龙虱科 （学名：Dytiscidae ），亦作龙虱科 ，为水甲虫 鞘翅目 之下的肉食亚目 。生活在田野、水沟、小溪等水体中，是一种掠食性 水生昆虫 。其学名源于希腊文δυτικός dytikos ），意思就是“肉食性的潜水甲虫”。本科物种几乎见于全世界所有淡水栖境，但亦有部分于落叶堆中栖息[ 1] Dytiscus latissimus Megadytes ducalis [ 1] [ 2] 澳大利亚 地下水中的Limbodessus [ 1] 橄榄绿 ，某些亚科 的物种会在其边沿带有金黄色的条状斑纹。
“
龙虱 ”重定向至此。“
龙虱 ”也可以指小型单人
龙舟 。
Quick Facts 龙虱科 化石时期：侏罗纪晚期至今, 科学分类 ...
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本科物种的成虫，特别是Cybister 可食用昆虫 。在美国内华达州 洪堡干湖 粪化石 内发现有Cybister explanatus [ 3] 墨西哥 ，他们在吃C. explanatus 时会以墨西哥夹饼 夹着来吃。在日本多个地区（例如：长野县 ）有以日本龙虱 （Cybister japonicus 中华人民共和国 广东省 ，除了日本龙虱以外，亦会食用C. bengalensis C. guerini C. limbatus C. sugillatus C. tripunctatus D. marginalis 台湾 、泰国 及新几内亚 亦有食用本科物种的习惯[ 4]
一只详细分类未明的本科物种 大但在陆地上很慢，成年后也不是特别凶猛，它们也被许多中型鸟类、哺乳动物和其他大型捕食者津津有味地吃掉。幼虫通常比较安全，由于它们的保护色和通过水柱逃跑的能力；它们可能相当难抓，可能成为小池塘的顶级掠食者。
本科包括下列各属[ 5] [ 6]
Acilius Leach, 1817
Aethionectes Sharp, 1882
Africodytes Biström, 1988
Africophilus Guignot, 1948
Agabetes Crotch, 1873
Agabinus Crotch, 1873
豆龙虱属 Agabus Leach, 1817[ 6] Agametrus Sharp, 1881
Agaporomorphus Zimmermann, 1921
Aglymbus Sharp, 1880
Agnoshydrus Biström, Nilsson & Wewalka, 1997
Allodessus Guignot, 1953
彩斑龙虱属 Allopachria Zimmermann, 1924
Amarodytes Régimbart, 1900
Amurodytes Fery & Petrov, 2013
Andex Sharp, 1882
Andonectes Guéorguiev, 1971
Anginopachria Wewalka, Balke & Hendrich, 2001
Anodocheilus Babington, 1841
Antiporus Sharp, 1882
Australphilus Watts, 1978
Austrelatus Shaverdo, Hájek, Hendrich, Surbakti, Panjaitan & Balke, 2023
Austrodytes Watts, 1978
Barrethydrus
Barretthydrus Lea, 1927
Barretthydrus geminatus Lea, 1927Barretthydrus stepheni Watts, 1978Barretthydrus tibialis Lea, 1927 Batrachomatus Clark, 1863
Belladessus K.B.Miller & Short, 2015
Bidessini Sharp, 1880
Bidessodes Régimbart, 1895
Bidessonotus Régimbart, 1895
Bidessus Sharp, 1882
Boreonectes (Fery & Ribera, 2018) Fery & Ribera, 2018
Boreonectes Angus, 2010
Borneodessus Balke, Hendrich, Mazzoldi & Biström, 2002
Brachyvatus Zimmermann, 1919
Brancuporus Hendrich, Toussaint & Balke, 2014
Bunites Spangler, 1972
Calicovatellus K.B.Miller & Lubkin, 2001
Canthyporus Zimmermann, 1919
Capelatus Turner & Bilton, 2015
Caperhantus Balke, Hájek & Hendrich, 2017
Carabdytes Balke, Hendrich & Wewalka, 1992
Carabhudrus
Carabhydrus Watts, 1978
Carrhydrus
Celina Aubé, 1837
Chostonectes Sharp, 1880
Clarkhydrus Fery & Ribera, 2018
Clemnius Villastrigo, Ribera, Manuel, Millán & Fery, 2017
Clypeodytes Régimbart, 1894
Coelambus Thomson, 1860
Coelambus cribrarius (Scudder, 1900)Coelambus derelictus (Scudder, 1900)Coelambus disjectus (Scudder, 1900)Coelambus dzieduszyckii (Lomnicki, 1894)Coelambus ganglbaueri (Lomnicki, 1894)Coelambus infernalis (Scudder, 1900)Coelambus latefasciatus (Lomnicki, 1894)Coelambus miocenus (Wickham, 1912)Coelambus niedzwiedzkii (Lomnicki, 1894)Coelambus ozokeriticus (Lomnicki, 1894)Coelambus picipoides (Lomnicki, 1894) Coelhydrus Sharp, 1882
Colymbetes Clairville, 1806
Colymbetes Schellenberg, 1806
Comaldessus Spangler & Barr, 1995
Copelatus Erichson, 1832
Copelatus abonnenci Guignot, 1939Copelatus acamas Guignot, 1955Copelatus adelbert Megna, Atthakor, Manaono, Hendrich & Balke, 2017Copelatus advena Sharp, 1882Copelatus aemulus Bilardo & Rocchi, 1995Copelatus aequatorius Régimbart, 1899Copelatus aethiopicus Régimbart, 1906Copelatus agaroensis Balke, 2008Copelatus agrias Guignot, 1954Copelatus alternatus Sharp, 1882Copelatus amaroides Guignot, 1952Copelatus amatolensis Omer-Cooper, 1965Copelatus amazonicus Régimbart, 1889Copelatus ambiguus Bertrand & Legros, 1975Copelatus amphibius Ranarilalatiana & Bergsten, 2019Copelatus anastomosans Guignot, 1952Copelatus andobonicus Guignot, 1960Copelatus