絛蟲綱(學名:Cestoda)是一個寄生性扁形動物門的一個綱[2][3]。人們常說的絛蟲其實是多個物種的泛稱。所有絛蟲綱的物種都是寄生蟲,其生活史可能各有不同,但這些物種的成蟲寄生於各類脊椎動物的消化道內,而其幼年期則寄生於其他動物的物種,例如:軟體動物。這些物種自身無消化道,體分節或不分節的扁形動物。
絛蟲的寄生歷程已有很長久歷史:科學家在一塊2.7億年前(二疊紀中期到後期)的鯊魚糞便化石(coprolites)裡發現了多叢絛蟲卵及一條發育中的蚴[1]。
分類
目前全球約有4000餘種,中國約有400種[4]。共分以下兩個亞綱:
- 單節絛蟲亞綱(Cestodaria)
- 多節絛蟲亞綱(Eucestoidea)
寄生於人體的絛蟲有30餘種,多屬圓葉目(Cyclophyllidea)和假葉目(Pseudophyllidea),最著名的種是引起人類絛蟲病的豬肉絛蟲[4]。
多節絛蟲亞綱物種的分類已澄清[5]。以下為現時的分類:
- 單殖綱 Monogenea
- Amphilinidea
- Caryophyllidea
- Spathebothriidea
- Caryophyllidea
- Amphilinidea
四葉目、假葉目(因為裂頭絛蟲科,Diphyllobothriidae)及圓葉目(因為Mesocestoididae科)均為並系群。
帶科(Taeniidae)可能是圓葉目之下12個目的最基底[5]。
The Tetraphyllidea, Lecanicephalidea, Proteocephalidea, Nippotaeniidea, Tetrabothriidea, and Cyclophyllidea are considered to be the 'higher' tapeworms.
海洋中的Trypanorhyncha目有277個已知物種,分為五個總科:Tentacularioidea, Gymnorhynchoidea, Otobothrioidea, Eutetrarhynchidae 和 Lacistorhynchidae.
- 在GBIF,本綱包括以下目:
- Amphilinidea
- Bothriocephalidea
- Caryophyllidea
- Cathetocephalidea
- Cathetocephalidae
- Disculicepitidae
- 圓葉目 Cyclophyllidea
- Diphyllidea
- Echinobothriidae
- 裂頭絛蟲目 Diphyllobothriidea
- Cephalochlamydidae
- 裂頭絛蟲科 Diphyllobothriidae
- Adenocephalus Nybelin, 1931
- Baylisia Markowski, 1952
- Baylisiella Markowski, 1952
- Dibothriocephalus Lühe, 1899
- Diphyllobothrium Cobbold, 1858
- Diplogonophorus Lönnberg, 1892
- Dubium Rudolphi, 1819
- Flexobothrium Yurakhno, 1989
- Glandicephalus Fuhrmann, 1921
- Hexagonoporus Gubanov, 1955
- Ligula Bloch, 1782
- Ligula alternans Rudolphi, 1810
- Ligula avium Bloch, 1782
- Ligula colymbi Zeder, 1803
- Ligula digramma Creplin, 1839
- Ligula interrupta Rudolphi, 1810
- Ligula intestinalis (Linnaeus, 1758) Gmelin, 1790[6]
- Ligula monogramma Creplin, 1839
- Ligula nemachili (Dubinina, 1957) Kuchta & Scholz, 2017
- Ligula pancerii Polonio, 1860
- Ligula pavlovskii Dubinina, 1959
- Multiductus Clarke, 1962
- Plicobothrium Rausch & Margolis, 1969
- Pyramicocephalus Monticelli, 1890
- Schistocephalus Creplin, 1829
- Sparganum Diesing, 1854
- Spirometra Faust, Campbell & Kellogg, 1929
- Tetragonoporus Skryabin, 1961
- Solenophoridae
- Gyrocotylidea
- Haplobothriidea
- Lecanicephalidea
- Aberrapecidae
- Cephalobothiidae
- Cephalobothriidae
- Eniochobothriidae
- Lecanicephalidae
- Paraberrapecidae
- Polycephalidae
- Polypocephalidae
- Anteropora Subhapradha, 1955
- Anthemobothrium Shipley & Hornell, 1906
- Corollapex Herzog & Jensen, 2017
- Hornellobothrium Shipley & Hornell, 1906
- Polypocephalus Braun, 1878[7]
- Polypocephalus affinis Subhapradha, 1951
- Polypocephalus alii Shinde & Jadhav, 1981
