肺螺类 (学名:Pulmonata ),亦作有肺类 ,旧作有肺目 、肺螺目 或肺螺亚纲 ,是腹足纲 异鳃类 支序之下的一个非正式群组。这个群组包括有螺 和蛞蝓 ,均可以透过其外套膜肺 (pallial lung)而不是鳃 来呼吸空气。本群组除了包括有蜗牛 和淡水螺 ,还包括多个海螺的科。
事实速览 科学分类, 类群 ...
有肺类
科学分类
界:
动物界 Animalia
门:
软体动物门 Mollusca
纲:
腹足纲 Gastropoda
演化支 :
直腹足类 Orthogastropoda
演化支 :
后生腹足类 Apogastropoda
亚纲:
异鳃亚纲 Heterobranchia
非正式类群 :
有肺类 Pulmonata Cuvier , 1814
类群
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传统定义的肺螺类分类单元 的物种在Jörger et al. (2010)被发现原来是个多系群 [ 1] 。肺螺类已知从石炭纪 开始就已经开始出现,直到现在[ 2] 。
肺螺类物种有一个心室和一个肾脏,还有一个相互连接、左右对称的神经系统。外套膜腔位于身体的右侧,而且没有鳃,而是演变成一个布满血管的肺。大多数物种都有壳 ,但没有口盖,而且本分类元还包括有几种没有外壳的蛞蝓。肺类类物种都雌雄间性 ,部分群组还拥有用作交配的恋矢 (love darts)[ 3]
在原来的《旁得和林德伯格的腹足类分类 (1997年) 》,本支序是一个目级分类元,包括下列各个分类元:
收眼亚目 Systellommatophora Pilsbry, 1948
基眼亚目 Basommatophora Keferstein in Bronn, 1864:淡水螺及池塘的螺
真有肺亚目 Eupulmonata Haszprunar & Huber, 1990
Infraorder Acteophila Dall , 1885 (= formerly Archaeopulmonata)
Superfamily Melampoidea Stimpson, 1851
Infraorder Trimusculiformes Minichev & Starobogatov, 1975
柄眼下目 Stylommatophora A. Schmidt, 1856 Shells of pulmonate styllomatophoran snails in a museum collection An artistic but scientifically incorrect version of various European land snails and slugs (one species here is not a pulmonate), their food plants and fungi, and a beetle that eats mollusks, bottom right.
直尿道小目 Orthurethra
Superfamily Achatinelloidea Gulick, 1873
Superfamily Cochlicopoidea Pilsbry, 1900
Superfamily Partuloidea Pilsbry, 1900
Superfamily Pupilloidea Turton, 1831
弯尿道小目 Sigmurethra
Superfamily Acavoidea Pilsbry, 1895
非洲大蜗牛总科 Achatinoidea Swainson, 1840
Superfamily Aillyoidea Baker, 1960
Superfamily Arionoidea J.E. Gray in Turnton, 1840
Superfamily Buliminoidea Clessin, 1879
Superfamily Camaenoidea Pilsbry, 1895
烟管蜗牛总科 Clausilioidea Mörch , 1864
Superfamily Dyakioidea Gude & Woodward, 1921
Superfamily Gastrodontoidea Tryon, 1866
Superfamily Helicoidea Rafinesque , 1815
Superfamily Helixarionoidea Bourguignat, 1877
Superfamily Limacoidea Rafinesque , 1815
Superfamily Oleacinoidea H. & A. Adams, 1855
Superfamily Orthalicoidea Albers-Martens, 1860
Superfamily Plectopylidoidea Moellendorf, 1900
Superfamily Polygyroidea Pilsbry, 1894
Superfamily Punctoidea Morse, 1864
Superfamily Rhytidoidea Pilsbry, 1893
Superfamily Sagdidoidera Pilsbry, 1895
Superfamily Staffordioidea Thiele , 1931
Superfamily Streptaxoidea J.E. Gray , 1806
Superfamily Strophocheiloidea Thiele , 1926
Superfamily Trigonochlamydoidea Hese, 1882
Superfamily Zonitoidea Mörch , 1864
? Superfamily Athoracophoroidea P. Fischer, 1883 = Tracheopulmonata
? Superfamily Succineoidea Beck, 1837 = Heterurethra
Examples of Pulmonata: Achatina fulica top right, Bielzia coerulans top left, Praticolella berlandieriana center right, Megalobulimus oblongus in the center, Euglandina rosea center left, Helix pomatia bottom right & Ashmunella levettei bottom left
根据《布歇特和洛克罗伊的腹足类分类 (2005年) 》,Glacidorboidea 被搬到异鳃类 支序之下的下异鳃类 群组,本分类余下下列各个分类[ 4] :
基眼类 (Basommatophora)非正式群组包括喜湿螺类 (Hygrophila)支序和下列两个总科:
两栖螺总科 Amphiboloidea J.E. Gray , 1840
松螺总科 Siphonarioidea J.E. Gray , 1840
松螺科 Siphonariidae
† Family Acroreiidae
喜湿螺类 (Hygrophila)支序包括下列四个总科:
斜顶螺总科 Acroloxoidea Thiele , 1931
斜顶螺科 Acroloxidae Thiele, 1931
Superfamily Chilinoidea Dall, 1870
Family Chilinidae Dall, 1870
Family Latiidae Hutton, 1882
椎实螺总科 (Lymnaeoidea Rafinesque , 1815)
扁蜷总科 (Planorboidea Rafinesque, 1815)
真有肺类 (Eupulmonata)支序包括the clades Systellommatophora and Stylommatophora
收眼类 支序(Systellommatophora = Gymnomorpha )之下有两总科[ 5] ,分别为:
石磺总科 Onchidioidea Rafinesque , 1815
石磺科 Onchidiidae Rafinesque, 1815
皱足蛞蝓总科 Veronicelloidea Gray, 1840
皱足蛞蝓科 Veronicellidae Gray, 1840
拉索蛞蝓科 Rathouisiidae Heude, 1885
Contains the subclades Elasmognatha , Orthurethra and the informal group Sigmurethra
Two superfamilies belongs to clade Sigmurethra, but they are not in the limacoid clade.
Superfamily Arionoidea
Family Arionidae
Family Anadenidae
Family Ariolimacidae
Family Binneyidae
Family Oopeltidae
Family Philomycidae
Superfamily Helicoidea
Family Helicidae
Family Bradybaenidae
Family Camaenidae
Family Cepolidae
Family Cochlicellidae
Family Elonidae
Family Epiphragmophoridae
Family Halolimnohelicidae
Family Helicodontidae
Family Helminthoglyptidae
Family Humboldtianidae
Family Hygromiidae
Family Monadeniidae
Family Pleurodontidae
Family Polygyridae
Family Sphincterochilidae
Family Thysanophoridae
Family Trissexodontidae
Family Xanthonychidae
Jörger et al. (2010)[ 1] analyzed major groups within the Heterobranchia using genetic data and found that Pulmonata as traditionally defined was polyphyletic , for instance some pulmonates were more closely related to Sacoglossa and Acochlidia . They proposed the more inclusive taxon Panpulmonata to unite the clades Siphonarioidea , Sacoglossa , Glacidorboidea , Pyramidelloidea , Amphiboloidea , Hygrophila , Acochlidia and Eupulmonata .[ 1]