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吸蟲(學名:Trematoda)是寄生蟲的一種,為扁形動物門吸蟲綱動物的總稱,也稱為瓜仁蟲。一些吸蟲也被稱為二口蟲(拉丁語:Distoma)。其名由來是因為它的口吸盤和腹吸盤都被認為是口而致,拉丁語中,Di代表「二」,stoma代表「口」。

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吸蟲綱
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Botulus microporus,一種生活在帆蜥魚腸子內的大型吸蟲
科學分類 編輯
界: 動物界 Animalia
門: 扁形動物門 Platyhelminthes
亞門: 被桿體亞門 Rhabditophora
演化支 無中心體類 Acentrosomata
演化支 裂腸新皮類 Bothrioneodermata
總綱: 新皮總綱 Neodermata
綱: 吸蟲綱 Trematoda
Karl Rudolphi英語Rudolphi, 1808
亞綱

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概述

吸蟲是非常有名的一種寄生蟲,大部分為雌雄同體[1],可自體受精;生活史中可能有一個或多個中間宿主,以魚類人體等多種脊椎動物為最終宿主;成蟲產卵發育成纖毛幼蟲(Miracidium,亦作毛蚴),纖毛幼蟲入侵中間宿主後,增殖並發育成尾動幼蟲(cercaride,亦作尾蚴),再被最終宿主食入前會發育成曩狀幼蟲(metacerariae,亦作曩蚴[2]。吸蟲的成蟲有口吸盤及腹吸盤,其卵有卵蓋(operculum)[2],蟲體在人體中最久可活30年,由生食淡水魚而感染。貓、狗為一種名為保蟲的吸蟲宿主,成蟲寄生於膽道,大量感染時可在膽囊中發現蟲體。曩狀幼蟲存在於淡水魚時,醃漬、煙燻或脫水皆不能將其殺死,輕微感染時無症狀,重度感染會造成膽道阻塞、膽囊炎結石肝膿痬急性胰臟炎等,蟲卵約於感染後1個月後可於糞便中發現[3]

分類

吸蟲綱物種原來分為單殖目(Monogena)、複殖目(Digenea)、盾腹目(Aspidogastrea,亦作盾盤目楯盤目)三個目[1]。單殖、複殖所指的是物種的雄性生殖系統的輸精管的數目(單殖1個、複殖2個);而盾腹目的命名則沿於其特徵,在其腹口吸盤周圍有一個盾牌狀的襯墊。當時的單殖目可再細分為三個亞目[1]:259-260,但現時單殖目已被提升至綱級,成為單殖綱。餘下的兩個目合計的物種,推算約 18,000[4]到 24,000[5]個物種。有文獻把這兩個目提升到亞綱的地位。

複殖目又名二生目,是吸蟲綱絕大多數物種所屬,之下再分為腹口亞目(Gasterostomata,寄生於魚腸內)及前口亞目(Prosostomata,有59科)[1],其物種泛稱複殖吸蟲(digenetic trematode)是以寄生人體為對象的吸蟲。複殖吸蟲的基本結構和發育過程大致上相同,種類相當繁多且形態各自相差頗大,生活史有性世代(寄生於軟體動物,如螺螄、蚌類的階段)與無性世代(寄生於脊椎動物體內階段),基本生活型態是以卵生且多種幼蟲(毛蚴胞蚴雷蚴尾蚴囊蚴)之後在成長為成蟲[6]。這些吸蟲中以人類為宿主[1]的種類,有肺吸蟲(衛氏肺吸蟲)、肝吸蟲(中華肝吸蟲牛羊肝吸蟲)、腸道吸蟲橫川吸蟲血吸蟲日本血吸蟲)、布氏薑片蟲(Fasciolopsis buski)等。

楯盤目只有一科四屬,直接發育,無需中間宿主。今提升至亞綱級,是為楯吸蟲亞綱楯盤亞綱

本綱包括以下目:
  • Aspidogastrida
    • Aspidogasteridae
    • Aspidogastridae
    • Multicalycidae
    • Rugogastridae
  • 有壁目 Diplostomida
  • 斜睪目 Plagiorchiida
  • Stichocotylida
    • Stichocotylidae
目的地位未定的科:
  • Acanthostomatidae
  • Albulatrematidae
  • Areobiotrematidae
  • Arnolidae
  • Gekkonotrematidae
  • Halipegidae
  • Jubilariidae
  • Lecithochiridae
  • Lobatovitelliovariidae
  • Oesophagicolidae
  • Sclerodistomatidae
  • Tetrasteridae
目、科的地位未定的屬:
  • Acanthocephala Diesing, 1858
  • Achillurbainia Dollfus, 1939
  • Adolescaria Sinitzin, 1911
  • Afrobilharzia Le Roux, 1958
  • Agamodistomum Stossich, 1892
  • Ahemiuris Chauhan, 1954
  • Allomicrophallus Yamaguti, 1971
  • Allopseudopiclidophora Yamaguti, 1965
  • Alloschistorchis Yamaguti, 1970
  • Allotelorchis Goodman, 1988
  • Amphibothrium Frey & Leuckart, 1847
  • Amphistomulum Brandes, 1892
  • Amphocleithrium Price & Romero, 1969
  • Anatirenicola Odening, 1962
  • Ankyrocotyle Vlassenko, 1928
  • Antepharyngeum Witenberg, 1925
  • Antodera Pigulevsky, 1952
  • Aototrema Bean-Knudsen, Uhazy & Wagner, 1987
  • Aspalacitrema Deblock & Rausch, 1965
  • Aspina Tripathi, 1956
  • Astomum Schlotthauber, 1860
  • Asymphylodoroides Yamaguti, 1971
  • Athesmioides Cribb & Spratt, 1992
