Orthalicidae (orthalicid land snails) are a family of tropical air-breathing land snails, terrestrial pulmonate gastropod mollusks are classified in the subfamily Orthalicoidea of the order Stylommatophora.[2]
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They are medium-sized to large snails, from about 3 cm (about 1.2 inches) to 9 cm (about 3.5 inches) in shell length
In former times, this family was often known as the Bulimulidae, but this term may also denote what today is the subfamily Bulimulinae. The subfamily Bulimulinae replaces the former family Bulimulidae Crosse & P. Fischer, 1873.
2005 taxonomy
The Orthalicidae belong to the Orthalicoidea, a superfamily in the order Sigmurethra. Like other stylommatophorans, the Sigmurethra belong to the suborder Helicina. Among the three subfamilies of Orthalicidae, the Bulimulinae are the most diverse, consisting of three tribes:[3]
- subfamily Amphibuliminae P. Fischer, 1873 - synonym Peltellinae Gray, 1855 (check decision of International Commission on Zoological Nomenclature for Peltellinae) - 17 species[4]
- subfamily Bulimulinae Tryon, 1867 - 1243 species[4]
- tribe Bulimulini Tryon, 1867 - synonyms: Bulimidae Guilding, 1828 (inv.); Berendtiinae P. Fischer & Crosse, 1872; Bothiembryontidae Iredale, 1937
- tribe Odontostomini Pilsbry & Vanatta, 1898 - 125 species[4]
- tribe Simpulopsini Schileyko, 1999 - synonym: Tomogeridae Jousseaume, 1877
- subfamily Orthalicinae Albers, 1850 - synonym: Liguidae Pilsbry, 1891 - 83 species[4]
2012 taxonomy
Breure & Romero (2012)[6] confirmed previous results from 2010, additionally they elevated Simpulopsini to Simpulopsidae.[6]
Genera within the family Orthalicidae include:
- Aposcutalus Dutra & Leme, 1985
- Clathrorthalicus Strebel, 1909
- Corona Albers, 1850[7][6][8]
- † Cortana Salvador & Simone, 2013
- Hemibulimus Martens, 1885[8]
- Kara Strebel, 1910[9]
- Liguus Montfort, 1810
- Orthalicus Beck, 1837 - type genus of the family Orthalicidae[3][6]
- Paeniscutalus Wurtz, 1947
- Porphyrobaphe Shuttleworth, 1856[6][8]
- Quechua Strebel, 1910
- Scholvienia Strebel, 1910
- Sultana Shuttleworth, 1856[8]
Albers J. C. 1860. (Manuscript edited by Eduard von Martens) Die Heliceeen nach natürlicher Vervandtschaft systematisch geordnet. Edition 2, Engelmann, Leipzig, xviii + 359 pp., Orthalicidae is at page 209. Breure A., Groenenberg D. & Schilthuizen M. (2010). "Gondwana revisited: New insights in the phylogenetic relationships within the Orthalicoidea (Mollusca, Stylommatophora)". poster. PDF Archived 2012-03-09 at the Wayback Machine. Breure A. S. H., Groenenberg D. S. J. & Schilthuizen M. (2010). "New insights in the phylogenetic relations within the Orthalicoidea (Gastropoda, Stylommatophora) based on 28S sequence data". Basteria 74(1-3): 25-31. Breure A. S. H. (2011) "Annotated type catalogue of the Orthalicoidea (Mollusca, Gastropoda) in the Royal Belgian Institute of Sciences, Brussels, with descriptions of two new species". ZooKeys 101: 1–50. doi:10.3897/zookeys.101.1133.
- (in German) Neubert E. & Janssen R. (2004). "Die Typen und Typoide des Natur-Museums Senckenberg, 84: Mollusca: Gastropoda: Pulmonata: Orthalicoidea: Bulimulidae (2), Orthalicidae, Placostylidae". Archiv für Molluskenkunde 133: 193–297. abstract.
- Ramirez J. & Ramírez R. (2010). "Análisis de la estructura secundaria del LSU rRNA mitocondrial de caracoles terrestres peruanos (Orthalicidae: Gastropoda) / Analysis of the secondary structure of mitochondrial LSU rRNA of Peruvian land snails (Orthalicidae: Gastropoda)." Revista Peruana de Biología 17(1): 53–57. PDF.
- Ramirez J., Ramírez R., Romero P., Chumbe A. & Ramírez P. (2009). "Posición evolutiva de caracoles terrestres peruanos (Orthalicidae) entre los Stylommatophora (Mollusca: Gastropoda) / Evolutionary position of Peruvian land snails (Orthalicidae) among Stylommatophora (Mollusca: Gastropoda)". Revista Peruana de Biología 16(1): 51–56. PDF.
