Dryosauridae

Not to be confused with Dyrosauridae.

Dryosauridae was a family of primitive iguanodonts, first proposed by Milner & Norman in 1984. They are known from Middle Jurassic to Early Cretaceous rocks of Africa, Europe, and North America.^[1][2][3][4]

Dryosauridae Temporal range: Middle Jurassic – Early Cretaceous, 164–115 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Dysalotosaurus skeletal mount in Berlin
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Dinosauria
Clade:	†Ornithischia
Clade:	†Neornithischia
Clade:	†Ornithopoda
Clade:	†Iguanodontia
Clade:	†Dryomorpha
Superfamily:	†Dryosauroidea Milner & Norman, 1984
Family:	†Dryosauridae Milner & Norman, 1984
Subgroups
†Callovosaurus? †Dryosaurus †Dysalotosaurus †Elrhazosaurus †Eousdryosaurus? †Iyuku? †Kangnasaurus? †Valdosaurus †Vectidromeus?

Classification

Until recently, many dryosaurids have been regarded as dubious (Callovosaurus and Kangnasaurus) or as species of the type member, Dryosaurus (Dysalotosaurus, Elrhazosaurus and Valdosaurus). However, more recent studies redescribe these genera as valid.^{[3][4][5][6][7]} Under the Phylocode, Madzia et al. (2021) formally defined Dryosauridae as "the largest clade containing Dryosaurus altus but not Iguanodon bernissartensis."^[8] The cladogram below follows their chosen reference phylogeny, taken from the description of Orthomerus dolloi.^[9]

Iguanodontia	Rhabdodontidae Rhabdodon spp. Zalmoxes robustus Mochlodon vorosi Zalmoxes shqiperorum Tenontosaurus tilletti Tenontosaurus dossi Dryomorpha Ankylopollexia Dryosauridae Eousdryosaurus nanohallucis "Camptosaurus" valdensis Callovosaurus leedsi Dysalotosaurus lettowvorbecki Dryosaurus altus Valdosaurus canaliculatus Elrhazosaurus nigeriensis

References

1. Norman, David B.; Weishampel, David B. (1990). "Iguanodontidae and related ornithopods". In Weishampel, David B.; Dodson, Peter; Osmólska Halszka (eds.). The Dinosauria. Berkeley: University of California Press. pp. 510–533. ISBN 0-520-06727-4.
2. Norman, David B. (2004). "Basal Iguanodontia". In Weishampel, D.B.; Dodson, P.; Osmólska H. (eds.). The Dinosauria (2nd ed.). Berkeley: University of California Press. pp. 413–437. ISBN 0-520-24209-2.
3. Ruiz-Omeñaca, José Ignacio; Pereda Suberbiola Xabier; Galton, Peter M. (2007). "Callovosaurus leedsi, the earliest dryosaurid dinosaur (Ornithischia: Euornithopoda) from the Middle Jurassic of England". In Carpenter Kenneth (ed.). Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs. Bloomington and Indianapolis: Indiana University Press. pp. 3–16. ISBN 978-0-253-34817-3.
4. Galton, Peter M. (2009). "Notes on Neocomian (Lower Cretaceous) ornithopod dinosaurs from England – Hypsilophodon, Valdosaurus, "Camptosaurus", "Iguanodon" – and referred specimens from Romania and elsewhere" (PDF). Revue de Paléobiologie. 28 (1): 211–273.
5. McDonald, A.T.; Kirkland, J.I.; DeBlieux, D.D.; Madsen, S.K.; Cavin, J.; Milner, A.R.C.; Panzarin, L. (2010). "New Basal Iguanodonts from the Cedar Mountain Formation of Utah and the Evolution of Thumb-Spiked Dinosaurs". PLOS ONE. 5 (11): e14075. Bibcode:2010PLoSO...514075M. doi:10.1371/journal.pone.0014075. PMC 2989904. PMID 21124919.
6. Andrew T. McDonald (2011). "The taxonomy of species assigned to Camptosaurus (Dinosauria: Ornithopoda)" (PDF). Zootaxa. 2783: 52–68. doi:10.11646/zootaxa.2783.1.4.
7. Paul M. Barrett; Richard J. Butler; Richard J. Twitchett; Stephen Hutt (2011). "New material of Valdosaurus canaliculatus (Ornithischia: Ornithopoda) from the Lower Cretaceous of southern England". Special Papers in Palaeontology. 86: 131–163.
8. Madzia, Daniel; Arbour, Victoria M.; Boyd, Clint A.; Farke, Andrew A.; Cruzado-Caballero, Penélope; Evans, David C. (2021-12-09). "The phylogenetic nomenclature of ornithischian dinosaurs". PeerJ. 9: e12362. doi:10.7717/peerj.12362. ISSN 2167-8359. PMC 8667728. PMID 34966571.
9. Madzia, Daniel; Jagt, John W. M.; Mulder, Eric W. A. (2020-04-01). "Osteology, phylogenetic affinities and taxonomic status of the enigmatic late Maastrichtian ornithopod taxon Orthomerus dolloi (Dinosauria, Ornithischia)". Cretaceous Research. 108: 104334. Bibcode:2020CrRes.10804334M. doi:10.1016/j.cretres.2019.104334. ISSN 0195-6671.

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