andreinii Régimbart, 1905Copelatus andrewesi J.Balfour-Browne, 1939Copelatus angolensis Peschet, 1924Copelatus angouei Bilardo & Rocchi, 2011Copelatus angustus Gschwendtner, 1932Copelatus ankaratra Ranarilalatiana & Bergsten, 2019Copelatus annobomensis J.Balfour-Browne, 1939Copelatus anthracinus Régimbart, 1895Copelatus antoniorum Hájek & Brancucci, 2011Copelatus aphroditae Balke, 2003Copelatus apicinotatus Bilardo & Rocchi, 2018Copelatus assalyi Bilardo & Rocchi, 2010Copelatus assimilis Régimbart, 1895Copelatus ateles Guignot, 1955Copelatus atrosulcatus Régimbart, 1906Copelatus babyrousa Hájek, Hendrich & Balke, 2018Copelatus bacchusi Wewalka, 1981Copelatus bacillifer Guignot, 1955Copelatus baculiformis Guignot, 1955Copelatus badeni Sharp, 1882Copelatus bakewelli J.Balfour-Browne, 1939Copelatus baoulicus Bilardo & Pederzani, 1978Copelatus bapounouensis Bilardo & Rocchi, 2019Copelatus baranensis Hájek, Shaverdo, Hendrich & Balke, 2021Copelatus barbouri Young, 1942Copelatus basifasciatus Régimbart, 1895Copelatus basilewskyi Bilardo & Pederzani, 1979Copelatus bechynei Guignot, 1953Copelatus befasicus Guignot, 1956Copelatus bengalensis Guignot, 1955Copelatus bertrandi Nilsson, Bilardo & Rocchi, 1996Copelatus betampona Ranarilalatiana & Bergsten, 2019Copelatus bezdeki Sheth, Ghate & Hájek, 2018Copelatus bibulus Guignot, 1948Copelatus bicolor (Fabricius, 1792) Copelatus biformis Sharp, 1882Copelatus bimaculatus (Resende & Vanin, 1991) Copelatus binaghii Bilardo & Pederzani, 1978Copelatus biolleyi Guignot, 1952Copelatus biroi Guignot, 1956Copelatus birougouensis Bilardo & Rocchi, 2018Copelatus biseriatus J.Balfour-Browne, 1939Copelatus biswasi Mukherjee & Sengupta, 1986Copelatus blancasi Guignot, 1958Copelatus blatchleyi Young, 1953Copelatus bolivianus Guignot, 1957Copelatus bombycinus Guignot, 1952Copelatus bonvouloiri Sharp, 1882Copelatus bottegoi Régimbart, 1895Copelatus bougainvillensis Hájek, Shaverdo, Hendrich & Balke, 2021Copelatus boukali Hendrich & Balke, 1998Copelatus brancuccii Rocchi, 1979Copelatus brasiliensis Zimmermann, 1921Copelatus brendelli Hájek, Hendrich & Balke, 2018Copelatus brevicornis (Sharp, 1882) Copelatus brevistrigatus Guignot, 1959Copelatus brivioi Rocchi, 1976Copelatus bromeliarum (Scott, 1912) Copelatus brullei Aubé, 1838Copelatus brunneus J.Balfour-Browne, 1939Copelatus buqueti Aubé, 1838Copelatus burgeoni Gschwendtner, 1930Copelatus caelatipennis Aubé, 1838Copelatus caelatipennis Chevrolat, 1863Copelatus caelatus Guignot, 1952Copelatus caffer J.Balfour-Browne, 1939Copelatus calaquei Bilardo & Rocchi, 2008Copelatus camerunensis Guignot, 1941Copelatus capensis Sharp, 1882Copelatus carayoni Legros, 1949Copelatus carinatus Sharp, 1882Copelatus celinoides Guignot, 1952Copelatus cessaima Caetano, Bená & Vanin, 2013Copelatus chevrolati Aubé, 1838Copelatus chibcha Guignot, 1952Copelatus chinensis Regimbart, 1899Copelatus chipiriricus Guignot, 1957Copelatus chloroticus Régimbart, 1899Copelatus cinnamomeus Régimbart, 1895Copelatus clarki Sharp, 1882Copelatus collarti Gschwendtner, 1932Copelatus compertus Guignot, 1956Copelatus concii Bilardo, 1982Copelatus concolor Sharp, 1882Copelatus concolorans J.Balfour-Browne, 1939Copelatus confinis Bilardo & Rocchi, 1999Copelatus congo Gschwendtner, 1938Copelatus consimilis Bilardo & Rocchi, 2002Copelatus consors Sharp, 1882Copelatus cooperae Bilardo & Pederzani, 1972Copelatus cordovai Megna & Epler, 2012Copelatus cordylinoides J.Balfour-Browne, 1938Copelatus coxalis Sharp, 1882Copelatus crassus Régimbart, 1895Copelatus cryptarchoides Régimbart, 1899Copelatus cubaensis Schaeffer, 1908Copelatus curtistriatus