The Phaneropterinae , the sickle-bearing bush crickets or leaf katydids , are a subfamily of insects within the family Tettigoniidae .[2] [3] false katydids [4] round-headed katydids .[2]
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The name Phaneropterinae is based upon the Old World genus Phaneroptera P. falcata [5] Barbitistini and Odonturini have become brachypterous .
Adult Phaneropterinae species with eggs
The legs of individuals in this subfamily vary from genus to genus, but, as in nearly all Orthoptera , the posterior (rear) legs are adapted to leaping, and as such are always much longer than other legs. Phaneropterinae are generally well-camouflaged with green and brown colors being most prevalent, but there are exceptions including certain Aganacris Scaphura Batesian mimics of wasps.[6]
The Phaneropterinae are largely arboreal in habitat.[2] [2] meadow grasshoppers , so much so that they seldom come to human notice.[2]
The ovipositor and male genitalia vary according to the genus. The Phaneropterinae differ from other subfamilies of Tettigoniidae (and other Orthoptera) in their oviposition; their eggs are rarely deposited in the earth, but are either glued in double rows to twigs, or are inserted in the edges of leaves.[2]
The subfamily Phaneropterinae was first erected in 1838 by German zoologist Hermann Burmeister .[3] As of January 2024 [ update] [3]
Acrometopini
Auth.: Brunner von Wattenwyl, 1878 – Africa, Europe, the Middle East
Acrometopa Fieber, 1853 Altihoratosphaga Hemp, Voje, Heller, Warchalowska-Sliwa & Hemp, 2010 Conchotopoda Karsch, 1887 Horatosphaga Schaum, 1853 Lamecosoma Ragge, 1960 Peronura Karsch, 1889 Peronurella Hemp, 2018 Prosphaga Ragge, 1960 Tenerasphaga Hemp, 2018 Amblycoryphini
Auth.: Brunner von Wattenwyl, 1878 – Americas, Africa
Amblycorypha insolita Orophus tessellatus
Subtribe Plangiina Cadena-Castañeda, 2015 Madagascarantia Massa, 2017 Monteiroa Karsch, 1889 Paraplangia Heller, 2018 Plangia Stål, 1873 Pseudoplangia Massa, 2014 Agaurella Uvarov, 1939 Amblycorypha Stål, 1873 Eurycoplangiodes Massa, 2020 Eurycorypha Stål, 1873 Orophus Saussure, 1859 Oxygonatium Ragge, 1980 Paraeurycorypha Massa, 2020 Plangiodes Chopard, 1954
Barbitistini
Auth.: Jacobson, 1905 – Europe to central Asia
Male Barbitistes obtusus Isophya
Ancistrura Uvarov, 1921 - monotypic Ancistrura nigrovittata Andreiniimon Capra, 1937 - monotypic Andreiniimon nuptialis Barbitistes Charpentier, 1825 Dasycercodes Bei-Bienko, 1951 - monotypic Dasycercodes iranicus Euconocercus Bei-Bienko, 1950 Isoimon Bei-Bienko, 1954 - monotypic Isoimon riabovi Isophya Brunner von Wattenwyl, 1878 Kurdia Uvarov, 1916 Leptophyes Fieber, 1853 Metaplastes Ramme, 1939 Orthocercodes Bei-Bienko, 1951 - monotypic Orthocercodes zarudnyi Phonochorion Uvarov, 1916 Poecilimon Fischer, 1853 Polysarcus Fieber, 1853
