Haldane, J.B.S. (1924) "A mathematical theory of natural and artificial selection" Part 1 Transactions of the Cambridge philosophical society: 23: 19-41 link (pdf file) (页面存档备份,存于互联网档案馆)
Hamilton, W.D. (1964) "The evolution of social behavior" Journal of Theoretical Biology 1:...
Hartl, D. L. A Primer of Population Genetics. Sinauer, Sunderland, Massachusetts, 1981.
Maynard Smith, J. Evolutionary Genetics. Oxford University Press, 1998.
延伸阅读
Sober, E. (2001). The Two Faces of Fitness. In R. Singh, D. Paul, C. Krimbas, and J. Beatty (Eds.), Thinking about Evolution: Historical, Philosophical, and Political Perspectives. Cambridge University Press, pp.309-321. Full text