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proteins）的反式作用因子。在哺乳动物中，UPF2和UPF3为外显子连接复合体（英语：Exon junction complex）（英語：exon-exon junction complex）与在RNA剪接（英語：RNA Splicing）后mRNA连接的部分。其它诸如eIF4AIII、
Pedersen JS, Séraphin B, Le Hir H, Andersen GR. Structure of the exon junction core complex with a trapped DEAD-box ATPase bound to RNA. Science. 2006, 313: