鱉甲蝸牛科

鱉甲蝸牛科
	Five worn shells of Erepta setiliris at Réunion island
科學分類
界：	動物界 Animalia
門：	軟體動物門 Mollusca
綱：	腹足綱 Gastropoda
目：	柄眼目 Stylommatophora
亞目：	旋蝸牛亞目 Helicina
下目：	蛞蝓下目 Limacoidei
總科：	勇蝸總科 Helicarionoidea
科：	鱉甲蝸牛科 Helicarionidae; Bourguignat, 1877

鱉甲蝸牛科（學名：Helicarionidae），又名勇蝸科，鱉甲螺科^[2]，是一個由陸地上的蝸牛及半蛞蝓（英語：Semi-slug）組成的科，皆為真有肺類柄眼類支序的成員，屬於鱉甲蝸牛總科（Helicarionoidea）^[3]。

快速預覽 鱉甲蝸牛科, 科學分類 ...

關閉

本科物種分佈於古北界東部，馬爾加什（馬達加斯加、留尼汪及科摩羅）、印度、東南亞及澳大利亞^[4]。

本科的蝸牛都會在交配時伸出幾丁質的交配器戀矢（love dart）插進對方的生殖孔，以交換精子^[5]。

此外，本科物種的單倍體染色體介乎21到30之間^[6]。

本科包括以下屬：

Advena Gude, 1913
Aenigmatoconcha
C.Tumpeesuwan & S.Tumpeesuwan, 2017
- Aenigmatoconcha clivicola C.Tumpeesuwan & S.Tumpeesuwan, 2017^[7]
- Aenigmatoconcha eunetis Pholyotha & Panha, 2021^[8]
- Aenigmatoconcha mitis (L.Pfeiffer, 1863)
- Aenigmatoconcha sumonthai C.Tumpeesuwan & S.Tumpeesuwan, 2018
Amenixesta
Iredale, 1941
- Amenixesta fifensis Iredale, 1941
- Amenixesta hunsteini (E.A.Smith, 1887)
Antiquarion
Stanisic, 2010
- Antiquarion boondah Hyman & Köhler, 2020
- Antiquarion chooreechillum Hyman & Köhler, 2020
- Antiquarion collinsae J.Stanisic, 2010
- Antiquarion evelynensis Hyman & Köhler, 2020
- Antiquarion limbus Hyman & Köhler, 2020
- Antiquarion occultus Hyman & Köhler, 2020
- Antiquarion ravenshoe Hyman & Köhler, 2020
- Antiquarion swausi Hyman & Köhler, 2020
Apothapsia D.T.Holyoak & G.A.Holyoak, 2020
Attenborougharion Hyman & Köhler, 2017
奧氏蛞蝓屬 Austenia G.Nevill, 1878
Bathia Robson, 1914
Belopygmaeus Solem, 1989
Belopygmeus Baker, 1938
Brevisentis Hyman, 2007
Buffetia Iredale, 1945
Burmochlamys Pholyotha & Panha, 2022
Burnettia Stanisic, 2010
Caldwellia H.Adams, 1873
Caracolla Ehrenberg, 1831
疑難螺屬 Chalepotaxis
Ancey, 1887
- 黑線小鱉甲蝸牛 Chalepotaxis spadix cinctus
Colmanarion Stanisic, 2010
隱奧氏蛞蝓屬 Cryptaustenia Cockerell, 1891
Ctenoglypta Ancey, 1904
Ctenophila Ancey, 1882
Cucullarion Stanisic, 1998
Dancea Zilch, 1960
Dedrolamellaria
Delinitesta Iredale, 1933
Dendrolamellaria Preston, 1913
Dendronitor Iredale, 1933
Dupontia Godwin-Austen, 1908
小杜螺屬 Durgella
W.T.Blanford, 1863
- Durgella aborense Godwin-Austen, 1916
- Durgella assamica Gondwin-Austen, 1881
- Durgella birmanica (Pfeiffer, 1847)
- Durgella christianae (Theobald, 1864)
- Durgella concinna Blanford & Godwin-Austen, 1908
- Durgella densestriata Vermeulen, T.-S.Liew & Schilthuizen, 2015
- Durgella dwipana (Gude, 1903)
- Durgella edeana Godwin-Austen, 1910
- Durgella erratica (Godwin-Austen, 1888)
- Durgella hosei Godwin-Austen, 1891
- Durgella khasiaca Godwin-Austen, 1883
- Durgella kumahensis (Theobald & Stoliczka, 1872)
- Durgella levicula (W.H.Benson, 1859)
- Durgella libas Solem, 1966
- Durgella mairangensis Godwin-Austen, 1898
- Durgella naharaniensis Godwin-Austen, 1914
- Durgella nulla Pholyotha & Panha, 2024^[9]
- Durgella pentata Pholyotha & Panha, 2024^[9]
- Durgella rhaphiellus (E.von Martens, 1867)