- Polypocephalus barsami Roudsari & Haseli, 2020
- Polypocephalus bombayensis Shinde, Dhule & Jadhav, 1991
- Polypocephalus budhadebae Jadhav, 2007
- Polypocephalus caribbensis (Gardner & Schmidt, 1984) Jensen, 2005
- Polypocephalus coronatus Subhapradha, 1951
- Polypocephalus digholensis Deshmukh, Jadhav & Shinde, 1982
- Polypocephalus djeddensis Jadhav & Shinde, 1989
- Polypocephalus elongatus (Southwell, 1912) Jensen, 2005
- Polypocephalus harnesis Lanka, Pathan & Jadhav, 2006
- Polypocephalus helmuti Jensen, 2005
- Polypocephalus himanshui Pramanik & Manna, 2006
- Polypocephalus indicus Deshmukh, Jadhav & Shinde, 1982
- Polypocephalus karbharii Deshmukh, Jadhav & Shinde, 1982
- Polypocephalus katpurensis Shinde & Jadhav, 1981
- Polypocephalus kuhlii Vankara, Vijayalakshmi & Vijayalakshmi, 2006
- Polypocephalus lintoni Subhapradha, 1951
- Polypocephalus maharashtra Deshmukh, Jadhav & Shinde, 1982
- Polypocephalus medusia (Linton, 1890) Southwell, 1925
- Polypocephalus mirkarwarensis Dandawate & Jadhav, 2009
- Polypocephalus moretonensis Butler, 1987
- Polypocephalus prathibhai Deshmukh, Jadhav & Shinde, 1982
- Polypocephalus radiatus Braun, 1878
- Polypocephalus ratnagiriensis Jadhav, Shinde & Sarwade, 1986
- Polypocephalus rhinobatidis Subhapradha, 1951
- Polypocephalus rhynchobatidis Subhapradha, 1951
- Polypocephalus saoudi Hassan, 1982
- Polypocephalus singhii Shinde & Jadhav, 1981
- Polypocephalus thapari Shinde & Jadhav, 1981
- Polypocephalus vesicularis Yamaguti, 1960
- Polypocephalus visakhapatnamensis Vankara, Lakshmi & Lakshmi, 2007
- Polypocephalus vitellaris Subhapradha, 1951
- Polypocephalus waltairensis Mote, 2012
- Seussapex Jensen & Russell, 2014
- Thysanobothrium Shipley & Hornell, 1906
- Tetragonocephalidae
- Zanobatocestidae
- Litobothriidea
- Nippotaeniidea
- Onchoproteocephalidea
- Phyllobothriidea
- Chimaerocestidae
- Phyllobothriidae
- Alexandercestus Ruhnke & Workman, 2013
- Anindobothrium
- Anthobothrium
- Aocobothrium
- Bibursibothridium McKenzie & Caira, 1998
- Bibursibothrium McKenzie & Caira, 1998
- Biporophyllaeus Subramaniam, 1939
- Calyptrobothrium Monticelli, 1893
- Cardiobothrium McKenzie & Caira, 1998
- Caulobothrium
- Caulopatera Cutmore, Bennett & Cribb, 2010
- Clistobothrium Dailey & Vogelbein, 1990
- Clistobothrium amyae Caira, Hayes & Jensen, 2020
- Clistobothrium carcharodoni Dailey & Vogelbein, 1990[8]
- Clistobothrium gabywalterorum Caira, Hayes & Jensen, 2020
- Clistobothrium montaukense Ruhnke, 1993
- Clistobothrium tumidum (Linton, 1922) Ruhnke, 1993