  • Atrioglabrum Gracenea, Montoliu & Deblock, 1993
  • Atriospinosum Deblock, 1973
  • Australodiscus Sey, 1983
  • Axine Yamaguti, 1938
  • Baiohelmins Cribb & Pearson, 1988
  • Bashkinovitrema Skrjabin, 1944
  • Bathycola Fuhrmann, 1928
  • Baughbilharzia Chauhan, Srivastava & Chauhan, 1973
  • Beauchampia Yamaguti, 1963
  • Bengaliniella Deblock, Mukherjee & Ghosh, 1970
  • Bilateracotyloides Ramalingam, 1961
  • Bilorchis Mehra, 1937
  • Biplacunella Unnithan, 1971
  • Bispina Tripathi, 1956
  • Bistellavulva Unnithan, 1971
  • Bivulvostella Unnithan, 1971
  • Blairium Brooks & Holcman, 1993
  • Bothriophoron Stiles & Goldberger, 1910
  • Brachionus Schrank, 1776
  • Brachylecithoides Yamaguti, 1971
  • Brachysaccus Johnston, 1912
  • Bravotrema Groschaft, 1970
  • Bullulina Hammerschmidt, 1838
  • Buxifrons Fukui, 1929
  • Buxifrons Näsmark, 1937
  • Bychowskicotyle Unnithan, 1971
  • Calicotyle Nybelin, 1941
  • Calicotylides Nybelin, 1941
  • Canaania Travassos, 1944
  • Capronia Richard, 1966
  • Carangixenus Unnithan, 1962
  • Catoptroides Looss, 1902
  • Caudorchis Talbot, 1933
  • Cauliorchis Stiles & Goldberger, 1910
  • Cesartrema Travassos & Kohn, 1964
  • Ceylonotrema Gussev, 1963
  • Chamaeleophilus Yamaguti, 1971
  • Chaseostrigea Ukoli, 1967
  • Cheilostomum Diesing, 1850
  • Chelonella Beneden & Hesse, 1863
  • Chinhuta Lal, 1937
  • Chiropterotarbus Bay-Schmith, 1972
  • Chiropterotrema Velez & Thatcher, 1992
  • Chiroptodendrium Skarbilovitch, 1943
  • Choanocotyle Jue Sue & Platt, 1998
  • Choerocotyle Baer, 1959
  • Choerocotyloides Prudhoe, Yeh & Khalil, 1964
  • Choristogonoporus Stunkard, 1938
  • Cladorchis Wang & Zhou, 1989
  • Cochinocotyle Gupta & Gupta, 1971
  • Coelomotrema Angel, 1970
    • Coelomotrema aegyptiaca[7]
  • Coitocoecum Nicoll, 1915
  • Combesia Mas-Coma, Roset & Mantoliu, 1985
  • Conspicum Bhalerao, 1936
  • Coryphogonimus Cribb & Spratt, 1990
  • Cotylegaster Cobbold, 1877
  • Cryptocephalus Srivastava, 1944
  • Cryptostomum Nitzsch, 1832
  • Cychlorchis Lühe, 1908
  • Cyclostoma Otto, 1823
  • Cymbiforma Yamaguti, 1933
  • Dactylodiscus Kamegai, 1971
  • Dasyurotrema Cribb, 1992
  • Decacotylus Mayer, 1841
  • Dermophthiroides Cheung & Nigrelli, 1983
  • Diagenites Poinar & Boucot, 2006
  • Diastomum Packard, 1882
  • Dicotyle Sonsino, 1898
  • Dicranocercaria Lutz, 1919
  • Dicranocercaria Szidat, 1932
  • Digenites Poinar & Boucot, 2006
  • Dinotaster Unnithan, 1971
  • Diplobothrium Chandler, 1942
  • Diplostomum Brandes, 1888
  • Diplotrema Tripathi, 1959
  • Diporpa Dujardin, 1844
  • Distemum
  • Distoma Retzius, 1786
  • Distomulum Brandes, 1892
  • Distomum Diesing, 1850
  • Distomus Laennec, 1807
  • Divitelluses Cai & Li, 1993
  • Dolichodemas Looss, 1900
  • Dolichosomum Looss, 1899
  • Dozenkotrema Onkarnatha, 1964
  • Duttiella Srivastava, Prasad & Maurya, 1980
  • Echinella van Beneden & Hesse, 1864
  • Edcaballerotrema Teixeira de Freitas, 1960
  • Eroliostrigea Yamaguti, 1971
  • Eucercaria Diesing, 1855
  • Eucercarioides Tang & Tang, 1992
  • Eucradium Szidat, 1954
  • Eurobilharzia Le Roux, 1958
  • Eurymetra Odhner, 1910
  • Eustemma Diesing, 1850
  • Eustomos MacCallum, 1921
  • Evandrocotyle Jansen, 1941
  • Evanorchis Skrjabin, 1944
  • Exorchocoelium Thapar, 1958
  • Falciungus Akmerov, 1952
  • Festucaria Schrank, 1788
  • Flabellodiscus Johnston & Tiegs, 1922
  • Fontius Lotz & Palmieri, 1985
  • Gastrothylacias Stiles & Goldberger, 1910
  • Gastrothylacides Stiles & Goldberger, 1910
  • Geaterinia Lebedev, 1969
  • Gemellicotyle Prudhoe, 1975
  • Gibsonium Brooks & Holcman, 1993
  • Gigantatrium Yamaguti, 1958
  • Gigantobilharziella Khalifa, 1974
  • Gigantopora Looss, 1907
  • Glenocercaria Diesing, 1858
  • Globostomum Oshmarin, 1970
  • Globularia Rolando, 1841
  • Glossidioides Yamaguti, 1958
  • Glossocotyle Van Beneden & Hesse, 1863
  • Glyptamphistoma Yamaguti, 1958
  • Gotocotyla Meserve, 1938
  • Gymnocephala Diesing, 1858
  • Gymnocephala Leidy, 1877
  • Haematobium Dunglison, 1893
  • Haplostomum Burmeister, 1856
  • Harmostomum Witenberg, 1925
  • Hawkesius Stiles & Goldberger, 1910
  • Hemistephanus Looss, 1901
  • Hemixenotrema Baer, 1971
  • Hepatotrema Stunkard, 1923
  • Hephostomum Burmeister, 1856
  • Heptostomum Schomburgk, 1844
  • Heteracanthus Diesing, 1836
  • Heteraxine Linton, 1940
  • Heterobdella Beneden & Hesse, 1864
  • Heterocoelium Travassos, 1921
  • Heterocotyloides Yamaguti, 1963
  • Heterolope Looss, 1899
  • Heterostoma De Filippi, 1837
  • Hexastoma Kuhn, 1829
  • Hexastoma Rudolphi, 1809
  • Hexathyridium Treutler, 1793
  • Histerolecitha Linton, 1910
  • Histrionella Bory de St.Vincent, 1823
  • Histrionella Ehrenberg, 1828
  • Histrionellina Diesing, 1858
  • Holostoma Nitzsch, 1816
  • Hormocercaria Diesing, 1855
  • Hydricuculus McCrady, 1873
  • Hydrochoeristrema Sinkoc, Muller & Brum, 1998
  • Hypostoma Rudolphi, 1809
  • Indopleurogenes Yamaguti, 1971
  • Indopodocotyle Gupta & Puri, 1981
  • Indopodocotyloides Gupta & Sayal, 1979
  • Ishiella Unnithan, 1971
  • Ityogonismus
  • Jamuartrema Kumar Lal, 1976
  • Kaurma Chatterji, 1936
  • Khalilloossia Hilmy, 1949
  • Klepsitromis Hammerschmidt, 1835
  • Kunckelia Künstler, 1881
  • Labontes MacCallum, 1927
  • Labriferoides Ganapati, Hanumantha & Lakshmana Rao, 1962
  • Laterostrigea Yang, 1962
  • Leiperocotyle Eduardo, 1980
  • Leipertrema Sandosham, 1951
  • Lepidotes Johnston & Tiegs, 1922
  • Lepustomum Jain, 1953
  • Levinseniella Deblock & Pearson, 1971
  • Levinsensia Looss, 1928
  • Limatuloides Dubois, 1964
  • Limosina Manter, 1944
  • Lintonia Monticelli, 1904
  • Lobostoma Berthold, 1827
  • Longitrema Chen, 1954
  • Lophocercaria Diesing, 1858
  • Loriformis Goeze, 1782
  • Lutianicotyle Mamaev, 1990
  • Lutziella Rohde, 1966
  • Luxitrema Bray, 1987
  • Lyperosomoides Yamaguti, 1971
  • Macallumia Chatterji, 1938
  • Macrechinostomum Odhner, 1911
  • Macrocercus Müller, 1773
  • Macrocraeoides
  • Macrogenotrema Skrjabin & Baskakov, 1925
  • Macrurochaeta Costa, 1864
  • Maillardiella Brooks & Holcman, 1993
  • Malleolus Ehrenberg, 1838
  • Manidicola Graber & Gevrey, 1981
  • Maritremopsis Deblock, 1973
  • Maxtrema Travassos, Teixeira de Freitas & Kohn, 1969
  • Mazzantia Travassos & Kohn, 1964
  • Mediantestis Issaitschikow, 1928
  • Mediodera Pigulevsky, 1952
  • Mediopharyngeum Witenberg, 1925
  • Mediorchis Panin, 1984
  • Mediorima Nicoll, 1914
  • Meditypus Barton, 1994
  • Megacetabulum Oshmarin, 1964
  • Megacustis Bennett, 1935
  • Megalatriotrema Rao, 1969
  • Megalophalloides Ching & Ibanez Herrera, 1976
  • Mehraformis Bhardwaj, 1963
  • Melanoglena Eichwald, 1850
  • Mesodendrium Faust, 1919
  • Mesogonimus Monticelli, 1885
  • Metacercaria
  • Metadelphis Travassos, 1944
  • Metagorgoderina Fernandes, 1958
  • Metagyrodactylus Yamaguti, 1963
  • Metahelicometra Yamaguti, 1971
  • Microcotyle Tripathi, 1956
  • Microderma Mehra, 1931
  • Microphallus Deblock & Pearson, 1969
  • Mixophthalmus Karyakarte, 1967
  • Monarrhenos Deblock & Pearson, 1971
  • Monocaecum Deblock & Pearson, 1970
  • Monostoma Rudolphi, 1809
  • Monostoma Zeder, 1800
  • Monostomeus Rafinesque, 1815
  • Monostomulum Brandes, 1892
  • Morenodendrium Odening, 1969
  • Muhlingina Mehra, 1950
  • Multiglandularis Schulz & Skworzow, 1931
  • Multigonadus Bilqees, 1970
  • Multilecithum Oshmarin & Demshin, 1972
  • Musalia Shipley & Hornell, 1904
  • Muttua Shipley & Hornell, 1904
  • Nannoenteron Ozaki, 1924
  • Navigiolum Onji & Nishio, 1924
  • Neodactylodiscus Kamegai, 1972
  • Neodermophtherius Price, 1963
  • Neodicrocoelium Agrawal & Sharma, 1990
  • Neodiplostomum Dubois, 1937
  • Neoeuparadistomum Farooq & Azra Qamar, 1996
  • Neogorgoderina Pereira & Cuocolo, 1940
  • Neogyrodactylus Baugh, 1957
  • Neohamatopeduncularia Gupta & Varsha Sachdeva, 1987
  • Neolepoderma Mehra, 1937
  • Neomicrophallus Farooq & Yousuf, 1986
  • Neophroechinostoma Oshmarin & Belous, 1951
  • Neopisthorchis Chatterji & Kruidenier, 1961
  • Neoprosotocus Bourgat & Combes, 1977
  • Neorenifer Byrd & Denton, 1938
  • Nephrocephala Diesing, 1858
  • Neuronaia Goodsir, 1845
  • Nicollina Deshmukh, 1988
  • Nidarosia
  • Nidrosia Mortensen, 1933
  • Niviventertrema Dronen, Homesley & Cleveland, 1995
  • Notaster Unnithan, 1971
  • Octostoma Otto, 1823
  • Octothilus Hermann, 1782
  • Oligolecithus Vercammen-Grandjean, 1960
  • Olssoniella Travassos, 1944
  • Olveria Sinha, 1941
  • Olveria Thapar & Singha
  • Omnipharynx Nicholls & Brooks, 1969