- Durgella rogersi Godwin-Austen, 1907
- Durgella russeola (Morelet, 1865)
- Durgella salius (W.H.Benson, 1859)
- Durgella seposita (W.H.Benson, 1859)
- Durgella siamensis Möllendorff, 1902
- Durgella sundana B.Rensch, 1930
Durgellina Thiele, 1928
Echonitor Iredale, 1937
Eddiella Stanisic, 2010
Einasleighana Stanisic, 2010
Elatonitor Stanisic, 2010
Ellarion Iredale, 1941
Epiglypta Pilsbry, 1893
Erepta Albers, 1850
Eufretum H.B.Baker, 1941
Eurychlamys Godwin-Austen, 1899
Expocystis Iredale, 1937
Fastosarion Iredale, 1933
Fenestrarion Stanisic, 2010
Fijia Gude, 1913
吉拉蛞蝓屬 Girasia Gray, 1855
Gudeconcha
Gudeoconcha Iredale, 1944
Haleakala Baker, 1940
Harmogenanina Germain, 1918
勇蝸屬 Helicarion Férussac, 1821
Hiloaa Baker, 1940
Hiona Cooke, 1940
Hionarion Baker, 1940
Hionella Baker, 1940
Holkeion Godwin-Austen, 1908
Howearion Iredale, 1944
伊比蛞蝓屬 Ibycus
Heynemann, 1863
- 小玉蝸牛 Ibycus hiraseanus
Insulorbis Baker, 1940
Irenella Gude, 1913
Kalidos Gude, 1911
Kermarion Iredale, 1944
Kipua Baker, 1940
Laconia Gray, 1855
Laocaia Kuzminykh, 1999
Levidens Hyman, 2007
Limpidarion Hyman & Köhler, 2020
Luinarion Iredale, 1933
Malandena Iredale, 1933
Mauka Baker, 1940
Medyla Martens, 1860
Minororbis Baker, 1940
Mistarion Iredale, 1941
Montanocystis Stanisic, 2010
Muangnua Solem, 1966
Mysticarion Iredale, 1941
Nesaecia Gude, 1911
Nesarion Baker, 1940
Nesocyclus Baker, 1940
Nesonanina C.R.Boettger, 1918
Neutra Baker, 1940
Nitor Gude, 1911
Nukupiena Baker, 1940
Oafatua Baker, 1940
Opara Baker, 1940
圓鱉甲蝸牛屬 Ovachlamys Habe, 1946
Pachystyla Mörch, 1852
Palmervillea Stanisic, 2010
Panulena
Parmacochlea E.A.Smith, 1884
Parmavitrina Iredale, 1937
Parmella H.Adams, 1867
Parmellops Iredale, 1944
Peloparion Iredale, 1937
Periclocystis Iredale, 1937
扁楯蛞蝓屬 Petalochlamys Godwin-Austen, 1907，葉楯蛞蝓屬
Pitcairnia Baker, 1940
Pittoconcha Preston, 1913
Plegma Gude, 1911
Pravonitor Iredale, 1937
Pseudaustenia Cockerell, 1891
Pseudoguppya Baker, 1925
Pseudophasis Germain, 1918
Pseudosaphtia de Winter, 2008
Pseudosesara Solem, 1962
Quirosella Clench, 1958
Rapafila Baker, 1940
曙螺屬 Rasama Laidlaw, 1932
Rhadella Godwin-Austen, 1914
Rhyssotopsis Ancey, 1887
若通螺屬 Rotungia Godwin-Austen, 1918
Saphtia de Winter, 2008
Satiella W.T.Blanford & Godwin-Austen, 1908
Sesara Martens, 1860
Sheaia Hyman, 2007
Sigaloeista Shea & Griffiths, 2010
昔塔螺屬 Sitala H.Adams, 1865
Sitalarion H.B.Baker, 1941
Sivella W.T.Blanford, 1863
Sodaleta Iredale, 1937
Sophina Benson, 1859
Stanisicarion Hyman & Ponder, 2010
Tarocystis Iredale, 1937
Teraia Solem, 1966
Thularion Stanisic, 1993
Torrecystis Stanisic, 2010
Tropicystis Stanisic, 2010
Uafatua Baker, 1940
Uapuneutra Baker, 1940
Ubiquitarion Hyman, Lamborena & Köhler, 2017
Urazirochlamys Habe, 1946
Vanmolia de Winter, 2008
Vercularion Iredale, 1933
Waihoua Baker, 1940
Westracystis Iredale, 1939
Zagmena Iredale, 1941