- Crossobothrium Linton, 1889
- Cyatocotyle Mola, 1908
- Dinobothrium
- Flexibothrium McKenzie & Caira, 1998
- Gastrolecithus
- Guidus Ivanov, 2006
- Hemipristicola Cutmore, Theiss, Bennett & Cribb, 2011
- Hoaleshwaria
- Kowsalyabothrium Muralidhar, Shinde & Jadhav, 1987
- Marsupiobothrium Yamaguti, 1952
- Monorygma Diesing, 1863
- Orygmatobothrium Diesing, 1863
- Pelichnibothrium Monticelli, 1889
- Pentaloculum Alexander, 1963
- Phanobothrium
- Phormobothrium Alexander, 1963
- Phyllobothrium Van Beneden, 1850
- Pillersium Southwell, 1927
- Pithophorus Southwell, 1925
- Polipobothrium
- Rockacestus Caira, Bueno & Jensen, 2021
- Ruhnkebothrium Caira, Bueno & Jensen, 2021
- Scyphophyllidium Woodland, 1927
- Shindeiobothrium
- Thysanocephalum Linton, 1890
- Tiarabothrium Shipley & Hornell, 1906
- Yamaguticestus Caira, Bueno & Jensen, 2021
- Zyxibothrium Hayden & Campbell, 1981
- Rhinebothriidea
- Spathebothriidea
- Acrobothriidae
- Diplocotylidae
- Spathebothriidae
- Tetrabothriidea
- Tetrabothriidae
- Anophryocephalus Baylis, 1922
- Bothridiotaenia Lönnberg, 1896
- Chaetophallus Nybelin, 1916
- Dendrometra Jordano & Diaz-Ungria, 1956
- Priapocephalus Nybelin, 1922
- Strobilocephalus Baer, 1932
- Tetrabothrium Blainville, 1828
- Tetrabothrius Rudolphi, 1819
- Trigonocotyle Baer, 1932
- Tetrabothriidae
- 四葉目 Tetraphyllidea
- 錐吻目 Trypanorhyncha
- 目的地位未定的科:
- Ballianiidae
- Pseudophyllidae
- Trilocularidae
- 目和科的地位未定的屬:
- Acanthorhynchus Diesing, 1850
- Acrostoma Le Sauvage, 1827
- Alyselminthus Zeder, 1800
- Amphicephalus Beneden, 1871
- Aprostatandrya Spasski, 1951
- Atriotaenia Spasski, 1951
- Balanoforus Briganti, 1825
- Bicorne Sultzer, 1801
- Bicornius Rafinesque, 1815
- Bothriocephalum Lühe, 1899
- Bothriorhynchus Lidth de Jeude, 1829
- Bothriotaenia Railliet, 1892
- Bothryocephalus Cuvier, 1817
- Butleroides Lordello & Zamith, 1959
- Caryophillinus Schrank, 1788
- Caryophyllus Rudolphi, 1809
- Catenaria Zeder, 1800
- Cephalocotyleum Diesing, 1850
- Cercocystis Villot, 1882
- Cestoideum Diesing, 1854
- Cestoscolex Parona, 1887
- Chitonorecta Meggitt, 1927
- Cisticercoideum Parona, 1901
- Corynesoma Leuckart, 1843
- Cotylogenes Lühe, 1902
- Cryptocephalus van Beneden, 1849
- Cryptocystis Villot, 1883
- Cucurbilepsis Sadovskaya, 1965
- Cucurbitina Alexander, 1833
- Cucurbitinus Gleichen, 1779
- Cucurbitinus Kniphof, 1759
- Cylindrocorpus Rühm, 1959
- Cysticercus Zeder, 1800
- Cysturus Oken, 1815
- Davainoides Fuhrmann, 1921
- Dibothria Rudolphi, 1809
- Dibothriorhyncus Chiaje, 1831
- Dibothrium Diesing, 1850