  • Ompoktrema Varma & Singh, 1986
  • Onchiodiscus Kulkarni, 1969
  • Onchogaster Claparède, 1863
  • Ophiorchis Mehra, 1937
  • Opisthoarchiotrema Gupta & Sharma, 1974
  • Opisthobrachylecithum Yamaguti, 1971
  • Opisthogenes Nicoll, 1914
  • Opistholecithum Baer, 1970
  • Opistocotyle LeBlond, 1836
  • Oreochromogyrus Ferdousi & Chandra, 2002
  • Orientoacanthostomum Zaidi & Khan, 1977
  • Orthorchis Mödlinger, 1925
  • Orthorchis Travassos, 1944
  • Oryzomytrema Thatcher, 1982
  • Oswaldoia Travassos, 1919
  • Overstreetium Brooks & Holcman, 1993
  • Palamphistomum Srivastava & Tripathi, 1987
  • Palitrema Cogate, 1939
  • Pancreaticum Bhalerao, 1936
  • Pancreatrema Oshmarin, 1952
  • Papillatrium Richard, 1966
  • Papilloleucochloridium Bakke, 1980
  • Parabascoides Stunkard, 1938
  • Parabenedenia Johnston, 1929
  • Paracanthatrium Dubois, 1961
  • Paracephalogoniomus Skrjabin, 1949
  • Paraconcinnum Vassiliades & Richard, 1970
  • Paradistomoidella Brooks & Palmieri, 1981
  • Paradistomoides Travassos, 1944
  • Paraibatrema Ueta, Deberaldini, da Silva Cordeiro & de Toledo Artigas, 1981
  • Paraindopleurogenes Kalyankar & Palladwar, 1977
  • Paraleyogonimus Vaucher, 1968
  • Paralutztrema Faust, 1967
  • Parametadelphis Travassos, 1955
  • Paramonostomoides Yamaguti, 1971
  • Paramphistomacotyle Mehra, 1980
  • Paramphistomoides Yamaguti, 1958
  • Paraoistosomum Tkach, 2011
  • Paraplagiorchis Dollfus, 1924
  • Paraschistosomatium Price, 1929
  • Parepibdella Johnston, 1929
  • Paryphostomum Jain, 1952
  • Pentadion Saint-Rémy, 1891
  • Petalodistomum Pigulevsky, 1952
  • Philline Parona & Perugia, 1890
  • Phillines Westwood, 1840
  • Phrynobatrachotrema Goodman, 1986
  • Placotrema Looss, 1907
  • Plagiorchoides Olsen, 1937
  • Platyamphistoma Yamaguti, 1958
  • Platycotyle Beneden & Hesse, 1863
  • Platymetra Mehra, 1931
  • Platynosomoides Yamaguti, 1971
  • Plenosominoides Ke & Liang, 1984
  • Pleurolobatus Kaw, 1944
  • Pleuropsoloides Goodman, 1990
  • Polyporus Grube, 1840
  • Postodera Pigulevsky, 1952
  • Postpharyngeum Witenberg, 1925
  • Praeorchitrema Oshmarin, 1952
  • Primatotrema Premvati, 1958
  • Proacetabulorchoides Oshmarin, 1970
  • Probolitrema Pigulevsky, 1952
  • Prochiorchis Dollfus, 1963
  • Procyonophilus Thatcher, 1982
  • Proechimytrema Thatcher, 1982
  • Proeputiodiplostomum Gupta, 1962
  • Progonimodiscus Vercammen-Grandjean, 1960
  • Promicrophallus Fischthal & Kuntz, 1981
  • Pronopharynx Cohn, 1904
  • Proschistosoma Grétillat, 1962
  • Proshystera Korkhaus, 1930
  • Prosostephanus Lutz, 1935
  • Prosthometra Looss, 1896
  • Prostorchigenes Tubangui, 1928
  • Protofasciola Odhner, 1926
  • Protofasciolopsis Odhner
  • Prouterina Foreyt, Schell & Beyer, 1996
  • Prudhoella Beverley-Burton, 1960
  • Prymnoprion Looss, 1899
  • Pseudalassostoma Yamaguti, 1958