Catoptropterigini
Auth.: Massa, 2016 – Africa
Catoptropteryx Karsch, 1890 Griffinipteryx Massa, 2016 Ducetiini
Auth.: Brunner von Wattenwyl, 1878 – Africa, India, China, Indochina to Australia
Ducetia Shirakisotima japonica Abaxisotima Gorochov, 2005 Agnapha Brunner von Wattenwyl, 1891 Bulbistridulous Xia & Liu, 1991 Ducetia Stål, 1874 Kuwayamaea Matsumura & Shiraki, 1908 Noia Walker, 1870 Paraducetia Gorochov & Kang, 2002 Paragnapha Willemse, 1923 Prohimerta Hebard, 1922 Shirakisotima Furukawa, 1963 Subibulbistridulous Shi, 2002 Dysoniini
Dysonia sp.Markia arizae Auth.: Rehn, 1950 – tropical Americas
Genus group Dysonia (temporary)
Dysonia White, 1862 Lichenodentix Cadena-Castañeda, 2011 Lichenomorphus Cadena-Castañeda, 2011
Genus group Markia Cadena-Castañeda, 2013
Apolinaria Rehn, 1950 Lichenodraculus Braun, 2011 Machima Brunner von Wattenwyl, 1878 Machimoides Rehn, 1950 Markia White, 1862
Genus group Paraphidniae Cadena-Castañeda, 2016
Anaphidna Gorochov & Cadena-Castañeda, 2012 Paraphidnia Giglio-Tos, 1898
Genus group not assigned
Hammatofera Brunner von Wattenwyl, 1878 Quiva Hebard, 1927 Yungasacris Rehn, 1950
Ectemnini
Auth.: Cadena-Castañeda, 2015 – tropical Americas
Ectemna Brunner von Wattenwyl, 1878 Euthyrrhachis Brunner von Wattenwyl, 1878 Elimaeini
Auth.: Brunner von Wattenwyl, 1891 – Asia
Elimaea punctifera Ectadia Brunner von Wattenwyl, 1878 Elimaea Stål, 1874 Hemielimaea Brunner von Wattenwyl, 1878 Orthelimaea Karny, 1926 Holochlorini
Auth.: Brunner von Wattenwyl, 1878 – Africa, Asia-Pacific
Holochlora japonica Poecilopsyra octoseriata
Ancylecha Serville, 1838 Arantia Stål, 1874 Arnobia Stål, 1876 Calopsyra Brunner von Wattenwyl, 1891 Casigneta Brunner von Wattenwyl, 1878 Cesasundana Koçak & Kemal, 2009 Dapanera Karsch, 1889 Elbenia Stål, 1876 Furnia Stål, 1876 Gonatoxia Karsch, 1889 Holochlora Stål, 1873 Leucopodoptera Rentz & Webber, 2003 Liotrachela Brunner von Wattenwyl, 1878 Molpa Walker, 1870 Parapsyra Carl, 1914 Phaulula Bolívar, 1906 Phygela Stål, 1876 Poecilopsyra Dohrn, 1892 Pseudopsyra Hebard, 1922 Psyrana Uvarov, 1940 Rectimarginalis Liu & Kang, 2007 Ruidocollaris Liu, 1993 Sinochlora Tinkham, 1945 Stibaroptera Bolívar, 1906 Stictophaula Hebard, 1922 Sympaestria Brunner von Wattenwyl, 1878 Tapiena Bolívar, 1906
Insarini
Auth.: Rehn, & Hebard, 1914 – tropical & subtropical Americas
Arethaea Arethaea Stål, 1876 Brachyinsara Rehn & Hebard, 1914 Insara Walker, 1869 Psilinsara Hebard, 1932 Kevaniellini
Auth.: Massa, 2017 – eastern Africa
Kevaniella Chopard, 1954 Microcentrini
Auth.: Brunner von Wattenwyl, 1878 – Americas
Eggs of Microcentrum
