Acrocerinae

Acrocerinae is a subfamily of small-headed flies in the family Acroceridae. Their larvae are endoparasites of araneomorph spiders, with the exception of Carvalhoa appendiculata which can develop as ectoparasitoids on their host spiders.^[1] Traditionally, the subfamily included the genera now placed in Cyrtinae and Ogcodinae, but the subfamily in this sense was found to be polyphyletic and was split up in 2019.^[1][2]

Acrocerinae
Acrocera orbiculus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Diptera
Family:	Acroceridae
Subfamily:	Acrocerinae Leach, 1815
Genera
See text

Systematics

The subfamily includes two extant genera and one extinct:^[1]

• Acrocera Meigen, 1803
• Carvalhoa Koçak & Kemal, 2013^[3]
• †Schlingeromyia Grimaldi & Hauser in Grimaldi, Arillo, Cumming & Hauser, 2011^[4]

The extinct fly genus †Burmacyrtus Grimaldi & Hauser in Grimaldi, Arillo, Cumming & Hauser, 2011^[4] was originally placed in this subfamily as well, but according to Gillung & Winterton (2017) it is not considered an acrocerid.^[5]

References

1. Gillung, Jessica P.; Winterton, Shawn L. (2019). "Evolution of fossil and living spider flies based on morphological and molecular data (Diptera, Acroceridae)". Systematic Entomology. 44 (4): 820–841. Bibcode:2019SysEn..44..820G. doi:10.1111/syen.12358.
2. Stubbs, A.; Drake, M. (2014). British Soldierflies and their Allies (2 ed.). Wokingham: British Entomological and Natural History Society. p. 528. ISBN 9781899935079.
3. Koçak, A. Ö.; Kemal, M. (2013). "A nomenclatural note in the family Acroceridae (Diptera)". Centre for Entomological Studies Ankara Miscellaneous Papers (159): 3–4.
4. Grimaldi, DA; Arillo, A; Cumming, JM; Hauser, M (2011). "Brachyceran Diptera (Insecta) in Cretaceous ambers, Part IV, Significant New Orthorrhaphous Taxa". ZooKeys (148): 293–332. Bibcode:2011ZooK..148..293G. doi:10.3897/zookeys.148.1809. PMC 3264415. PMID 22287902.
5. Gillung, Jessica P.; Winterton, Shaun L. (2017). "A review of fossil spider flies (Diptera: Acroceridae) with descriptions of new genera and species from Baltic Amber". Journal of Systematic Palaeontology. 16 (4): 325–350. doi:10.1080/14772019.2017.1289566. S2CID 90493326.

Further reading

• Schlinger, Evert I.; Gillung, Jessica P.; Borkent, Christopher J. (2013). "New spider flies from the Neotropical Region (Diptera, Acroceridae) with a key to New World genera". ZooKeys (270): 59–93. Bibcode:2013ZooK..270...59S. doi:10.3897/zookeys.270.4476. PMC 3668421. PMID 23730188.
• Winterton, Shaun L.; Gillung, Jéssica P. (2012). "A new species of spider fly in the genus Sabroskya Schlinger from Malawi, with a key to Acrocerinae world genera (Diptera, Acroceridae)". ZooKeys (171): 1–15. Bibcode:2012ZooK..171....1W. doi:10.3897/zookeys.171.2137. PMC 3288680. PMID 22423193.