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relationships among arecoid palms (Arecaceae: Arecoideae). Annals of Botany（英语：Annals of Botany）, 108(8), 1417-1432. doi:10.1093/aob/mcr020 PDF Dransfield
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Seaweeds of the British Isles. Volume 3 Fucophyceae (Phaeophyceae) British Museum (Natural History)ISBN 0-565-00992-3 Jones, V. A. 2012. "Annals of Botany. July
Phylogenetic relationships among arecoid palms (Arecaceae: Arecoideae). Annals of Botany（英语：Annals of Botany）, 108(8), 1417-1432. doi:10.1093/aob/mcr020 PDF
J.M. & Chase, Mark W. Trends and concepts in fern classification. Annals of Botany. 2014, 113 (9): 571–594. PMC 3936591 . PMID 24532607. doi:10.1093/aob/mct299
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pdf （页面存档备份，存于互联网档案馆） Annals of Botany: The tannosome is an organelle forming condensed tannins in the chlorophyllous organs of Tracheophyta 色素体 鞣质 叶绿体