Richard Swann Lull (November 6, 1867 April 22, 1957) was an American paleontologist and Sterling Professor at Yale University who is largely remembered for championing a non-Darwinian view of evolution, whereby mutation(s) could unlock presumed "genetic drives" that, over time, would lead populations to increasingly extreme phenotypes (and perhaps, ultimately, to extinction).

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Richard Swann Lull
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Born(1867-11-06)November 6, 1867
DiedApril 22, 1957(1957-04-22) (aged 89)
Alma materRutgers College
Columbia University
Scientific career
FieldsPaleontology
InstitutionsMassachusetts Agricultural College
Yale University
Doctoral advisorHenry Fairfield Osborn
Notable studentsGeorge Gaylord Simpson[1]
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American Museum party at Bone-Cabin Quarry, 1899. Seated, left to right, Walter Granger, Professor H.F. Osborn, Dr. W.D. Matthew; standing, F. Schneider, Professor R.S. Lull, Albert Thomson, Peter Kaison

Lull was born in Annapolis, Maryland, the son of naval officer Edward Phelps Lull and Elizabeth Burton, daughter of General Henry Burton. He married Clara Coles Boggs, and he has a daughter, Dorothy. He majored in zoology at Rutgers College, where he received both his undergraduate and master's degrees (M.S. 1896). He worked for the Division of Entomology of the United States Department of Agriculture but in 1894 became an assistant professor of zoology at the State Agricultural College in Amherst, Massachusetts (now the University of Massachusetts Amherst). Lull's interest in fossil footprints began at Amherst College, renowned for its collection of fossil footprints, and eventually led him to switch from entomology to paleontology.

In 1899, Lull worked as a member of the American Museum of Natural History's expedition to Bone Cabin Quarry, Wyoming, helping to collect that museum's brontosaur skeleton. In 1902 he again joined an American Museum team in Montana, then studied under Columbia University professor Henry Fairfield Osborn. In 1903 he received his Ph.D. from Columbia University, and in 1906, after a brief time at Amherst, he was named assistant professor of Vertebrate Paleontology in Yale College and Associate Curator of Vertebrate Paleontology at the Peabody Museum of Natural History. He stayed at Yale for the next 50 years. In 1933, Lull was awarded the Daniel Giraud Elliot Medal from the National Academy of Sciences.[2]

One famous example he used to support his non-Darwinian evolution theory concerned the enormous antlers of the Irish elk: he argued that these could not possibly be the result of natural selection, and instead reflected one of his "unlocked genetic drives" toward ever-increasing antler size. The poor elk, coping in each generation with ever-bigger antlers were eventually driven extinct.[3] His evolutionary theory was a form of orthogenesis.[4]

His book Organic Evolution (1917) received positive reviews and was described as an "excellent summary of the theories, facts, and factors of evolution."[5][6]

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Time cover, 1 Jun 1925

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