Bilardo & Rocchi, 1995Copelatus curvispinis Bilardo & Rocchi, 2008Copelatus cynthiae Bilardo & Rocchi, 2008Copelatus daemeli Sharp, 1882Copelatus danyi Megna & Epler, 2012Copelatus darlingtoni Young, 1942Copelatus davidi Wewalka, 2017Copelatus debilis Sharp, 1882Copelatus deccanensis Sheth, Ghate & Hájek, 2018Copelatus decellei Bilardo, 1982Copelatus decemsulcatus Régimbart, 1895Copelatus deceptor Bilardo & Rocchi, 1995Copelatus dentatipenis Jiang, Hajek & Jia, 2022Copelatus depressus Sharp, 1882Copelatus descarpentriesi Bertrand & Legros, 1975Copelatus diffisus Guignot, 1939Copelatus dimorphus Sharp, 1882Copelatus distinctus Aubé, 1838Copelatus distinguendus Régimbart, 1903Copelatus diversistriatus Jiang, Hajek & Jia, 2022Copelatus djiboutensis Wewalka & Jäch, 2017Copelatus dolosus Guignot, 1956Copelatus doriae Sharp, 1882Copelatus doudouensis Bilardo & Rocchi, 2015Copelatus duodecimstriatus Aubé, 1838Copelatus duponti Aubé, 1838Copelatus efoutensis Bilardo & Rocchi, 1995Copelatus ejactus Omer-Cooper, 1965Copelatus ellai Bilardo & Rocchi, 1995Copelatus elutus Guignot, 1958Copelatus enganensis Guignot, 1940Copelatus epactus Guignot, 1948Copelatus erichsonii Guérin-Méneville, 1847Copelatus espinhasso Hájek, Alarie, Benetti, Hamada, Springer, Hendrich, Villastrigo, Torres, Basantes & Balke, 2024[ 7] Copelatus esteriensis Bilardo & Pederzani, 1978Copelatus eucritus Guignot, 1952Copelatus evanidus Bilardo & Rocchi, 1995Copelatus exaratus Sharp, 1882Copelatus externus Kirsch, 1873Copelatus falciformis Pederzani & Schizzerotto, 2017Copelatus fasciatus Bilardo & Rocchi, 1995Copelatus fastidiosus Guignot, 1959Copelatus feae Régimbart, 1888Copelatus felicis Hajek, Jiang & Jia, 2022Copelatus fernandensis Guignot, 1952Copelatus ferruginicollis Régimbart, 1895Copelatus ferus Guignot, 1953Copelatus festae Régimbart, 1899Copelatus fidschiensis Zimmermann, 1928Copelatus fijiensis Guignot, 1955Copelatus filiformis Sharp, 1882Copelatus fizpaci Bilardo & Rocchi, 2013Copelatus flavicans Guignot, 1952Copelatus flavidus Régimbart, 1895Copelatus florae Hájek, Alarie, Benetti, Hamada, Springer, Hendrich, Villastrigo, Torres, Basantes & Balke, 2024[ 7] Copelatus fluviaticus Guignot, 1957Copelatus fontanus J.Balfour-Browne, 1950Copelatus fossilis Říha, 1974 Copelatus fractistriatus Bilardo & Rocchi, 1995Copelatus freudei Guignot, 1955Copelatus fryi J.Balfour-Browne, 1939Copelatus fulviceps J.Balfour-Browne, 1938Copelatus fuscipennis Sharp, 1882Copelatus fuscomaculatus Guignot, 1952Copelatus gabonicus Bilardo & Pederzani, 1978Copelatus galapagoensis G.R.Waterhouse, 1845Copelatus garambanus Guignot, 1955Copelatus gardineri Scott, 1912Copelatus geayi Régimbart, 1904Copelatus geniculatus Sharp, 1882Copelatus gentilis Sharp, 1882Copelatus gestroi (Sharp, 1882) Copelatus gigas Bilardo & Rocchi, 2019Copelatus glyphicus (Say, 1823) Copelatus griffinii Régimbart, 1899Copelatus gschwendtneri Guignot, 1939Copelatus guadelupensis Legros, 1948Copelatus guerini Aubé, 1838Copelatus guineensis Guignot, 1955Copelatus haemorrhoidalis Régimbart, 1883Copelatus hararensis Guignot, 1952Copelatus hardenbergi J.Balfour-Browne, 1950Copelatus hebeter Guignot, 1953Copelatus heterogynus Régimbart, 1899Copelatus hydroporoides (Murray, 1859) Copelatus ibrahimi Zalat, Saleh, Angus & Kaschef, 2000Copelatus iguelaensis Bilardo & Rocchi, 2002Copelatus ilybioides Régimbart, 1895Copelatus imitator Bilardo & Rocchi, 2002Copelatus inaequalis Sharp, 1882Copelatus incognitus Sharp, 1882Copelatus indicus Sharp, 1882Copelatus inopinatus Bilardo & Rocchi, 1995Copelatus inornatus Sharp, 1882Copelatus insidiosus Bilardo & Rocchi, 