Citation. The Royal Society. 2004 [21 October 2012]. (原始内容存档于2020-02-07). Charlesworth, Deborah. Citation. The Royal Society. 2005 [21 October 2012]. (原始内容存档于2020-02-07)
Grafting. 2008-05-20. ISSN 0735-2689. doi:10.1080/07352680802053940. Charlesworth, Deborah; Willis, John H. The genetics of inbreeding depression. Nature