The following two subfamilies were recognized in the taxonomy of Bouchet & Rocroi (2005):^[3]

亞科 Helicarioninae Bourguignat, 1877 - synonyms: Pseudotrochatellinae A. J. Wagner, 1905;^[10] Ereptinae Godwin-Austen, 1908;^[11] Xestinae Gude & B. B. Woodward, 1921;^[12] Sesarinae Thiele, 1931;^[13] Nitoridae Iredale, 1937;^[14] Epiglyptidae Iredale, 1944;^[15] Gudeoconchidae Iredale, 1944^[15]
亞科 Durgellinae Godwin-Austen, 1888^[16]
- 族 Durgellini Godwin-Austen, 1888 - synonyms: Sitalinae Godwin-Austen, 1900;^[17] Sophininae W. Blanford & Godwin-Austen, 1908;^[18] Satiellini Schileyko, 2003^[19]
- 族 Girasiini Collinge, 1902^[20]

屬

下列本科所有的屬：

Helicarioninae亞科

Epiglypta Pilsbry, 1893^[3]
Erepta Albers, 1850^[3]
Gudeoconcha Iredale, 1914^[3]
Helicarion Férussac, 1821 - type genus of the family Helicarionidae^[3]
Nitor Gude, 1911^[3]
Pseudotrochatella G. Nevill, 1881^[3]
Sesara Albers, 1860^[3]
Wilhelminaia Preston, 1913 ^[21]
Xesta Albers, 1850^[3]

Durgellinae亞科

族 Durgellini

Durgella Blanford, 1863 - type genus of the subfamily Durgellinae^[3]
Satiella Godwin-Austen, 1908^[3]
Sitala H. Adams, 1865^[3]
Sophina Benson, 1859^[3]

族 Girasiini

Girasia Gray, 1855 - 本族的模式屬^[3]

亞科未定

Advena
Atrichotoxon
Buffetia
Diastole
Dolapex
Elisolimax
Harmogenanina
Iredaleoconcha
恰里螺屬 Kaliella
Lutilodix
Mathewsoconcha
Mysticarion
Nancibella
圓鱉甲蝸牛屬 Ovachlamys Habe, 1946，卵楯蛞蝓屬
Panulena
Philonesia
Pittoconcha
Quintalia
Ryssota
Thapsia
Trichotoxon
Zingis