1995Copelatus insolitus Chevrolat, 1863Copelatus instabilis Régimbart, 1897Copelatus instriatus Zimmermann, 1921Copelatus insuetus Guignot, 1941Copelatus insulanus Guignot, 1939Copelatus integer Sharp, 1882Copelatus internus Régimbart, 1904Copelatus interstriatus J.Balfour-Browne, 1939Copelatus inuber Guignot, 1952Copelatus ipiformis Régimbart, 1895Copelatus irinus Régimbart, 1899Copelatus irregularis W.J.MacLeay, 1871Copelatus ischius Guignot, 1956Copelatus jactator Guignot, 1955Copelatus jaechi Bilardo & Rocchi, 2016Copelatus jamaicensis Young, 1942Copelatus japonicus Sharp, 1884Copelatus jarrigei Legros, 1954Copelatus javanus Régimbart, 1883Copelatus jocosus Guignot, 1955Copelatus johannis Guignot, 1954Copelatus kalaharii Gschwendtner, 1935Copelatus kammuriensis Tamu & Tsukamoto, 1955Copelatus kaszabi Guignot, 1956Copelatus kely Ranarilalatiana & Bergsten, 2019Copelatus kietensis Hájek, Shaverdo, Hendrich & Balke, 2021Copelatus kilimandjarensis Bilardo & Pederzani, 1972Copelatus kindianus Guignot, 1954Copelatus kongouensis Bilardo & Rocchi, 1999Copelatus koreanus Mori, 1932Copelatus laccophilinus Sharp, 1882Copelatus laeticulus Sharp, 1882Copelatus laevipennis Hájek, Shaverdo, Hendrich & Balke, 2021Copelatus lamottei Legros, 1954Copelatus lanzai Bilardo & Rocchi, 1995Copelatus laraensis Bilardo & Rocchi, 1995Copelatus lasckonyi Bilardo & Rocchi, 1995Copelatus latens Guignot, 1952Copelatus laticollis Régimbart, 1899Copelatus latifasciatus Bilardo & Rocchi, 1999Copelatus latipes Sharp, 1882Copelatus latus J.Balfour-Browne, 1950Copelatus leonardii Bilardo & Rocchi, 1999Copelatus leonensis Legros, 1954Copelatus lepersonneae Gschwendtner, 1943Copelatus lignosus Guignot, 1955Copelatus lineatipennis Guignot, 1955Copelatus lineatus (Guérin-Méneville, 1838) Copelatus longicornis Sharp, 1882Copelatus lootensi Guignot, 1955Copelatus louayensis Bilardo & Rocchi, 2004Copelatus luctuosus Guignot, 1939Copelatus luridescens Régimbart, 1889Copelatus luteocinctus Guignot, 1955Copelatus luteomaculatus Guignot, 1956Copelatus luzonicus Guignot, 1952Copelatus macellus Guignot, 1950Copelatus madoni Guignot, 1952Copelatus mahajanga Pederzani & Hájek, 2005Copelatus mahleri Holmen & Vazirani, 1990Copelatus makokouensis Bilardo & Rocchi, 1995Copelatus malaisei Guignot, 1954Copelatus malavergnorum Manuel & Ramahandrison, 2022Copelatus mancus Sharp, 1887Copelatus marginatus Sharp, 1882Copelatus marginipennis (Laporte, 1835) Copelatus martinbaehri Hendrich, Shaverdo, Hájek & Balke, 2019Copelatus martini Manuel, Deler-Hernández, Megna & Hájek, 2018Copelatus masculinus Régimbart, 1899Copelatus massaicus Guignot, 1941Copelatus mathani Guignot, 1952Copelatus maushomi Sheth, Ghate & Hájek, 2018Copelatus mbokoensis Bilardo & Rocchi, 2006Copelatus melanogrammus Régimbart, 1883Copelatus mendax Bilardo & Rocchi, 2011Copelatus miaowangi Hajek, Jiang & Jia, 2022Copelatus minimus Bilardo & Rocchi, 1995Copelatus minor Bilardo & Pederzani, 1978Copelatus minutissimus J.Balfour-Browne, 1939Copelatus mocquerysi Régimbart, 1895Copelatus mohelicus Guignot, 1959Copelatus monticola Guignot, 1951Copelatus montivagus Young, 1942Copelatus mopanshanensis Jiang, Zhao & Hajek, 2022Copelatus mulangensis Bameul, 2003Copelatus mundus Sharp, 1882Copelatus mutabilis Bilardo & Rocchi, 1999Copelatus mvoungensis Bilardo & Rocchi, 2004Copelatus mysorensis Vazirani, 1970Copelatus nakamurai Guéorguiev, 1970Copelatus nangaensis Guignot, 1952Copelatus nanlingensis Hajek, Jiang & Jia, 2022Copelatus nauclerus Guignot, 1939Copelatus neavei J.Balfour-Browne, 1950Copelatus neelumae Vazirani, 1973Copelatus neogestroi Balke, 2008Copelatus