Ulidiidae

The Ulidiidae (formerly Otitidae) or picture-winged flies are a large and diverse cosmopolitan family of flies (Diptera), and as in related families, most species are herbivorous or detritivorous. They are often known as picture-winged flies, along with members of other families in the superfamily Tephritoidea that have patterns of bands or spots on the wings. Some species share with the Tephritidae an unusual elongated posteroapical projection of the anal cell in the wing, but can be differentiated by the smoothly curving subcostal vein. Two species, Tetanops myopaeformis and Euxesta stigmatias, are agricultural pests.

Tropical Ulidiidae

Callopistromyia annulipes

Ulidiidae
Tritoxa incurva
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Diptera
Superfamily:	Tephritoidea
Family:	Ulidiidae Macquart, 1835
Subfamilies
Otitinae Aldrich, 1932[1] Ulidiinae Macquart, 1835
Synonyms
Ortalidae Otitidae

Systematics

The Ulidiidae are divided into two subfamilies.^[2]

Subfamily Otitinae

Tribe Cephaliini Schiner, 1864

• Acrostictella Hendel, 1914^[3]
• Cephalia Meigen, 1826^[4]
• Delphinia Robineau-Desvoidy, 1830^[5]
• Myiomyrmica Steyskal, 1961^[6]
• Myrmecothea Hendel, 1910^[7]
• Proteseia Korneyev & Hernandes, 1998^[8]
• Pterotaenia Rondani, 1868^[9]
• Tritoxa Loew, 1873^[10]

Tribe Myennidini Kameneva & Korneyev, 2006^[11]

• Acatochaeta Enderlein, 1921^[12]
• Arborotites Barraclough, 2000^[13]
• Callopistromyia Hendel, 1907^[14]
• Dyscrasis Aldrich, 1932^[1]
• Myennis Robineau-Desvoidy, 1830^[5]
• Namibotites Barraclough, 2000^[13]
• Neodyscrasis Kameneva & Korneyev, 2006^[11]
• Oedopa Loew, 1868^[15]
• Paroedopa Coquillett, 1900^[16]
• Pseudodyscrasis Hernández-Ortiz, 1988^[17]
• Pseudotephritina Malloch, 1931^[18]
• Pseudotephritis Johnson, 1902^[19]
• Stictoedopa Brèthes, 1926^[20]
• Stictomyia Bigot, 1885^[21]
• Ulidiotites Steyskal, 1961^[6]

Tribe Otitini Aldrich, 1932^[1]

• Ceroxys Macquart, 1835
• Dorycera Meigen, 1830
• Herina Robineau-Desvoidy, 1830^[5]
• Hiatus Cresson, 1906
• Melieria Robineau-Desvoidy, 1830^[5]
• Otites Latreille, 1804
• Tetanops Fallén, 1820
• Ulidiopsis Hennig, 1941

Otitinae incertae sedis

• Curranops Harriot, 1942
• Diacrita Gerstäcker, 1860
• Haigia Steyskal, 1961^[6]
• Idana Loew, 1873^[10]
• Psaeropterella Hendel, 1914
• Pseudomelieria Brèthes, 1921
• Tetropismenus Loew, 1876
• Tujunga Steyskal, 1961^[6]

Subfamily Ulidiinae

Tribe Lipsanini Enderlin, 1838

• Acrosticta Loew, 1868
• Acrostictomyia Blanchard, 1938
• Aspistomella Hendel, 1909
• Axiologina Hendel, 1909
• Cenchrometopa Hendel, 1909
• Chaetopsis Loew, 1868
• Euacaina Steyskal, 1963
• Eumecosomyia Hendel, 1909
• Eumetopiella Hendel, 1907
• Euphara Loew, 1868
• Euxesta Loew, 1868
• Heterodoxa J. R. Malloch, 1832
• Hypoecta Loew, 1868
• Lipsana Enderlein, 1938
• Neoeuxesta Malloch, 1930
• Notogramma Loew, 1868
• Paraphyola Hendel, 1909
• Pareuxesta Coquillett, 1901
• Perissoneura J. R. Malloch, 1832
• Polyteloptera Hendel, 1909
• Pseudeuxesta Hendel, 1910^[7]
• Siopa Hendel, 1909
• Steneretma
• Stenomyia Loew, 1868
• Texasa
• Ulivellia Speiser, 1929
• Vladolinia Kameneva, 2005
• Zacompsia Coquillett, 1901

