Capoeta

Capoeta, also known as scrapers, is a genus of fish in the family Cyprinidae found in Western Asia. The distribution extends from Turkey to the Levant, to Transcaucasia, Iraq, Turkmenistan, in Armenia, particularly in lake Sevan and northern Afghanistan.^[1][2] This genus is most closely related to Luciobarbus and in itself is divided into three morphologically, biogeographically and genetically distinct groups or clades: the Mesopotamian clade, the Anatolian-Iranian clade and the Aralo-Caspian clade.^[1][3][2] The Mesopotamian clade was split off to Paracapoeta in 2022.^[4]

Capoeta
Capoeta buhsei
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	Cypriniformes
Family:	Cyprinidae
Subfamily:	Cyprininae
Genus:	Capoeta Valenciennes, 1842
Type species
Cyprinus capoeta Güldenstädt, 1773
Synonyms
Scaphiodon Heckel, 1843

Species

There are currently 28 recognized species in this genus:^[5]

• Capoeta aculeata (Valenciennes, 1844)
• Capoeta alborzensis Jouladeh-Roudbar, Eagderi, Ghanavi & Doadrio, 2016^[6]
• Capoeta angorae (Hankó (hu), 1925) (Ankara barb)
• Capoeta antalyensis (Battalgil, 1944) (Pamphylian scraper, Antalya barb)
• Capoeta baliki Turan, Kottelat, Ekmekçi & İmamoğlu, 2006 (fourbarbel scraper, Sakarya barb)
• Capoeta banarescui Turan, Kottelat, Ekmekçi & İmamoğlu, 2006 (Colchic scraper, Banarescu's barb)
• Capoeta bergamae M. S. Karaman (sr), 1969 (Aegean scraper)
• Capoeta buhsei Kessler, 1877 (Namak scraper)
• Capoeta caelestis Schöter, Özuluğ & Freyhof, 2009 (Taurus scraper)
• Capoeta capoetoides Pellegrin, 1938
• Capoeta capoeta (Güldenstädt, 1773) (Caucasian scraper)
• Capoeta coadi Alwan, Zareian & Esmaeili, 2016^[7]
• Capoeta damascina (Valenciennes, 1842) (Levantine scraper, Mesopotamian barb)
• Capoeta ekmekciae Turan, Kottelat, Kırankaya & Engin, 2006 (Grusinian scraper)
• Capoeta ferdowsii Jouladeh-Roudbar, Eagderi, Murillo-Ramos, Ghanavi & Doadrio, 2017^[8]
• Capoeta fusca A. M. Nikolskii, 1897
• Capoeta gracilis (Keyserling, 1861)
• Capoeta kaput Levin, Prokofiev & Roubenyan 2019^[9]
• Capoeta kosswigi M. S. Karaman (sr), 1969 (Van scraper, Van barb)
• Capoeta mauricii Küçük, Turan, Şahin & Gülle, 2009 (longsnout scraper)
• Capoeta pestai (Pietschmann, 1933) (Eğirdir longsnout scraper, Eğirdir barb)
• Capoeta pyragyi Jouladeh-Roudbar, Eagderi, Murillo-Ramos, Ghanavi & Doadrio, 2017^[8]
• Capoeta razii Jouladeh-Roudbar, Eagderi, Ghanavi & Doadrio, 2017^[10]
• Capoeta saadii (Heckel, 1847)
• Capoeta sevangi De Filippi, 1865
• Capoeta shajariani Jouladeh-Roudbar, Eagderi, Murillo-Ramos, Ghanavi & Doadrio, 2017^[11]
• Capoeta sieboldii (Steindachner, 1864) (nipple-lip scraper)
• Capoeta tinca (Heckel, 1843) (western fourbarbel scraper, Anatolian khramulya)
• Capoeta umbla (Heckel, 1843)^[12] (Tigris scraper)