neoguineensis Zimmermann, 1919Copelatus nguemai Bilardo & Rocchi, 2011Copelatus nigricans Sharp, 1882Copelatus nigricollis Paulino de Oliveira, 1880 Copelatus nigrolineatus Sharp, 1882Copelatus nigropennis Zimmermann, 1927Copelatus nigrostriatus Régimbart, 1895Copelatus nilssoni Wewalka, 1982Copelatus nimbaensis Legros, 1958Copelatus nitens Bilardo & Rocchi, 1999Copelatus nitidus Sharp, 1882Copelatus nodieri Régimbart, 1895Copelatus normalis Erichson, 1847Copelatus notius Omer-Cooper, 1965Copelatus nzei Bilardo & Rocchi, 1999Copelatus oblitus Sharp, 1882Copelatus obscurus Sharp, 1882Copelatus occultus Bilardo & Rocchi, 1995Copelatus ogasawarensis Kamiya, 1932Copelatus onorei Pederzani & Rocchi, 1982Copelatus ornatipennis Zimmermann, 1926Copelatus owas Régimbart, 1895Copelatus pachys Guignot, 1955Copelatus pallidus Régimbart, 1895Copelatus paludorum Guignot, 1952Copelatus pandanorum Scott, 1912Copelatus panguana Hájek, Alarie, Benetti, Hamada, Springer, Hendrich, Villastrigo, Torres, Basantes & Balke, 2024[ 7] Copelatus pantosi Guignot, 1958Copelatus papuensis J.Balfour-Browne, 1939Copelatus parabaptus Guignot, 1955Copelatus parallelipipedus Régimbart, 1895Copelatus parallelus Zimmermann, 1920Copelatus pardii Rocchi, 1990Copelatus parisii Guignot, 1959Copelatus parumstriatus Gschwendtner, 1934Copelatus paryphes Guignot, 1955Copelatus pederzanii Bilardo & Rocchi, 1995Copelatus pereirai Guignot, 1955Copelatus peridinus Guignot, 1955Copelatus pinnifer Guignot, 1952Copelatus piriensis Omer-Cooper, 1965Copelatus platynotus Régimbart, 1895Copelatus polystrigus Sharp, 1882Copelatus ponomarenkoi Říha, 1974 Copelatus portior Guignot, 1956Copelatus posticatus (Fabricius, 1801) Copelatus poungai Bilardo & Rocchi, 2006Copelatus predaveterus Miller, 2003Copelatus prolixus Sharp, 1882Copelatus prolongatus Sharp, 1882Copelatus propino Guignot, 1960Copelatus propinquus Régimbart, 1895Copelatus proximus Sharp, 1882Copelatus pseudostriatus Ranarilalatiana & Bergsten, 2019Copelatus pulchellus (Klug, 1834) Copelatus pulicarius Régimbart, 1895Copelatus pumilus Régimbart, 1895Copelatus punctatus Bilardo & Rocchi, 1995Copelatus punctulatus Aubé, 1838Copelatus puzhelongi Jiang, Hajek & Jia, 2022Copelatus quadrisignatus Régimbart, 1877Copelatus quadristriatus Pederzani & Schizzerotto, 2017Copelatus racenisi Guignot, 1951Copelatus ragazzii Régimbart, 1887Copelatus regimbarti Branden, 1885Copelatus restrictus Sharp, 1882Copelatus rimosus Guignot, 1952Copelatus rivalis Guignot, 1952Copelatus rocchii Bilardo, 1982Copelatus romani Zimmermann, 1924Copelatus rosulae Hajek, Jiang & Jia, 2022Copelatus royi Legros, 1958Copelatus rubiginosus Guignot, 1954Copelatus ruficapillus Régimbart, 1895Copelatus ruteri Legros, 1954Copelatus saegeri Guignot, 1955Copelatus safiotra Ranarilalatiana & Bergsten, 2019Copelatus sahlbergi J.Balfour-Browne, 1939Copelatus sallaei Sharp, 1882Copelatus sanfilippoi Bilardo & Pederzani, 1972Copelatus saverii Pederzani & Schizzerotto, 2017Copelatus scalptus Guignot, 1954Copelatus scaphites Guignot, 1955Copelatus schaefferi Young, 1942Copelatus schereri Wewalka, 1981Copelatus schuhi Hendrich & Balke, 1998Copelatus scutatus Guignot, 1952Copelatus scytalotus Guignot, 1956Copelatus senegalensis Lagar & Hernando, 1991Copelatus sexstriatus Sharp, 1882Copelatus sharpi Branden, 1885Copelatus sibelaemontis Hájek, Hendrich, Hawlitschek & Balke, 2010Copelatus silvestrii Régimbart, 1903Copelatus simoni Régimbart, 1889Copelatus singularis Bilardo & Rocchi, 1995Copelatus sociennus J.Balfour-Browne, 1952Copelatus solitarius Sharp, 1882Copelatus sordidipennis Régimbart, 1895Copelatus