Tribe Pterocallini Loew, 1868

• Aciuroides Hendel, 1914
• Apterocerina Hendel, 1909
• Chondrometopum Hendel, 1909
• Coscinum Hendel, 1909
• Cymatozus Enderlein, 1912
• Cyrtomostoma Hendel, 1909
• Dasymetopa Loew, 1868
• Elapata Hendel, 1909
• Goniaeola Hendel, 1909
• Lathrostigma Enderlein
• Megalaemyia Hendel, 1909
• Micropterocerus Hendel, 1914
• Neomyennis Hendel, 1914
• Ophthalmoptera Hendel, 1909
• Paragoniaeola?
• Paragorgopis Giglio-Tos, 1893
• Parophthalmoptera?
• Perissoza?
• Plagiocephalus Wiedemann, 1830
• Pterocalla Rondani, 1848
• Pterocerina Hendel, 1909
• Rhyparella Hendel, 1909
• Sympaectria Hendel, 1909
• Terpnomyennis Kameneva, 2004
• Terpnomyia Hendel, 1909
• Tetrapleura Schiner, 1868
• Xanthacrona Wulp, 1899

Tribe Seiopterini Kameneva and Korneyev, 1994

• Homalocephala Zetterstedt, 1838

Syn.: Psairoptera

• Pseudoseioptera Stackelberg 1955
• Seioptera Kirby, 1817

Syn.: Ortalis Fallén, 1810

Tribe Ulidiini Macquart, 1835

• Physiphora Fallén, 1810
• Timia Wiedemann, 1824
• Ulidia Meigen, 1826^[4]

Others

• Anacampta
• Blainvillia
• Califortalis
• Carlottaemyia
• Eupterocerina
• Euxestina
• Heramyia
• Hypochra
• Idanophana
• Macheirocera
• Meckelia
• Megaloprepemyia
• Metopocampta
• Myrmecomyia
• Ophryoterpnomyia
• Ortalis
• Paragorgopsis
• Phaeosoma
• Platyeuxesta
• Prionella
• Rhadinomyia
• Schnusimyia
• Systata
• Tephronota
• Terelliosoma
• Vespomima