- Dibothrium Rudolphi, 1839
- Dibothrius Rudolphi, 1819
- Dibothrydus
- Diorchis Matevosyan, 1946
- Diplasia Holten, 1802
- Diporus Diesing, 1863
- Diprognopolus Anon., 1904
- Dithyridium Rudolphi, 1819
- Dithyridium Valenciennes, 1844
- Dittocephalus Parona, 1887
- Echinobothria Rudolphi, 1810
- Echinobothrion Rudolphi, 1809
- Echinoproboscilepsis Sadovskaya, 1965
- Fascia Blanchard, 1888
- Furia Linnaeus, 1758
- Gieystoria
- Globus Scopoli, 1772
- Gopalaia Dixit, Capoor & Rengaragu, 1980
- Guptaia Malhotra, 1985
- Gvosdevileps Spasskii, 1953
- Gymnobothrion Rudolphi, 1809
- Gymnoscolex Diesing, 1853
- Gymnotenia Beneden, 1873
- Gyrarthrix
- Halysis Zeder, 1803
- Heterodera Skarbilovich, 1959
- Hydatodes Abildgaard, 1790
- Ilisha Southwell & Prashad, 1918
- Lakistorhynchus Pintner, 1913
- Luheella Baer, 1924
- Maccallumiella Yamaguti, 1952
- Mastacembellophyllaeus Shinde & Chincholikar, 1977
- Merocestus Cohn, 1902
- Metabelea Mettrick, 1963
- Monocercus Villot, 1882
- Monogramma Cholodkovsky, 1915
- Notobothrium Linstow, 1905
- Octobothrium Diesing, 1850
- Onchoscolex Diesing, 1853
- Oobothrium Linton, 1904
- Orygmatoscolex Diesing, 1854
- Palaia Shipley, 1900
- Pancrates Gistl, 1848
- Parvitaenio Burt, 1940
- Piestocystis Diesing, 1850
- 平槽屬 Platybothrium Hornell & Nayudu, 1924
- Plerocercoides
- Proglottis Dujardin, 1843
- Prosthecocotyle Monticelli, 1892
- Pseudobothrium
- Pseudocapingentoldes Verma, 1971
- Pseudodiscus Polonio, 1860
- Pseudonybelinla Dollfus, 1966
- Pseudoscicus Polonio, 1860
- Sagittula Lamarck, 1816
- Skrjabinoparaksis Krotov, 1949
- Slossia Meggitt, 1931
- Solium Blanchard, 1886
- Spasskijela Tenora, 1959
- Sphairidion Goodsir, 1845
- Steganobothrium Diesing, 1854
- Stenotenia Gervais, 1870
- Strobilocercus Sambon, 1924
- Synbia Tschudi, 1837
- Taeniea Braun, 1894
- Taxocyclocotyle Diesing, 1854
- Terratas Blanchard, 1886
- Tetrabothriorhynchus Diesing, 1850
- Tetracisdicotyla Fuhrmann, 1907
- Tetracotylus Monticelli, 1892
- Tetrarhyncha
- Tetrastoma Forbes, 1840
- Tetrathyridium Rudolphi, 1819
- Tetrathyrus Creplin, 1851
- Thornonema Andrássy, 1959
- Urocystidium Beddard, 1912
- Versicaria Baird, 1853
特徵
成蟲扁長如腰帶,分節,白色或乳白色,體長因蟲種不同可從數毫米至數米不等。由頭節(scolex)、頸部(neck)、節片(proglottid)組成,前端細。 頭節上有槽、吸盤或鉤,藉此附著在宿主腸壁上;節片由頸分裂而來,節片由一至數千不等,越往後越寬大;每一節片相當一個體,稱鏈體(stroblius),鏈體是蟲體最顯著的部分,其中至少有一套雌雄生殖器官,其他器官退化,營養靠體表吸收宿主的養料;隨著生殖器官的發育,節片愈後愈成熟,有卵的節片稱為孕節,最後逐節或整段脫落,隨糞便排出宿主體外。
生活史
卵內具有鉤的胚,胚逸出後侵入中間宿主,發育為實心或囊狀的蚴,蚴被最終宿主吞食後,即在小腸內成長為成蟲;例如寄生人體內的豬肉絛蟲、牛肉絛蟲。
參考文獻
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