speciosus Régimbart, 1892Copelatus spoliatus Guignot, 1955Copelatus stavropolitanus Říha, 1974 Copelatus stillicidicola Bilardo & Rocchi, 1995Copelatus striatellus Boheman, 1848Copelatus striaticollis Lucas, 1857Copelatus striatopterus Aubé, 1838Copelatus striatulus Aubé, 1838Copelatus strigipennis (Laporte, 1835) Copelatus strigosulus (Fairmaire, 1878) Copelatus strinatii Guignot, 1958Copelatus striolatus Peschet, 1917Copelatus stygis Guignot, 1958Copelatus subdeficiens Régimbart, 1902Copelatus subsimilis Guignot, 1958Copelatus substriatus Kirsch, 1873Copelatus subterraneus Guéorguiev, 1978Copelatus sudrei Bameul, 2003Copelatus sulcatus Sharp, 1882Copelatus sulcipennis (Laporte, 1835) Copelatus sumbawensis Régimbart, 1899Copelatus suppar Guignot, 1956Copelatus supplementaris Régimbart, 1895Copelatus sylvaticoides Bilardo & Rocchi, 2013Copelatus sylvaticus Guignot, 1952Copelatus takakurai Satô, 1985Copelatus tanaus Guignot, 1953Copelatus taprobanicus Wewalka & Vazirani, 1985Copelatus tenebrosis Regimbart, 1880Copelatus tenebrosus Régimbart, 1880Copelatus tengchongensis Hajek, Jiang & Jia, 2022Copelatus teranishii Kamiya, 1938Copelatus terminalis Sharp, 1882Copelatus ternatensis Régimbart, 1899Copelatus terzanii Bilardo & Saverio, 2016Copelatus thiriouxi Peschet, 1917Copelatus thrasys Guignot, 1952Copelatus tibialis Sharp, 1882Copelatus tinctor Guignot, 1954Copelatus togoensis Régimbart, 1895Copelatus tomokunii Satô, 1985Copelatus tondangoyei Bilardo & Rocchi, 2013Copelatus tostus J.Balfour-Browne, 1950Copelatus trifilis Guignot, 1958Copelatus triglyphus Guignot, 1955Copelatus trilobatus Régimbart, 1895Copelatus tschaga Bilardo & Pederzani, 1972Copelatus tucuchiensis J.Balfour-Browne, 1939Copelatus tulagicus Guignot, 1942Copelatus uludanuensis Hendrich & Balke, 1995Copelatus undecimstriatus Aubé, 1838Copelatus urceolus Hájek, Shaverdo, Hendrich & Balke, 2021Copelatus usagarensis Zimmermann, 1926Copelatus vagatus Guignot, 1952Copelatus vagestriatus Zimmermann, 1919Copelatus vagus Bilardo & Rocchi, 1995Copelatus validus Sharp, 1882Copelatus vancli Hájek, Hendrich & Balke, 2018Copelatus vanninii Bilardo & Rocchi, 1999Copelatus variegatus Régimbart, 1895Copelatus variistriatus Hájek, Shaverdo, Hendrich & Balke, 2021Copelatus venustus Bilardo & Rocchi, 1995Copelatus vigintistriatus Fairmaire, 1869Copelatus vigintisulcatus Régimbart, 1895Copelatus villiersi Guignot, 1950Copelatus virungaensis Bilardo, 1982Copelatus vitraci Legros, 1948Copelatus vivax Guignot, 1953Copelatus vokoka Ranarilalatiana & Bergsten, 2019Copelatus wallacei J.Balfour-Browne, 1939Copelatus waltoni J.Balfour-Browne, 1950Copelatus wayanadensis Manivannan & Madani, 2011Copelatus wewalkai Holmen & Vazirani, 1990Copelatus weymarni J.Balfour-Browne, 1947Copelatus xanthocephalus Régimbart, 1899Copelatus xiniudong Jiang, Hajek & Jia, 2022Copelatus yacumensis Guignot, 1957Copelatus yaguarete Scheers, 2018Copelatus yombensis Bilardo & Rocchi, 2015Copelatus zadiensis Bilardo & Rocchi, 1995Copelatus zanabato Manuel & Ramahandrison, 2022Copelatus zanatanensis Ranarilalatiana & Bergsten, 2019Copelatus zimmermanni Gschwendtner, 1934 Copelatus Sheth, Ghate & Hájek, 2018
Coptotomus Say, 1830
Cretodytes Ponomarenko, 1977
Crinodessus K.B.Miller, 1997
大龙虱属 Cybister Curtis, 1827，真龙虱属
Darwinhydrus Sharp, 1882
Deronectes Sharp, 1882
Derovatellus Sharp, 1882
Desmopachria Babington, 1841
Deuteronectes Guignot, 1945
Deuteronectus
Dimitshydrus Uéno, 1996
Dyschirioides
Dyticus Geoffroy, 1762
Dytiscus Linnaeus, 1758
Electruphilus Balker, Toledo & Hendrich, 2019
Ereboporus K.B.Miller, Gibson & Alarie, 2009
Eretes Laporte, 1833