以下的系統發生學分支圖顯示了根據Hausdorf (2000)所描述蛞蝓下目各科之間的親緣關係^[4]：

蛞蝓下目 Limacoidei

Staffordioidea（英語：Staffordioidea）

Staffordiidae

Dyakioidea（英語：Dyakioidea）

Dyakiidae

Gastrodontoidea（英語：Gastrodontoidea）

Pristilomatidae（英語：Pristilomatidae）

克氏螺科 Chronidae

	真蛞蝓科 Euconulidae

	Trochomorphidae（英語：Trochomorphidae）

	腹齒螺科 Gastrodontidae

	Oxychilidae（英語：Oxychilidae）

Parmacelloidea（英語：Parmacelloidea）

	Trigonochlamydidae（英語：Trigonochlamydidae）

	Parmacellidae（英語：Parmacellidae）

	Milacidae（英語：Milacidae）

帶螺總科 Zonitoidea

帶螺科 Zonitidae

鱉甲蝸牛總科（英語：Helicarionoidea） Helicarionoidea（英語：Helicarionoidea）

鱉甲蝸牛科 Helicarionidae

	擬阿勇蛞蝓科 Ariophantidae

	Urocyclidae（英語：Urocyclidae）

Limacoidea（英語：Limacoidea）

Vitrinidae（英語：Vitrinidae）

Boettgerillidae（英語：Boettgerillidae）

	蛞蝓科 Limacidae

	野蛞蝓科 Agriolimacidae

[1]
Bourguignat J. R. (1877). "Description de deux nouveaux genres algériens, suive d'une classification des families et des genres de Mollusques terrestres et fluviatiles du système européen". Bulletin de la Société des Sciences Physiques et Naturelles de Toulouse 3(1): 49-101. page 64.
[2]
施乃普. 綠島小形貝殼. 中國貝誌 (中華民國貝類學會). 1975, 2: 33–46 [2018-08-28] （中文（繁體））.
[3]
Bouchet, P.; Rocroi, J.-P.; Frýda, J.; Hausdorf, B.; Ponder, W.; Valdés, Á. & Warén, A. Classification and nomenclator of gastropod families. Malacologia (Hackenheim, Germany: ConchBooks). 2005, 47 (1–2): 1–397. ISBN 3-925919-72-4. OCLC 61351079. OL 42989175M. S2CID 83847814.
[4]
Hausdorf, B. Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal. Journal of Biogeography. 2000, 27 (2): 379–390. doi:10.1046/j.1365-2699.2000.00403.x （英語）.
[5]
記用蝸牛來作實驗. 2013-07-19 （中文（繁體））.
[6]
G. M. Barker. The Biology of Terrestrial Molluscs. CABI. 2001: 139. ISBN 978-0-85199-318-8.
[7]
Chanidaporn Tumpeesuwan and Sakboworn Tumpeesuwan. 2017. Discovery of An Overlooked Helicarionid Land Snail (Helicarionidae: Durgellinae) from northeastern Thailand, with Description of A New Genus and New Species, and Note on Radula Morphology and Genital System. RAFFLES BULLETIN OF ZOOLOGY. 65; 181–188.
[8]
Arthit Pholyotha, Chirasak Sutcharit, Piyoros Tongkerd and Somsak Panha. 2021. Systematic Revision of the Limestone Karst-restricted Land Snail Genus Aenigmatoconcha (Eupulmonata: Helicarionidae), with Description of A New Species. European Journal of Taxonomy. 767(1), 55-82. DOI: 10.5852/ejt.2021.767.1487
[9]
Arthit Pholyotha, Chirasak Sutcharit, Ngwe Lwin and Panha Somsak. 2024. Unveiling the Diversity of the Semislug Gnus Durgella Blanford, 1863 (Eupulmonata: Helicarionidae) from Thailand and Myanmar, with Description of Two New Species. Zool Stud. 63:14. DOI: 10.6620/ZS.2024.63-14.
[10]
Wagner A. J. (1905). Denkschriften der Mathematisch-Naturwissenschaftlichen Klasse der Kaiserlichen Akademie der Wissenschaften 77: 365.
[11]
Godwin-Austen H. H. (1908). "On the Animals of Genera and Species of Mascarene Land-Mollusca belonging to the family Zonitidae, collected by Monsieur E. Dupont". Annals and Magazine of Natural History (8)2: 422-436. 432. Plate 9-11.
[12]
Gude G. P. L. K. & Woodward B. B. (1921). "On Helicella, Férussac". Proceedings of the Malacological Society of London 14(5-6): 174-190. page 185.
[13]
Thiele (1931). Handbuch der systematischen Weichtierkunde 1(2): 620.
[14]
Iredale T. (1937). The Australian Zoologist 9(1): 2.
[15]
Iredale T. (1944). The Australian Zoologist 10(3): page 326 and page 328.
[16]
Godwin-Austen H. H. (1888). Land and freshwater Mollusca of India 1(6): 253.
[17]
Godwin-Austen H. H. (1900). In: Sykes Mollusca. In: Sharp (ed.) Fauna Hawaiiensis 2(4): 283.
[18]
Blanford W. T. & Godwin-Austen H. H. (1908). The fauna of British India. Mollusca. Testacellidae and Zonitidae, page 283.
[19]
Schileyko (2003). Treatise on Recent terrestrial pulmonate molluscs, part 10: 1319.
[20]
Collinge (1902). The Journal of Malacology 9(3): 71, 73.
[21]
{{cite journal |last=Ho |first=Wai Hoong |title=A Review Of The Land-Snail Fauna of Singapore |journal=[[The Raffles Bulletin of Zoology ||year=1995 |volume=43 |issue=1 |pages=91-113 |url=http://malaypeninsularsnail.myspecies.info/node/30 |language=en |access-date=2017-02-16 |archive-date=2017-02-16 |archive-url=https://web.archive.org/web/20170216224248/http://malaypeninsularsnail.myspecies.info/node/30 |dead-url=no }}