Acropsis Uvarov, 1939 Anapolisia Piza, 1980 Apoballa Brunner von Wattenwyl, 1878 Boroseiyla Mendes, Chamorro-Rengifo & Rafael, 2020 Caauara Mendes, Chamorro-Rengifo & Rafael, 2020 Capanema Mendes & Rafael, 2021 Capiguara Mendes, Chamorro-Rengifo & Rafael, 2020 Ctenophlebia Stål, 1873 Hyalipenna Mendes, Chamorro-Rengifo & Rafael, 2020 Ischyra Brunner von Wattenwyl, 1878 Lamprophyllum Hebard, 1924 Lobophyllus Saussure, 1859 Microcentrum Scudder, 1862 Petaloptera Saussure, 1859 Philophyllia Stål, 1873 Phoebolampta Brunner von Wattenwyl, 1878 Raggophyllum Nickle, 1967 Syntechna Brunner von Wattenwyl, 1878 Tuaia Mendes, Chamorro-Rengifo & Rafael, 2020 Tukunha Mendes & Rafael, 2021
Mirolliini
Auth.: Brunner von Wattenwyl, 1878 – Asia
Amirollia Ingrisch, 2011 Deflorita Bolívar, 1906 Hemimirollia Ingrisch, 2011 Hueikaeana Ingrisch, 1998 Mirollia Stål, 1873 Morgeniini
Auth.: Karsch, 1890 - central and western Africa
Mangomaloba Sjöstedt, 1902 Morgenia Karsch, 1890 Odonturini
Auth.: Brunner von Wattenwyl, 1878 – Americas, Iberian peninsula, Africa, Pacific islands
Obolopteryx castanea
Acanthorintes Cohn, Swanson & Fontana, 2014 Dichopetala Brunner von Wattenwyl, 1878 Gymnocerca Cohn, Swanson & Fontana, 2014 Mactruchus Cohn, Swanson & Fontana, 2014 Obolopteryx Cohn, Swanson & Fontana, 2014 Planipollex Cohn, Swanson & Fontana, 2014 Pterodichopetala Buzzetti, Barrientos-Lozano & Rocha-Sánchez, 2010 Rhabdocerca Cohn, Swanson & Fontana, 2014 Angara Brunner von Wattenwyl, 1891 Anisophya Karabag, 1960 Arachnitus Hebard, 1932 Arostratum Massa, 2015 Atlasacris Rehn, 1914 Cohnia Buzzetti, Fontana & Carotti, 2010 Monticolaria Sjöstedt, 1910 Nanoleptopoda Braun, 2011 Odontura Rambur, 1838 Odonturoides Ragge, 1980 Parangara Rehn, 1945 Paraperopyrrhicia Ebner, 1915 Peropyrrhicia Brunner von Wattenwyl, 1891 Xenicola Uvarov, 1940
Otiaphysini
Auth.: Karsch, 1889 - Africa
Debrona Walker, 1870 Drepanophyllum Karsch, 1890 Stenamblyphyllum Karsch, 1896 Tetraconcha Karsch, 1890 Pardalotini
Auth.: Brunner von Wattenwyl, 1878 - Africa
Percynini
Auth.: Cadena-Castañeda, 2015 – South America
Percyna Grant, 1964 Sictuna Walker, 1869 Phaneropterini
Auth.: Burmeister, 1838 – distributed worldwide
Phaneroptera nana nymph
Anaulacomerina Brunner von Wattenwyl, 1878 – Central and South America
Abrodiaeta Brunner von Wattenwyl, 1891 Anaulacomera Stål, 1873 Grammadera Brunner von Wattenwyl, 1878 Mendesius Piza, 1960 Montealegrezia Cadena-Castañeda, 2012 Phaneropteroides Piza, 1971 Rostellula Gorochov, 2018 Separatula Gorochov, 2018 Tenellulus Cadena-Castañeda, 2012
Pelecynotina Cadena-Castañeda, 2015 – South America
Viadanina Cadena-Castañeda, 2012 – Central and South America
Tomeophera Brunner von Wattenwyl, 1878 Viadana Walker, 1869
Subtribe not assigned