Etruscodytes Mazza, Cianferoni & Rocchi, 2013
Exocelina Broun, 1886
Fontidessus K.B.Miller & Spangler, 2008
Geodessus Brancucci, 1979
Gibbidessus Watts, 1978
Glareadessus Wewalka & Biström, 1998
Graphoderus Dejean, 1833
Graptodytes Seidlitz, 1887
Haideoporus Young & Longley, 1976
Hamadiana Benetti, Short & Michat, 2019
Hemibidessus Zimmermann, 1921
Heroceras Guignot, 1950
Heterhydrus Fairmaire, 1869
Heterosternuta Strand, 1935
Hintonia Young, 1980
Hoperius Fall, 1927
Hornectes Fery & Ribera, 2018
Hovahydrus Biström, 1982
Huxelhydrus Sharp, 1882
Hydaticus Leach, 1817
Hyderodes Hope, 1838
Hydrocolus Roughley & Larson, 2000
Hydrodessus J.Balfour-Browne, 1953
Hydrodytes K.B.Miller, 2001
短褶龙虱属 Hydroglyphus Motschulsky, 1853
Hydronebrius Jakovlev, 1897
Hydropeplus Sharp, 1882
Hydroporus Clairville, 1806
Hydroporus Schellenberg, 1806
Hydrotrupes Sharp, 1882
Hydrovatus Motschulsky, 1853
水龙虱属 Hygrotus Stephens, 1828
Hyphoporus Sharp, 1882
Hyphovatus Wewalka & Biström, 1994
异爪龙虱属 Hyphydrus Illiger, 1802
Hypodessus Guignot, 1939
Iberonectes Fery & Ribera, 2018
Iberoporus Castro & Delgado, 2001
Ilybiosoma Crotch, 1873
Ilybius Erichson, 1832
Incomptodessus K.B.Miller & García, 2011
Isonotus Houlbert, 1934
Japanolaccophilus Satô, 1972
Kakadudessus Hendrich & Balke, 2009
Kakadudessus tomweiri Hendrich & Balke, 2009 Kuschelydrus Ordish, 1976
Laccodytes Régimbart, 1895
Laccomimus Toledo & Michat, 2015
Lacconectus Motschulsky, 1855
粒龙虱属 Laccophilus Leach, 1815
Laccoporus J.Balfour-Browne, 1939
Laccoporus nigritulus (Gschwendtner, 1936) Laccornellus Roughley & Wolfe, 1987
Laccornis Gozis, 1914
Laccosternus Brancucci, 1983
Lancetes Sharp, 1882
Laodytes Queney, Lemaire & Ferrand, 2020
Larsonectes Fery & Ribera, 2018
Leconectes Fery & Ribera, 2018
Leiodytes Guignot, 1936
Leuronectes Sharp, 1882
Liadyxianus Prokin, Petrov, Wang & Ponomarenko, 2013
Limbodessus Guignot, 1939
Liodessus Guignot, 1939
Lioporeus Guignot, 1950
Liopterus Dejean, 1833
Madaglymbus Shaverdo & Balke, 2008
Matus Aubé, 1836
Megadytes Sharp, 1882
Megaporus Brinck, 1943
Meladema Laporte, 1835
Melanodytes Seidlitz, 1887
Meridiorhantus Balke, Hájek & Hendrich, 2017
Mesodytes Prokin, Petrov, Wang & Ponomarenko, 2013
Metaporus Guignot, 1945
Methles Sharp, 1882
Microdessus Young, 1967
Microdytes J.Balfour-Browne, 1946
Microhydrodytes K.B.Miller, 2002
Miodytiscus Wickham, 1911
Morimotoa Uéno, 1957
Mystonectes Fery & Ribera, 2018
Napodytes Steiner, 1981
Nartus Zaitzev, 1907
孔龙虱属 Nebrioporus Régimbart, 1906
Necterosoma Sharp, 1882
Necterosoma W.J.MacLeay, 1871
Nectoboreus Fery & Ribera, 2018
Nectomimus Fery & Ribera, 2018
Nectoporus Guignot, 1950
Neobidessodes Hendrich & Balke, 2009
Neobidessus Young, 1967
Neoclypeodytes Young, 1967
Neonectes J.Balfour-Browne, 1944
Neonectes Zimmermann, 1932
Neoporus Guignot, 1931
Neoscutopterus J.Balfour-Browne, 1943
彩裳粒龙虱属 Neptosternus Sharp, 1882
Notaticus Zimmermann, 1928
Novadessus K.B.Miller, 2016
Onychohydrus Schaum & White, 1847
Oreodytes Seidlitz, 1887
Pachydrus Sharp, 1882
Pachynectes Régimbart, 1903
Palaeodytes Ponomarenko, 1987
Papuadessus Balke, 2001
Paroster Sharp, 1882
Peschetius Guignot, 1942
Petrodessus K.B.Miller, 2012
Philaccolilus Guignot, 1937
Philaccolus Guignot, 1937
Philodytes J.Balfour-Browne, 1939
Phreatodessus Ordish, 1976
Platambus Thomson, 1859
Platydytes Biström, 1988
Platynectes Gueorguiev, 1972
Platynectes Régimbart, 1879
Porhydrus Guignot, 1945
Primospes Sharp, 1882
Procoelambus Théobald, 1937
Prodaticus Sharp, 1882
缝龙虱属 Pseuduvarus Biström, 1988
Psychopomporus Jean, Telles & K.B.Miller, 2012
Pteroporus Guignot, 1933
Queda Sharp, 1882
Regimbartina Chatanay, 1911