References

1. Aldrich, J. M. (1932). "New Diptera, or two-winged flies, from America, Asia, and Java, with additional notes". Proceedings of the United States National Museum. 81 (9). The United States National Museum: 1–28. doi:10.5479/si.00963801.81-2932.1. Retrieved 26 June 2022.
2. Elena P. Kameneva & Valery A. Korneyev (2006). "Myennidini, a new tribe of the subfamily Otitinae (Diptera: Ulidiidae), with discussion of the suprageneric classification of the family" (PDF). Israel Journal of Entomology. 35–36: 497–586.
3. Hendel, F. (1914). "Die Bohrfliegen Sudamerikas". Abhandlungen und Berichte des Königl. Zoologischen und Anthropologisch-Etnographischen Museums zu Dresden. (1912)14: 1–84, 4 pls.
4. Meigen, J.W. (1826). Systematische Beschreibung der bekannten europäische n zweiflugeligen Insekten. Hamm: Vierter Theil. Schulz-Wundermann. pp. xii + 412 pp., pls. 42–54.
5. Robineau-Desvoidy, André Jean Baptiste (1830). "Essai sur les myodaires". Mémoires presentés à l'Institut des Sciences, Lettres et Arts, par divers savants et lus dans ses assemblées: Sciences, Mathématiques et Physique. 2 (2): 1–813. Retrieved 15 July 2018.
6. Steyskal, G.C. (1961). "The genera of Platystomatidae and Otitidae known to occur in America north of Mexico (Diptera, Acalyptratae)". Annals of the Entomological Society of America. 54 (401–410).
7. Hendel, Friedrich Georg (1910). "Über die Nomenklatur der Acalyptratengattungen nach Th. Beckers Katalog der paläarktischen Dipteren, Bd. 4". Wiener entomologische Zeitung. 29: 307–313. Retrieved 27 January 2021.
8. Hernández-Ortiz, V.; Kameneva, E.P.; Korneyev, V.A. (1998). "A new genus and species of the picture-winged flies (Diptera: Ulidiidae) from Mexico". Zhur. Ukrayn. Ent. Tovar. 4 (73–79). Retrieved 25 June 2022.
9. Rondani, Camillo (1868). "Diptera aliqua in America meridionali lecta a Prof. P. Strobel annis 1866-67 distincta et annotata, novis aliquibus descriptis". Annuario della Societàdi naturalisti di Modena. 3: 24–40. Retrieved 17 March 2021.
10. Loew, Hermann (1873). "Monographs of the Diptera of North America. Part III". Smithsonian Miscellaneous Collections. 11: vii + 351 +XIII pp., 4 pls. Retrieved 26 January 2021.
11. Kameneva, Elena P.; Korneyev, Valery A. (2006). "Myennidini, a new tribe of the subfamily Otitinae (Diptera: Ulidiidae), with discussion of the suprageneric classification of the family". Israel Journal of Entomology. 35–36: 497–586. Retrieved 26 June 2022.
12. Günther, Enderlein (1921). "Zur Kenntnis der Pterocallinen". Zoologischer Anzeiger. 52 (8/9): 211–219.
13. Barraclough, D.A. (2000). "A review of the southern African Ulidiidae (Diptera: Schizophora), with description of two new genera". Annals of the Natal Museum. 41: 77–101.
14. Hendel, F. (1907). "Nomina nova fur mehrere Gattungen der acalyptraten Musciden". Wiener Entomologische Zeitung. 26: 98. Retrieved 26 June 2022.
15. Loew, Hermann (1868). "Die amerikanischen Ulidina". Berliner entomologische Zeitschrift. (1867) 11: 283–326, pl. 2. Retrieved 26 June 2022.
16. Coquillett, Daniel William (1900). "Notes and descriptions of Ortalidae". Journal of the New York Entomological Society. 8: 21–25. Retrieved 26 January 2021.
17. Hernández-Ortiz, V. (1988). "Reconsideración taxonómica del género Dyscrasis Aldrich y la descripción de Pseudodyscrasis gen. nov. (Diptera: Otitidae)". Folia Entomológica Mexicana. 74: 181–188.
18. Malloch, J.R. (1931). "Flies of the genus Pseudotephritis Johnson (Diptera: Ortalidae)" (PDF). Proceedings of the United States National Museum. 79 ((34)[2900]): 1–6. Retrieved 26 June 2022.
19. Johnson, Charles Willison (1902). "Remarks on Tephronota ruficeps and description of a new species". Entomological News. 13: 143–144. Retrieved 26 June 2022.
20. Brèthes, J. (1926). "Description d'un nouveau genre et nouvelle espèce d'Ulididae du Chile". Revta Chil. Hist. Nat. 30: 187–188.
21. Bigot, J.M.F. (1885). "Les diagnoses de deux genres nouveaux de diptères appartenant a la famille des Ortalidae". Bulletin (Bimensuel) de la Société Entomologique de France. 1885 (18): clxv–clxvi.

External links

• Images of Ulidiidae from Diptera.info
• Family Ulidiidae at EOL
• Fauna Europaea
• Nomina Insecta Nearctica
• Delphinia picta University of Florida / Institute of Food and Agricultural Sciences
• Euxesta stigmatias, cornsilk fly University of Florida / Institute of Food and Agricultural Sciences