References

1. Ghanavi, Hamid Reza; Gonzalez, Elena G.; Doadrio, Ignacio (2016). "Phylogenetic relationships of freshwater fishes of the genus Capoeta (Actinopterygii, Cyprinidae) in Iran". Ecology and Evolution. 6 (22): 8205–8222. Bibcode:2016EcoEv...6.8205G. doi:10.1002/ece3.2411. PMC 5108271. PMID 27878089.
2. Levin, Boris A.; Freyhof, Jörg; Lajbner, Zdeněk; Perea, Silvia; Abdoli, Asghar; Gaffaroğlu, Muhammet; Özuluğ, Müfit; Rubenyan, Haikaz R.; Salnikov, Vladimir B. (January 2012). "Phylogenetic relationships of the algae scraping cyprinid genus Capoeta (Teleostei: Cyprinidae)". Molecular Phylogenetics and Evolution. 62 (1): 542–549. Bibcode:2012MolPE..62..542L. doi:10.1016/j.ympev.2011.09.004. hdl:10261/76088. PMID 21967785.
3. Zareian, H.; Esmaeili, H. R.; Heidari, A.; Khoshkholgh, M. R.; Mousavi-Sabet, H. (2016). "Contribution to the molecular systematics of the genus Capoeta from the south Caspian Sea basin using mitochondrial cytochrome b sequences (Teleostei: Cyprinidae)". Molecular Biology Research Communications. 5 (2): 65–75. PMC 5219896. PMID 28097160.
4. Turan, Davut; Kaya, Cüneyt; Aksu, İsmail; Bektaş, Yusuf (2022). "Paracapoeta, a new genus of the Cyprinidae from Mesopotamia, Cilicia and Levant (Teleostei, Cypriniformes)". Zoosystematics and Evolution. 98 (2): 201–212. doi:10.3897/ZSE.98.81463.
5. Froese, Rainer; Pauly, Daniel (eds.). "Species in genus Capoeta". FishBase. June 2024 version.
6. Jouladeh-Roudbar, A., Eagderi, S., Ghanavi, H.R. & Doadrio, I. (2016): Taxonomic review of the genus Capoeta Valenciennes, 1842 (Actinopterygii, Cyprinidae) from central Iran with the description of a new species. FishTaxa, 1 (3): 166-175.
7. Alwan, N.H., Zareian, H. & Esmaeili, H.R. (2016): Capoeta coadi, a new species of cyprinid fish from the Karun River drainage, Iran based on morphological and molecular evidences (Teleostei, Cyprinidae). ZooKeys, 572: 155–180.
8. Jouladeh-Roudbar, A., Eagderi, S., Murillo-Ramos, L., Ghanavi, H.R. & Doadrio, I. (2017): Three new species of algae-scraping cyprinid from Tigris River drainage in Iran (Teleostei: Cyprinidae). FishTaxa, 2 (3): 134-155.
9. Levin, BA; Prokofiev, AM; Roubenyan, HR (2019). "A New Species of Algae Eaters Capoeta kaput sp. nov. (Teleostei, Cyprinidae) from Transcaucasia". Inland Water Biology. 12 (1): 32–41. Bibcode:2019InWB...12...32L. doi:10.1134/S1995082919010139. ISSN 1995-0829. S2CID 146102213.
10. Jouladeh-Roudbar, A., Eagderi, S., Ghanavi, H.R. & Doadrio, I. (2017): A new species of the genus Capoeta Valenciennes, 1842 from the Caspian Sea basin in Iran (Teleostei, Cyprinidae). ZooKeys, 682: 137-155.
11. Jouladeh-Roudbar, A., Eagderi, S., Murillo-Ramos, L., Ghanavi, H.R. & Doadrio, I. (2017): Three new species of algae-scraping cyprinid from Tigris River drainage in Iran (Teleostei: Cyprinidae). FishTaxa, 2 (3): 134-155.
12. Esmaeili, H.R., Zareian, H., Eagderi, S. & Alwan, N. (2016): Review on the taxonomy of Tigris scraper, Capoeta umbla (Heckel, 1843) and its confirmation record from the Iranian part of Tigris River, Persian Gulf basin (Teleostei: Cyprinidae). FishTaxa, 1 (1): 35-44.