Umbra (fish)

Umbra is a genus of mudminnows native to Europe and North America. It is the only extant member of the family Umbridae.

Umbra Temporal range: Late Eocene to present PreꞒ Ꞓ O S D C P T J K Pg N
European mudminnow, Umbra krameri
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	Salmoniformes
Family:	Umbridae
Genus:	Umbra W. H. Kramer, 1777
Type species
Umbra krameri Walbaum, 1792

Phylogenetic evidence suggests the North American and European clades split between the Late Cretaceous or early Paleogene, making this a rather ancient group.^[1]

Species

The currently recognized species in this genus are:^[2]

• Umbra krameri Walbaum, 1792 (European mudminnow)
• Umbra limi (Kirtland, 1840) (central mudminnow)
• Umbra pygmaea (DeKay, 1842) (eastern mudminnow)

The following fossil species are also known:

• †Umbra euronota Schwarzhans et al., 2024 - Middle Miocene of Ukraine [otolith]^[3]
• †Umbra irtyshensis Sytchevskaya, 1968^[4]
• †Umbra longidorsalis Böhme, 2004 - Early Miocene of the Czech Republic (Most Formation)^[5]
• †Umbra oderiensis Oberhlová, 1978 - Late Oligocene/Early Miocene of the Czech Republic^[5]
• †Umbra praekrameri Weinfurter, 1950 - Late Miocene of Austria [otolith]^[3]
• †Umbra prochazkai Oberhlová, 1978 - Late Oligocene of the Czech Republic^[1][5]

Indeterminate Umbra scales are known from the Late Eocene/Early Oligocene of the Czech Republic.^[6]