Tetraphalerus

Tetraphalerus is a genus of beetles in the family Ommatidae, It is currently known from two extant species native to South America and several fossil species from the Jurassic and Cretaceous of Asia.

Tetraphalerus Temporal range: Early Jurassic–Present PreꞒ Ꞓ O S D C P T J K Pg N
Tetraphalerus wagneri in side (top) and dorsal (bottom) view
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Family:	Ommatidae
Genus:	Tetraphalerus Waterhouse, 1901
Type species
Tetraphalerus wagneri Waterhouse, 1901

Systematics

According to Kirejtshuk, 2020.^[1]

Taxa labelled (?) are considered questionable by Kirejtshuk, 2020

• (?)†Tetraphalerus brevis Ponomarenko, 1964 – Karabastau Formation, Kazakhstan, Oxfordian
• Tetraphalerus bruchi Heller, 1913 – recent, Northern Argentina
• (?)†Tetraphalerus collaris Ponomarenko, 1997 – Dzun-Bain Formation, Mongolia, Aptian
• (?)†Tetraphalerus glabratus Ponomarenko, 1997 – Togo-Khuduk Formation, Mongolia, Bajocian/Bathonian
• (?)†Tetraphalerus mongolicus Ponomarenko, 1986 – Gurvan-Eren Formation, Mongolia, Aptian
• (?)†Tetraphalerus notatus Ponomarenko, 1997 – Dzun-Bain Formation, Mongolia, Aptian
• Tetraphalerus wagneri Waterhouse, 1901 – recent, South America

Other described fossil species:^[2]

• †Tetraphalerus largicoxa Lin, 1986 – Guanyintan Formation, China, Toarcian/Aalenian
• †Tetraphalerus penalveri Soriano & Delclòs, 2006 – La Pedrera de Rúbies Formation, Spain, Barremian
• †Tetraphalerus ponomarenkoi Soriano & Delclòs, 2006 – La Huérguina Formation, Spain, Barremian

Formerly assigned species

• †Tetraphalerus antiquus Ponomarenko, 1964 assigned to Allophalerus
• †Tetraphalerus bontsaganensis Ponomarenko, 1997 assigned to Allophalerus
• †Tetraphalerus brevicapitis Pononarenko & Martinez-Delclos, 2000 considered indeterminate by Kirejtshuk, 2020
• †Tetraphalerus fentaiensis (Ren, 1995) assigned to Monticupes
• †Tetraphalerus grandis Ponomarenko, 1964 assigned to Omma
• †Tetraphalerus incertus Ponomarenko, 1969 assigned to Allophalerus
• †Tetraphalerus laetus Lin, 1976 assigned to Notocupes
• †Tetraphalerus latus Tan et al. 2007 assigned to Allophalerus
• †Tetraphalerus lindae Jarzembowski et al. 2017 assigned to Bukhkalius
• †Tetraphalerus longicollis Ponomarenko, 1997 assigned to Omma
• †Tetraphalerus maximus Ponomarenko, 1968 assigned to Allophalerus
• †Tetraphalerus okhotensis Ponomarenko in Cromov et al., 1993 assigned to Allophalerus
• †Tetraphalerus oligocenicus (Crowson, 1962) assigned to Tetraphalerites
• †Tetraphalerus tenuipes Ponomarenko, 1964 assigned to Allophalerus
• †Tetraphalerus verrucosus Ponomarenko, 1966 assigned to Allophalerus
• †Tetraphalerus surrectus (Ren, 1995) assigned to Monticupes

References

1. Kirejtshuk, Alexander G. (2020-02-17). "Taxonomic Review of Fossil Coleopterous Families (Insecta, Coleoptera). Suborder Archostemata: Superfamilies Coleopseoidea and Cupedoidea". Geosciences. 10 (2): 73. Bibcode:2020Geosc..10...73K. doi:10.3390/geosciences10020073. ISSN 2076-3263.
2. Ommatidae Species List Archived 2012-10-23 at the Wayback Machine at Joel Hallan’s Biology Catalog. Texas A&M University. Retrieved on 11 May 2012.