維基物種上的相關資訊：鱉甲蝸牛科
Helicarionidae. NCBI （英語）.

Video of a Helicarionid in Sulawesi （頁面存檔備份，存於互聯網檔案館）

[1] [1]
Bourguignat J. R. (1877). "Description de deux nouveaux genres algériens, suive d'une classification des families et des genres de Mollusques terrestres et fluviatiles du système européen". Bulletin de la Société des Sciences Physiques et Naturelles de Toulouse 3(1): 49-101. page 64.

[shelldoc_000027-2] [2]
施乃普. 綠島小形貝殼. 中國貝誌 (中華民國貝類學會). 1975, 2: 33–46 [2018-08-28] （中文（繁體））.

[Bouchet_2005-3] [3]
Bouchet, P.; Rocroi, J.-P.; Frýda, J.; Hausdorf, B.; Ponder, W.; Valdés, Á. & Warén, A. Classification and nomenclator of gastropod families. Malacologia (Hackenheim, Germany: ConchBooks). 2005, 47 (1–2): 1–397. ISBN 3-925919-72-4. OCLC 61351079. OL 42989175M. S2CID 83847814.

[Hausdorf_2000-4] [4]
Hausdorf, B. Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal. Journal of Biogeography. 2000, 27 (2): 379–390. doi:10.1046/j.1365-2699.2000.00403.x （英語）.

[5] [5]
記用蝸牛來作實驗. 2013-07-19 （中文（繁體））.

[Barker-6] [6]
G. M. Barker. The Biology of Terrestrial Molluscs. CABI. 2001: 139. ISBN 978-0-85199-318-8.

[7] [7]
Chanidaporn Tumpeesuwan and Sakboworn Tumpeesuwan. 2017. Discovery of An Overlooked Helicarionid Land Snail (Helicarionidae: Durgellinae) from northeastern Thailand, with Description of A New Genus and New Species, and Note on Radula Morphology and Genital System. RAFFLES BULLETIN OF ZOOLOGY. 65; 181–188.