Rhantaticus Sharp, 1880
Rhantus Dejean, 1833
Rhithrodytes Bameul, 1989
Rompindessus Balke, Bergsten & Hendrich, 2017
Sandracottus Sharp, 1882
Sanfilippodytes Franciscolo, 1979
Scarodytes Gozis, 1914
Schistomerus Palmer, 1957
Sekaliporus Watts, 1997
Sharphydrus Omer-Cooper, 1958
Siamoporus Spangler, 1996
Siettitia Abeille de Perrin, 1904
Sinodytes Spangler, 1996
Spanglerodessus K.B.Miller & García, 2011
Spencerhydrus Sharp, 1882
Sternhydrus Brinck, 1945
Sternopriscus Sharp, 1880
Stictonectes Brinck, 1943
Stictotarsus Zimmermann, 1919
Stygoporus Larson & LaBonte, 1994
Suphisellus Zimmermann, 1921
Tassilodytes Fery & Bouzid, 2016
Tepuidessus Spangler, 1981
Thermonectus Dejean, 1833
Thermonectus alfredi Griffini, 1898Thermonectus basillaris (Harris, 1829)Thermonectus batesi Sharp, 1882Thermonectus circumscriptus (Latreille, 1809)Thermonectus cuneatus Sharp, 1882 Thermonectus depictus Sharp, 1882 Thermonectus duponti (Aubé, 1838)Thermonectus intermedius Crotch, 1873 Thermonectus laporti (Aubé, 1838)Thermonectus leprieuri J.Balfour-Browne, 1944Thermonectus margineguttatus (Aubé, 1838)Thermonectus marmoratus (Gray, 1831)Thermonectus nigrofasciatus (Aubé, 1838)Thermonectus nobilis Zimmermann, 1924Thermonectus sibleyi Goodhue-McWilliams, 1981Thermonectus simulator Sharp, 1882Thermonectus succinctus (Aubé, 1838)Thermonectus tremouillesi Michat & Torres, 2016[ 8] Thermonectus variegatus (Dejean, 1836)Thermonectus variegatus (Laporte de Castelnau, 1835)Thermonectus zimmermani Goodhue-McWilliams, 1981 Tikoloshanes Omer-Cooper, 1956
Tiporus Watts, 1985
Trichonectes Guignot, 1941
Trogloguignotus Sanfilippo, 1958
Tyndallhydrus Sharp, 1882
Typhlodessus Brancucci, 1985
Uvarus Guignot, 1939
Vatellus Aubé, 1837
Yola Gozis, 1886
Yolina Guignot, 1936
Zaitzevhydrus Fery & Ribera, 2018
Zimpherus K.B.Miller & Wheeler, 2015
G.N. Foster; D.T. Bilton. The Conservation of Predaceous Diving Beetles: Knowns, Unknowns and Anecdotes. D.A. Yee (编). Ecology, Systematics, and the Natural History of Predaceous Diving Beetles (Coleoptera: Dytiscidae). 2014: 437–462. ISBN 978-94-017-9109-0 Miller, Kelly; Bergsten, Johannes. Diving Beetles of the World: Systematics and Biology of the Dytiscidae. Baltimore: Johns Hopkins University Press. 3 October 2016: 20.
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Jiří Hájek, Yves Alarie, Cesar J. Benetti, Neusa Hamada, Monika Springer, Lars Hendrich, Adrián Villastrigo, Rodulfo Ospina Torres, Michael S. Basantes and Michael Balke. 2024. Underestimated Diversity and Range Size of Diving Beetles in Tank Bromeliads—Coleoptera of ‘hygrofloric’ lifestyle (Dytiscidae). Zoological Journal of the Linnean Society. 200(3); 720–735. DOI: 10.1093/zoolinnean/zlad093
Mariano C. Michat & Patricia L. M. Torres (2016) Thermonectus tremouillesi sp. nov. (Coleoptera: Dytiscidae: Aciliini): description of the adults and larvae and comparisons with other species of the genus, Journal of Natural History, 50:25-26, 1633-1648, DOI: 10.1080/00222933.2016.1145274
联邦科学与工业研究组织 (CSIRO)Family Dytiscidae （页面存档备份 ，存于互联网档案馆 ）. Version of 2004-JUL-02. Retrieved 2008-AUG-04De Foliart, Gene R. (2002): Chapter 26 - Eastern Asia: China, Japan, and other countries . In: The Human Use of Insects as a Food Resource: A Bibliographic Account in Progress .Jäch, Manfred A. (2003): Fried water beetles Cantonese style. American Entomologist 49 (1): 34-37. PDF fulltext Larson, D.J., Alarie, Y., and Roughley, R.E. (2000): Predaceous Diving Beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. NRC Research Press, Ottawa. ISBN 978-0-660-17967-4 . 
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