Agennis Brunner von Wattenwyl, 1891 Chloroscirtus Saussure & Pictet, 1897 Dioncomena Brunner von Wattenwyl, 1878 Dithela Karsch, 1890 Eucatopta Karsch, 1889 Melidia Stål, 1876 Nephoptera Uvarov, 1929 Parapyrrhicia Brunner von Wattenwyl, 1891 Phaneroptera Serville, 1831 Pigalua Massa, 2017 Pleothrix Ragge, 1980 Pseudophaneroptera Brunner von Wattenwyl, 1878 Symmetrokarschia Massa, 2015 Symmetroraggea Massa, 2015 Xenodoxus Carl, 1914
Phlaurocentrini
Auth.: Karsch, 1889 – Africa
Buettneria Karsch, 1889 Leiodontocercus Chopard, 1954 Phlaurocentrum Karsch, 1889 Phyllopterini
Auth.: Brunner von Wattenwyl, 1878
Phyllopterina Brunner von Wattenwyl, 1878 – Central and South America
Apocerycta Brunner von Wattenwyl, 1878 Arota Brunner von Wattenwyl, 1891 Cephalophylloptera Cadena-Castañeda, 2015 Hyperphrona Brunner von Wattenwyl, 1878 Hyperphorina Cadena-Castañeda, 2015 Itarissa Walker, 1869 Julchiella Cadena-Castañeda, 2015 Meneghelia Piza, 1980 Metaprosagoga Vignon, 1930 Phrixa Stål, 1874 Phylloptera Serville, 1831 Resecabimus Cadena-Castañeda, 2015
Uberabina Cadena-Castañeda, 2015 – South America
Polichnodes Giglio-Tos, 1898 Uberaba Bruner, 1915
Plagiopleurini
Auth.: Brunner von Wattenwyl, 1878 – South America
Diplophyllus Saussure, 1859 Parableta Brunner von Wattenwyl, 1878 Plagiopleura Stål, 1873 Plangiopsidini
Auth.: Cadena-Castañeda, 2015 – Africa
Plangiola Bolívar, 1906 Plangiopsis Karsch, 1889 Poreuomenini
Auth.: Brunner von Wattenwyl, 1878 – Africa
Cestromoecha Karsch, 1893 Paraporeuomena Massa, 2018 Poreuomena Brunner von Wattenwyl, 1878 Preussiini
Auth.: Karsch, 1890 – Africa
Enochletica Karsch, 1896 Preussia Karsch, 1890 Weissenbornia Karsch, 1888 Pycnopalpini
Auth.: Cadena-Castañeda, 2014 – Central and South America
Pycnopalpina Cadena-Castañeda, 2014
Hetaira Brunner von Wattenwyl, 1891 Montezumina Hebard, 1925 Pycnopalpa Serville, 1838 Topanga Walker, 1869
Theiina Cadena-Castañeda, 2014
Dolichocercus Rehn & Hebard, 1914 Oxyprorella Giglio-Tos, 1898 Theia Brunner von Wattenwyl, 1891 Theiella Cadena-Castañeda, 2014
Scudderiini
Auth.: Brunner von Wattenwyl, 1878 – Americas
Scudderia
Aganacris Walker, 1871 Caroliniella Cadena-Castañeda, 2015 (note : name conflict with Coleopteran)Ceraia Brunner von Wattenwyl, 1891 Ceraiaella Hebard, 1933 Ctenophorema Piza, 1967 Euceraia Hebard, 1927 Harroweria Hebard, 1927 Homotoicha Brunner von Wattenwyl, 1891 Inscudderia Caudell, 1921 Ligocatinus Rehn, 1901 Parascudderia Brunner von Wattenwyl, 1891 Phanerocercus Piza, 1980 Scudderia Stål, 1873 Theudoria Stål, 1874 Vellea Walker, 1869 Zenirella Piza, 1973
Steirodontini
Auth.: Brunner von Wattenwyl, 1878 – subtropical and tropical parts of the Americas
Stilpnochlora couloniana Cnemidophyllum Rehn, 1917 Emsleyfolium Cadena-Castañeda, Mendes & Alves-Oliveira, 2016 Nicklephyllum Cadena-Castañeda, 2016 Steirodon Serville, 1831 Stilpnochlora Stål, 1873 Terpnistrini