[8] [8]
Arthit Pholyotha, Chirasak Sutcharit, Piyoros Tongkerd and Somsak Panha. 2021. Systematic Revision of the Limestone Karst-restricted Land Snail Genus Aenigmatoconcha (Eupulmonata: Helicarionidae), with Description of A New Species. European Journal of Taxonomy. 767(1), 55-82. DOI: 10.5852/ejt.2021.767.1487

[Pholyotha-9] [9]
Arthit Pholyotha, Chirasak Sutcharit, Ngwe Lwin and Panha Somsak. 2024. Unveiling the Diversity of the Semislug Gnus Durgella Blanford, 1863 (Eupulmonata: Helicarionidae) from Thailand and Myanmar, with Description of Two New Species. Zool Stud. 63:14. DOI: 10.6620/ZS.2024.63-14.

[10] [10]
Wagner A. J. (1905). Denkschriften der Mathematisch-Naturwissenschaftlichen Klasse der Kaiserlichen Akademie der Wissenschaften 77: 365.

[11] [11]
Godwin-Austen H. H. (1908). "On the Animals of Genera and Species of Mascarene Land-Mollusca belonging to the family Zonitidae, collected by Monsieur E. Dupont". Annals and Magazine of Natural History (8)2: 422-436. 432. Plate 9-11.

[12] [12]
Gude G. P. L. K. & Woodward B. B. (1921). "On Helicella, Férussac". Proceedings of the Malacological Society of London 14(5-6): 174-190. page 185.

[13] [13]
Thiele (1931). Handbuch der systematischen Weichtierkunde 1(2): 620.

[14] [14]
Iredale T. (1937). The Australian Zoologist 9(1): 2.

[Iredale_1944-15] [15]
Iredale T. (1944). The Australian Zoologist 10(3): page 326 and page 328.

[16] [16]
Godwin-Austen H. H. (1888). Land and freshwater Mollusca of India 1(6): 253.

[17] [17]
Godwin-Austen H. H. (1900). In: Sykes Mollusca. In: Sharp (ed.) Fauna Hawaiiensis 2(4): 283.

[18] [18]
Blanford W. T. & Godwin-Austen H. H. (1908). The fauna of British India. Mollusca. Testacellidae and Zonitidae, page 283.

[19] [19]
Schileyko (2003). Treatise on Recent terrestrial pulmonate molluscs, part 10: 1319.

[20] [20]
Collinge (1902). The Journal of Malacology 9(3): 71, 73.

[21] [21]
{{cite journal |last=Ho |first=Wai Hoong |title=A Review Of The Land-Snail Fauna of Singapore |journal=[[The Raffles Bulletin of Zoology ||year=1995 |volume=43 |issue=1 |pages=91-113 |url=http://malaypeninsularsnail.myspecies.info/node/30 |language=en |access-date=2017-02-16 |archive-date=2017-02-16 |archive-url=https://web.archive.org/web/20170216224248/http://malaypeninsularsnail.myspecies.info/node/30 |dead-url=no }}

[2]

[3]

[1]

[4]

[5]

[6]

[7]

[8]

[9]

[10]

[11]

[12]

[13]

[14]

[15]

[16]

[17]

[18]

[19]

[20]

[21]

屬

Helicarioninae亞科

Durgellinae亞科

亞科未定

Wikiwand in your browser!

鱉甲蝸牛科

屬

Helicarioninae亞科

Durgellinae亞科

亞科未定

Wikiwand in your browser!

分佈

解剖

下級分類

親緣支序圖

參考文獻

外部連結

鱉甲蝸牛科

Five worn shells of Erepta setiliris at Réunion island
科學分類
界：	動物界 Animalia
門：	軟體動物門 Mollusca
綱：	腹足綱 Gastropoda
目：	柄眼目 Stylommatophora
亞目：	旋蝸牛亞目 Helicina
下目：	蛞蝓下目 Limacoidei
總科：	勇蝸總科 Helicarionoidea
科：	鱉甲蝸牛科 Helicarionidae Bourguignat, 1877^[1]