Coccinelloidea

Coccinelloidea is a superfamily of beetles in the order Coleoptera, formerly included in the superfamily Cucujoidea. There are more than 10,000 species in Coccinelloidea, including more than 6000 in the lady beetle family Coccinellidae.^[1][2][3]

Coccinelloidea Temporal range: Early Cretaceous–Recent PreꞒ Ꞓ O S D C P T J K Pg N
Seven-Spotted Ladybug (Coccinella septempunctata)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Suborder:	Polyphaga
Infraorder:	Cucujiformia
Superfamily:	Coccinelloidea Latreille, 1807
Families
see text

Morphology

Adult Coccinelloidea have a reduced tarsal formula (each tarsus with 4 or 3 segments), hind coxae separated by more than 1/3 their width, the intercoxal process of abdominal ventrite 1 usually broadly rounded or truncate, hindwings with reduced anal veins and lacking a closed radial cell, the adeagus resting on its side when retracted and the phallobase usually reduced.^[1]

Larval Coccinelloidea have a unisetose pretarsal claw, spiracles that are usually annular, and the sensory appendage of the second antennal segment usually as long as the third segment.^[1]

Families

The family constituency of this lineage has changed considerably over time, from as few as four or five recognized families in 1970^[4] to 15 in 2015.^[5]

Cucujiformia	Lymexyloidea Tenebrionoidea Cleroidea Chrysomeloidea Curculionoidea Cucujoidea Coccinelloidea Bothrideridae and allies Latridiidae Akalyptoischiidae Alexiidae Corylophidae and allies Endomychidae Coccinellidae

• Akalyptoischiidae Lord, Hartley, Lawrence, McHugh, Whiting & Miller, 2010
• Alexiidae Imhoff, 1856
• Anamorphidae Strohecker, 1953
• Bothrideridae Erichson, 1845 (dry bark beetles)
• Cerylonidae Billberg, 1820 (minute bark beetles)
• Coccinellidae Latreille, 1807 (ladybird beetles)
• Corylophidae LeConte, 1852 (minute hooded beetles)
• Discolomatidae Horn, 1878
• Endomychidae Leach, 1815 (handsome fungus beetles)
• Eupsilobiidae Casey, 1895
• Euxestidae Grouvelle, 1908 (well polished beetles)
• Latridiidae Erichson, 1842 (minute brown scavenger beetles)
• Murmidiidae Jacquelin du Val, 1858
• Mycetaeidae Jacquelin du Val, 1857
• Teredidae Seidlitz, 1888

References

1. Robertson, James; Slipinski, Adam; Moulton, Matthew; Shockley, Floyd; et al. (2015). "Phylogeny and classification of Cucujoidea and the recognition of a new superfamily Coccinelloidea (Coleoptera: Cucujiformia)". Systematic Entomology. 40 (4): 745–778. doi:10.1111/syen.12138. ISSN 0307-6970. S2CID 55206626.
2. "Coccinelloidea Superfamily Information". BugGuide.net. Retrieved 13 April 2019.
3. "Coccinelloidea Report". Integrated Taxonomic Information System. Retrieved 13 April 2019.
4. Donald Borror; Richard White (1970). A field guide to the insects of America north of Mexico. Houghton Mifflin. ISBN 978-0-395-07436-7.
5. Robertson, James A.; Ślipiński, Adam; Moulton, Matthew; et al. (2015). "Phylogeny and classification of Cucujoidea and the recognition of a new superfamily Coccinelloidea (Coleoptera: Cucujiformia)". Systematic Entomology. 40 (4): 745–778. doi:10.1111/syen.12138. S2CID 55206626.

Further reading

• Vandenberg, Natalia J. (2002). Arnett, Ross H. Jr.; Thomas, M.C.; Skelley, P.E.; Frank, J.H. (eds.). Family 93; Coccinellidae Latreille 1807. American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press. pp. 371–389. ISBN 0-8493-0954-9.
• Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; et al. (2011). "Family-group names in Coleoptera (Insecta)". ZooKeys (88): 1–972. doi:10.3897/zookeys.88.807. ISSN 1313-2989. PMC 3088472. PMID 21594053.