Auth.: Brunner von Wattenwyl, 1878 – Africa, Sri Lanka
Diogena Brunner von Wattenwyl, 1878 Gelotopoia Brunner von Wattenwyl, 1891 Terpnistria Stål, 1873 Terpnistrioides Ragge, 1980 Tropidophrys Karsch, 1896 Trachyzulphini
Auth.: Gorochov, 2014 – China, Indochina, Malesia
Trachyzulpha Dohrn, 1892 Trigonocoryphini
Auth.: Bei-Bienko, 1954 – Middle East, India, Malaysia
Cosmozoma Karsch, 1889 Megotoessa Karsch, 1889 Trigonocorypha Stål, 1873 Tylopsidini
Auth.: Brunner von Wattenwyl, 1878 – Africa, Europe, Middle East, western Asia
Tylopsis lilifolia Vossiini
Auth.: Cadena-Castañeda, 2015 – Africa, Asia
Vossiina Cadena-Castañeda, 2015 – Africa
Azamia Bolívar, 1906 Vossia Brunner von Wattenwyl, 1891
Xantiina Cadena-Castañeda, 2015 – Asia
Paraxantia Liu & Kang, 2009 – ChinaXantia Brunner von Wattenwyl, 1878 – Borneo
Zeuneriini
Zeuneria melanopeza Auth.: Karsh, 1890 - tropical Africa
Gravenreuthia Karsch, 1892 Zeuneria Karsch, 1889 Genus groups
Genus groups include:
Male Acripeza reticulata Caedicia simplex
Acridopezae Brunner von Wattenwyl, 1878 – Australia
Aegimiae Brunner von Wattenwyl, 1878 – Central and South America
Aniarae Brunner von Wattenwyl, 1878 – South America, E. Africa
Aniarella Bolívar, 1906 Burgilis Stål, 1873 Corymeta Brunner von Wattenwyl, 1878 Coryphoda Brunner von Wattenwyl, 1878 Hyperophora Brunner von Wattenwyl, 1878 Pseudoburgilis Brunner von Wattenwyl, 1878 Tetana Brunner von Wattenwyl, 1878
Centroferae Brunner von Wattenwyl, 1878 – South America
Centrofera Brunner von Wattenwyl, 1878
Cosmophylla Brunner von Wattenwyl, 1878 – South America, Madagascar, Australia
Cosmophyllum Blanchard, 1851 Engonia Brunner von Wattenwyl, 1878 Marenestha Brunner von Wattenwyl, 1878 Ozphyllum Rentz, Su & Ueshima, 2007 Paracosmophyllum Brunner von Wattenwyl, 1891 Stenophyllia Brunner von Wattenwyl, 1878
Dysmorphae Brunner von Wattenwyl, 1878 – Malesia
Dysmorpha Brunner von Wattenwyl, 1878
Ephippithytae Brunner von Wattenwyl, 1878 – Vietnam, Papua New Guinea, Australia
Alectoria superba Alectoria Brunner von Wattenwyl, 1879 Caedicia Stål, 1874 Currimundria Rentz, Su & Ueshima, 2008 Diastella Brunner von Wattenwyl, 1878 Dicorypha Krauss, 1902 Ephippitytha Serville, 1838 Kurandoptera Rentz, Su & Ueshima, 2008 Polichne Stål, 1874 Protina Brunner von Wattenwyl, 1879 Symmachis Brunner von Wattenwyl, 1878 Torbia Walker, 1869
Eurypalpae Brunner von Wattenwyl, 1878 – Eastern China, Vietnam, Sumatra, Australia
Ceratopompa Karsch, 1890 Zulpha Walker, 1870
Isopserae Brunner von Wattenwyl, 1878 – Eastern Africa, India, Indochina, Malesia
Isopsera Brunner von Wattenwyl, 1878 Pelerinus Bolívar, 1906 Pseudopyrrhizia Brunner von Wattenwyl, 1891
