Hydrangeaceae

Hydrangeaceae is a family of flowering plants in the order Cornales, with a wide distribution in Asia and North America, and locally in southeastern Europe.^[3]

Hydrangeaceae
Hydrangea macrophylla, a member of the subfamily Hydrangeoideae
Fendlera rupicola, a member of the subfamily Jamesioideae
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Eudicots
Clade:	Asterids
Order:	Cornales
Family:	Hydrangeaceae Dumort.[1]
Genera
See text
Synonyms[2]
Kirengeshomaceae Nakai Philadelphaceae Martinov

Hydrangea hydrangeoides, a member of the now-synonymised genus Schizophragma

Description

The genera are characterised by leaves in opposite pairs (rarely whorled or alternate), and regular, bisexual flowers with four (rarely 5–12) petals. The fruit is a capsule or berry containing several seeds, the seeds with a fleshy endosperm.^[4]

Genera

The following genera are accepted:^[5]

• Carpenteria Torr.
• Deutzia Thunb.
• Fendlera Engelm. & A.Gray
• Fendlerella A.Heller
• Hydrangea Gronov. ex L.
• Jamesia Torr. & A.Gray
• Kirengeshoma Yatabe
• Philadelphus L.
• Whipplea Torr.

Phylogeny

The family Hydrangeaceae has two subfamilies, namely Jamesioideae and Hydrangeoideae. The subfamily Jamesioideae comprises the genera Jamesia and Fendlera.^[6][7] They are the sister group to the remaining Hydrangeaceae.^[7][8] The subfamily Hydrangeoideae has two tribes:^[9][7][8] Hydrangeae consists of Hydrangea s.l.,^[9] and Philadelpheae consists of Philadelphus, Carpenteria, Deutzia, Kirengeshoma, Whipplea, and Fendlerella.^[7][8] Carpenteria is the sister group to Philadelphus. Deutzia is the sister group to Kirengeshoma, and Fendlerella is the sister group to Whipplea.^[8][6][7] However the relationships among those three clades within the tribe Philadelphae are a bit unclear.^[8] The following cladogram summarizes results from different studies, and for each node it is noted which studies support the sister group positions of the following branches:

Hydrangeaceae[7][8]	Hydrangeoideae[7][8] Hydrangeae[7] Hydrangea s.l. (including Broussaisia, Cardiandra, Decumaria, Deinanthe, Dichroa, Pileostegia, Platycrater, and Schizophragma)[9] Philadelpheae[7] [7][6][9][8] Philadelphus Carpenteria [7] [6][7][8] Deutzia Kirengeshoma [6][7][8] Whipplea Fendlerella Jamesioideae[7][6][8] Jamesia Fendlera