Gnatcatcher

The gnatcatchers are a family of small passerine birds in the family Polioptilidae. The 22 species occur in North and South America (except for the far south and the high Andean regions). Most species of this mainly tropical and subtropical group are resident, but the blue-grey gnatcatcher of the United States and southern Canada migrates south in winter. They are close relatives of the wrens.^[1]

Gnatcatchers
California gnatcatcher (Polioptila californica)
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Aves
Order:	Passeriformes
Superfamily:	Certhioidea
Family:	Polioptilidae Baird, 1858
Genera
Ramphocaenus Microbates Polioptila

Description

These dainty birds are intermediate between Old World warblers and wrens in their structure and habits, moving restlessly through foliage seeking insects. The gnatcatchers are mainly soft bluish grey in color, and have the typical insectivore's long sharp bill. Many species have distinctive black head patterns (especially males) and long, regularly cocked, black-and-white tails. The skulking gnatwrens are browner, more thickset, and with proportionally shorter tails and longer bills.^[1]

Distribution and habitat

They are distributed from North to South America, with the exception of the far south and high Andean regions. Gnatwrens typically occur in the undergrowth of dense, often humid, forest, while gnatcatchers, depending on the species involved, occur in anything from dry scrubby habitats (e.g. the California gnatcatcher) to the canopy of humid Amazonian forest (e.g. the Guianan gnatcatcher). The North American species nest in bushes or trees, but the breeding behavior of several of the Neotropical species is essentially unknown.^[1]

Taxonomy and systematics

Certhioidea Tichodromidae: wallcreeper – 1 species Sittidae: nuthatches – 29 species Salpornithidae: spotted creepers – 2 species Certhiidae: treecreepers – 9 species Polioptilidae: gnatcatchers – 22 species Troglodytidae: wrens – 96 species

Relationships among families in the superfamily Certhioidea.[2][3]

A species new to science, the critically endangered Iquitos gnatcatcher Polioptila clementsi, was first described in 2005.^[4] This species is a member of the Guianan gnatcatcher Polioptila guianensis complex, which recently has been proposed split into three species (four with the Iquitos gnatcatcher), but not all authorities have accepted this (e.g. SACC). Furthermore, other groups should possibly be split, notably the tropical gnatcatcher Polioptila plumbea and masked gnatcatcher Polioptila dumicola complexes, but at present scientific papers on these matters are lacking.

The family contains 22 species divided into 3 genera:^[3]

• Ramphocaenus (2 species) – gnatwrens
• Microbates (2 species) – gnatwrens
• Polioptila (18 species) – gnatcatchers