Karschiae Brunner von Wattenwyl, 1891
Karschia Brunner von Wattenwyl, 1891
Leptoderae Brunner von Wattenwyl, 1878 – Sri Lanka, Malesia
Leptoderes Serville, 1838
Pseudophaneropterae Brunner von Wattenwyl, 1878 – Sri Lanka, Sumatra
Pseudophaneroptera Brunner von Wattenwyl, 1878
ScambophyllaBrunner von Wattenwyl, 1878 – Indochina, Malesia
Scaphurae Westwood, 1838 – South America
Taeniomenae Brunner von Wattenwyl, 1878 – Australia
Elephantodeta Brunner von Wattenwyl, 1878 Tinzeda Walker, 1869
Turpiliae Brunner von Wattenwyl, 1878 – Central America, Caribbean
Turpilia Stål, 1874 Turpiliodes Hebard, 1932
These genera have not been placed in a tribe:
Alloducetia Xia & Liu, 1992 Anchispora Brunner von Wattenwyl, 1891 Angustithorax Massa, 2015 Anisotochra Karsch, 1889 Anormalous Liu, 2011 Aphroptera Bolívar, 1902 Astathomima Karny, 1929 Austrodontura Fontana & Buzzetti, 2004 Balneum Piza, 1967 Baryprostha Karsch, 1891 Bertius Piza, 1974 Bongeia Sjöstedt, 1902 Brinckiella Chopard, 1955 Brycoptera Ragge, 1981 Bueacola Sjöstedt, 1912 Chinensis Özdikmen, 2009 Chloroscirtus Saussure & Pictet, 1897 Choirorhynchus Piza, 1974 Conversifastigia Liu & Kang, 2008 Corycomima Karsch, 1896 Diastellidea Bolívar, 1902 Dicranopsyra Dohrn, 1892 Ectomoptera Ragge, 1980 Elbeniopsis Dohrn, 1906 Enthephippion Bruner, 1915 Ephippitythoidea Tepper, 1892 Eulioptera Ragge, 1956 Eulophophyllum Hebard, 1922 Euryastes Ragge, 1980 Euxenica Bruner, 1915 Execholyrus Henry, 1940 Gabonella Uvarov, 1940 Gatunella Uvarov, 1940 Gelatopoiidion Zacher, 1909 Gregoryella Uvarov, 1925 Harposcepa Karsch, 1896 Indogneta Ingrisch & Shishodia, 2000 Itokiia Sjöstedt, 1902 Ivensia Bolívar, 1890 Japygophana Carl, 1921 Khaoyaiana Ingrisch, 1990 Ladnea Walker, 1869 Lunidia Hemp, 2010 Macedna Karsch, 1891 Materuana Hemp, 2017 Meruterrana Sjöstedt, 1912 Milititsa Burr, 1900 Miltinobates Sjöstedt, 1902 Mimoscudderia Carl, 1914 Myllocentrum Ragge, 1962 Natricia Walker, 1869 Nesoscirtella Carl, 1914 Niphella Bolívar, 1900 Oxyecous Chopard, 1935 Paracaedicia Brunner von Wattenwyl, 1891 Parapelerinus Liu & Kang, 2008 Paraphylloptera Carl, 1914 Parapolichne Bolívar, 1902 Phaneroptila Uvarov, 1957 Physocorypha Karsch, 1896 Platycaedicia Hebard, 1922 Platylyra Scudder, 1898 Polygamus Carl, 1914 Procaedicia Bolívar, 1902 Pronomapyga Rehn, 1914 Pseudomacedna Liu, 2014 Pseudopreussia Hemp, 2017 Puerula Bolívar, 1906 Qinlingea Liu & Kang, 2007 Raggeiella Cadena-Castañeda, 2015 Raggophyllum Nickle, 1967 Sanabria Walker, 1869 Scolocerca Ragge, 1980 Semicarinata Liu & Kang, 2007 Sikoriella Carl, 1914 Stylomolpa Karny, 1926 Sympaestroides Willemse, 1942 Tamdaopteron Gorochov, 2005 Tropidonotacris Chopard, 1954
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