Los botremídidos (Bothremydidae) son una familia extinta de tortugas que esconden su cuello lateralmente. Estas pertenecen al suborden Pleurodira y al orden Testudines. Las botremídidas vivieron en ecosistemas de agua dulce y eran anfibias. Eran al parecer omnívoros activos.
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La familia se divide en dos subfamilias y un cierto número de tribus.[1]
Bothremydidae
- Subfamilia Bothremydinae Gaffney et al., 2006
- Tribu Bothremydini Gaffney et al., 2006
- Iberoccitanemys Pérez-García et al., 2012[2]
- Iberoccitanemys convenarum Pérez-García et al., 2012
- Puentemys Cadena et al., 2012[3]
- Puentemys mushaisaensis Cadena et al., 2012
- Khargachelys AbdelGawad et al., 2023[4]
- Khargachelys caironensis AbdelGawad et al., 2023
- Subtribu Bothremydina Gaffney et al., 2006
- Araiochelys Gaffney et al., 2006
- Araiochelys hirayamai Gaffney et al., 2006
- Bothremys Leidy 1865 (sinónimo más moderno: Karkaemys)
- Bothremys arabicus (Zalmout et al., 2005)
- Bothremys cooki Leidy 1865
- Bothremys kellyi Gaffney et al., 2006
- Bothremys maghrebiana Gaffney et al., 2006
- Chedighaii Gaffney et al., 2006
- Chedighaii barberi (Schmidt 1940)
- Chedighaii hutchisoni Gaffney et al., 2006
- Rosasia Costa 1940
- Rosasia soutoi Carrington da Costa 1940
- Zolhafah Lapparent de Broin y Werner 1998
- Zolhafah bella Lapparent de Broin y Werner 1998
- Subtribu Foxemydina Gaffney et al., 2006
- Elochelys Nopsca 1931
- Elochelys convenarum Laurent et al., 2002
- Elochelys perfecta Nopsca 1931
- Foxemys Tong et al., 1998[5]
- Foxemys mechinorum Tong et al., 1998
- Foxemys trabanti Rabi, Tong & Botfalvai, 2012[6]
- Polysternon Portis, 1882
- Polysternon atlanticum Lapparent de Broin y Murelaga, 1996
- Polysternon isonae Marni et al., 2012[7]
- Polysternon provinciale (Matheron, 1869)
- Tribu Taphrosphyini Gaffney et al., 2006
- Subtribu Nigermydina Gaffney et al., 2006
- Arenila Lapparent de Broin y Werner 1998
- Arenila krebsi Lapparent de Broin ay Werner 1998
- Subtribu Taphrosphyina Gaffney et al., 2006
- Azabbaremys Gaffney et al., 2001
- Azabbaremys moragjonesi Gaffney et al., 2001
- Phosphatochelys Gaffney y Tong 2003
- Phosphatochelys tedfordi Gaffney y Tong 2003
- Rhothonemys Gaffney et al., 2006
- Rhothonemys brinkmani Gaffney et al., 2006
- Taphrosphys Cope 1869 (sinónimos más modernos: Amblypeza, Bantuchelys, Prochonias)
- Taphrosphys congolensis (Dollo 1912)
- Taphrosphys dares Hay 1908 (nomen dubium)
- Taphrosphys ippolitoi Gaffney et al., 2006
- Taphrosphys sulcatus (Leidy 1856)
- Ummulisani Gaffney et al., 2006
- Ummulisani rutgersensis Gaffney et al., 2006
- Crassachelys (nomen dubium)
- Crassachelys neurirregularis (Bergounioux 1952)
- Eusarkia (nomen dubium)
- Gafsachelys Stefano 1903 (nomen dubium)
- Labrostochelys Gaffney et al., 2006
- Labrostochelys galkini Gaffney et al., 2006
- Tribu Cearachelyini Gaffney et al., 2006
- Cearachelys Gaffney et al., 2001
- Cearachelys placidoi Gaffney et al., 2001
- Galianemys Gaffney et al., 2002
- Galianemys emringeri Gaffney et al., 2002
- Galianemys whitei Gaffney et al., 2002
- Subfamilia Kurmademydinae Gaffney et al., 2006
- Tribu Kurmademydini Gaffney et al., 2006
- Kinkonychelys Gaffney et al., 2009
- Kinkonychelys rogersi Gaffney et al., 2009
- Kurmademys Gaffney et al., 2001
- Kurmademys kallamedensis Gaffney et al., 2001
- Sankuchemys Gaffney et al., 2003
- Sankuchemys sethnai Gaffney et al., 2003
A continuación un cladograma basado en los estudios de Gaffney et al. in 2006:[8]
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Chelidae |
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Pelomedusoides |
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| Euraxemydidae |
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| Podocnemidioidea |
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Podocnemidinura |
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| Bothremydidae |
| Kurmademydini |
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| Bothremydinae |
| Cearachelyini |
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| Bothremyodda |
| Bothremydini |
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Zolhafah |
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Rosasia |
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Araiochelys |
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Chedighaii hutchisoni |
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Chedighaii barberi |
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Boremys cooki |
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Boremys magrhebiana |
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Boremys kellyi |
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Boremys arabicus |
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| Taphrosphyini |
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Azabbaremys |
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Labrostochelys |
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Taphrosphys |
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Rhothonemys |
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Phosphatochelys |
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Ummulisani |
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A. Perez-Garcia, F. Ortega, and X. Murelaga. 2012. A new genus of Bothremydidae (Chelonii, Pleurodira) in the Cretaceous of southwestern Europe. Geobios 45:219-229.
E. A. Cadena, J. I. Bloch, and C. A. Jaramillo. 2012. New bothremydid turtle (Testudines, Pleurodira) from the Paleocene of Northeastern Colombia. Journal of Paleontology 86(4):688-698.
Tong, Haiyan, Eugene S. Gaffney, Eric Buffetaut (1998). "Foxemys, a new side-necked turtle (Bothremydidae, Pelomedusoides) from the late Cretaceous of France". American Museum Novitates 3251: 1-19.
Rabi, M., Tong, H. & Botfalvai, G. 2012. A new species of the side-necked turtle Foxemys (Pelomedusoides: Bothremydidae) from the Late Cretaceous of Hungary and the historical biogeography of the Bothremydini. Geological Magazine, 149: 662-674.
J. Marmi, A. H. Luján, V. Riera, R. Gaete, and O. Oms Galobart. 2012. The youngest species of Polysternon: A new bothremydid turtle from the uppermost Maastrichtian of the southern Pyrenees. Cretaceous Research 35:133-142.
Gaffney, E.S.; Haiyan, T.; Meylan, P.A. (2006). «Evolution of the side-neck turtles: the families Bothremydidae, Euraxemydidae, Andararipemydidae». Bulletin of the American Museum of Natural History 300: 700.
