opecoelidae). Acta Zootaxonomica Sinica. 1989, 14 (1): 2–16.
52. Yamaguti, S. (1939). Studies on the helminth fauna of Japan. Part 26. Trematodes of fishes, VI. Japanese Journal of Zoology 8, 211–230.
53. Montgomery, W. R. (1957). Studies on Digenetic Trematodes from Marine Fishes of La Jolla, California. Transactions of the American Microscopical Society 76(1), 13–36.
54. Stafford, J. (1904). I. Wissenschaftliche Mitteilungen. 1. Trematodes from Canadian Fishes. Zoologischer Anzeiger XXVII(16–17), 481–495.
55. Dujardin, F. (1845). Histoire naturelle des Helminthes ou vers intestinaux. Paris: Libraire encyclopédique de Roret (Suites a Buffon).
56. David I. Gibson (1986) Podocotyle araii sp. nov. (Digenea: Opecoelidae) from Sebastes spp. off the coast of British Columbia, with a key to the species of Podocotyle Dujardin, 1845, occurring off the Pacific coast of North America, Journal of Natural History, 20:3, 735-743, DOI: 10.1080/00222938600770511
57. Hassanine, R. M. E. (2006). Trematodes from Red Sea fishes: Proneohelicometra aegyptensis gen. nov., sp. nov. (Opecoelidae Ozaki, 1925) and Neohypocreadium gibsoni sp. nov. (Lepocreadiidae Odhner, 1905). Acta Parasitologica 51(4), 249–254.
58. Fischthal, Jacob H. & Kuntz, Robert E. (1964). Digenetic Trematodes of Fishes from Palawan Island, Philippines. Part I. Families Acanthocolpidae, Angiodictyidae, Cryptogonimidae, Fellodistomidae, and Gyliauchenidae. The Journal of Parasitology 50(2), 248–252.
59. Abdou N E , Heckmann R A , Beltagy S M ,et al.Pseudoplagioporus interruptus Durio and Manter, 1968 and Hamacreadium agyptia sp. n. (Trematoda: Opecoeliidae) from the Red Sea Fish in Egypt[J].[2023-10-24].
60. Saad-Fares, A.; Maillard, C. Trématodes des poissons des côtes du Liban. I: Pseudopycnadena fischthali n.g. n.sp. (Trematoda Opistholebetidae) parasite des Diplodus (Teleostei) des côtes du Liban. Annales de Parasitologie Humaine et Comparée. 1986, 61 (3): 303–309 [2018-01-31]. ISSN 0003-4150. doi:10.1051/parasite/1986613303. （原始內容存檔於2020-08-09）.
61. Rudolphi, C. A. (1819). Entozoorum synopsis, cui accedunt mantissa duplex et indices locupletissimi. Cum tab. III aeneis. Berolini: Sumtibus Augusti Rücker.
62. Bilqees, F. & Khatoon, N. (2004). A new trematode species Thynstenopera lobata n. gen., n. sp. (Trematoda: Opecoelidae: Plagioporinae) another new trematode from the marine fish Thynnus thunnia of Karachi Coast, Pakistan. Pakistan Journal of Biological Sciences 7, 1343–1345.
63. Martin, S. B., Cutmore, S. C., & Cribb, T. H. (2017). Revision of Neolebouria Gibson, 1976 (Digenea: Opecoelidae), with Trilobovarium n. g., for species infecting tropical and subtropical shallow-water fishes. Systematic Parasitology 94(3), 307–338.
64. Ozaki, Y. (1927). Two new genera of fish trematodes. Japanese Journal of Zoology 1, 157–164.
65. Tang, C., Hsu, C., Huang, M. & Lu, S. (1975). Studies on the parasitic disease of Chinese razor clam (Sinonovacula contricta Lamarck) in the northern estuary of Chiulung River, Fukien. Journal of Xiamen University (Natural Science) 13, 161–177.
66. Bolaños, L. L. & Salas, E. M. (1982). Villarrealina peruanus gen. et sp. n. (Digenea: Allocreadiidae) en el pez marino Sciaena gilberti. Investigación. Ciencia. Universidad Federico Villarreal 1, 51–53.
67. Yamaguti, S. (1970). Digenetic trematodes of Hawaiian fishes. Tokyo: Keigaku Publishing Co.
68. Bray, R. A., Waeschenbach, A., Dyal, P., Littlewood, D. T. & Morand, S. (2014). New digeneans (Opecoelidae) from hydrothermal vent fishes in the south eastern Pacific Ocean, including one new genus and five new species. Zootaxa 3768(1), 73–87.
69. Isaaitcshikow, I. M. (1928). Zur Kenntnis der parasitischen Würmer einiger Gruppen von Wirbeltieren der russischen Arktis. Trucdy Morskogo nauchnogo instituta 3(2), 5–79.
70. Baeva, O. M. (1965). [Helminth fauna of Far Eastern mackerel. In Parasitic Worms of Domestic and Wild Animals]. In: Leonov, V. A., Mamaev, Y. L. & Oshmarin, P. G. (Eds.), Parasitic worms of domestic and wild animals. Papers on helminthology presented to Prof. A. A. Sobolev on the 40th anniversary of his scientific and teaching activity. Vladivostok: Dalnevostochnii Gosudarstvennii Universitet. 23–32.
71. Srivastava, H. D. 1942. New allocreadiids (Trematoda) from Indian marine food fishes. Part IV. The morphology and systematic position of a new genus, Horatrema, of digenetic trematodes. Parasitology 34(1) 128–132.
72. Yamaguti, S. (1942). Studies on the helminth fauna of Japan. Part 39. Trematodes of fishes mainly from Naha. Transactions of the Biogeographical Society of Japan 3, 329–398.
73. Aken'Ova, T. L. & Cribb, T. H. (2003). Scorpidotrema longistipes ng, n. sp.(Digenea: Opecoelidae) from Scorpis georgiana (Teleostei: Scorpididae) from southern Western Australia. Systematic Parasitology 54(2), 97-102
74. Polypipapiliotrema stenometra. Parasite of the Day. [2024-03-13]. （原始內容存檔於2024-03-13）.