  • Pseudallassostomoides Yamaguti, 1971
  • Pseudallobenedenia Yamaguti, 1966
  • Pseudathesmia Travassos, 1942
  • Pseudoallocreadium Yamaguti, 1971
  • Pseudobianium Sey, 1996
  • Pseudocalceostoma Yamaguti, 1963
  • Pseudodiplodiscoides Murty, 1970
  • Pseudodiscus Sonsino, 1895
  • Pseudoechinostomum
  • Pseudoganeo Khomenovskij, 1971
  • Pseudomagnivitellinum Dronen & Underwood, 1980
  • Pseudomerizocotyle Kay, 1942
  • Pseudomorcheides Wang, 1982
  • Pseudoorientocreadium Agrawal & Sharma, 1990
  • Pseudoparamphistoma Yamaguti, 1958
  • Pseudoplagiorchis Yamaguti, 1971
  • Pseudoquinqueserialis Sutton, 1981
  • Pseudorchis Yamaguti, 1971
  • Pseudorenifer Price, 1935
  • Pseudospelotrematoides Yamaguti, 1939
  • Pterocotyle Beneden & Hesse, 1863
  • Pteronella Beneden & Hesse, 1863
  • Pterygotomaschalos Stunkard, 1924
  • Ptyasiorchis Mehra, 1937
  • Quasimaritrema Deblock, 1973
  • Rallitrema Travassos & Kohn, 1964
  • Redia Filippi, 1837
  • Reduxotrema Pigulevsky, 1952
  • Renicola Sudarikov, 1951
  • Rhacophotrema Uchida, Itagaki & Inoue, 1980
  • Rhodobilharzia Le Roux, 1958
  • Rhopalocerca Diesing, 1850
  • Rudolphiella Travassos, 1924
  • Rutschurutrema Baer, 1959
  • Sarumitrema Beverley-Burton, 1963
  • Sawqirahcotyle Lebedev, 1976
  • Schisturus Rudolphi, 1809
  • Schizocerca Diesing, 1858
  • Scintillovulva Unnithan, 1971
  • Serpentinotrema Travassos & Kohn, 1964
  • Sigmapera Nicoll, 1918
  • 華畢屬 Sinobilharzia Dutt & Srivastava, 1955
  • Sinobilharzia Le Roux, 1958
  • Sinuosoides Denton & Krissinger, 1975
  • Skrjabinodendrium Skarbilovitch, 1943
  • Skrjabinosomum Skrjabin, 1944
  • Solenocerca Diesing, 1855
  • Solenocotyle Diesing, 1850
  • Sorexegliphe Sadovskaya, 1954
  • Spinoderetrema Bray, 1987
  • Spinometra Mehra, 1931
  • Sporocerca Diesing, 1855
  • Sporocystis Filippi, 1854
  • Sporocystophora Diesing, 1858
  • Sporonema Diesing, 1855
  • Sporotherium Diesing, 1858
  • Srivastavaia Singh, 1970
  • Staffordiella Mehra, 1966
  • Stephanolecithus Nakagawa, 1919
  • Stephanopharynx Fischoeder, 1901
  • Stichocotyle Faust & Tang, 1936
  • Strzeleckia Cribb & Spratt, 1991
  • Sturniratrema Velez & Thatcher, 1992
  • Suchocyathocotyle Dubois, 1984
  • Sureshiella Srivastava, Maurya & Prasad, 1980
  • Symmetricatesticula Krasnolobova, 1976
  • Synethes Stiles & Goldberger, 1910
  • Tagumaea Fukui, 1930
  • Taiwantrema Fischthal & Kuntz, 1981
  • Telogonella Mehra & Negi, 1928
  • Tenuifasciola Yamaguti, 1971
  • Testisaculus Bhalerao, 1927
  • Tetrapapillatrema Ralph, 1938
  • Tetrastoma Chiaje, 1833
  • Thailandobilharzia Baugh, 1977
  • Thylogaster Stiles & Goldberger, 1910
  • Topsiturvitrema Velez & Thatcher, 1992
  • Travassasella Faust & Tang, 1938
  • Travassodendrium Skarbilovitch, 1943
  • Travassosina Vaz, 1932
  • Travassostrema Rebecq, 1962
  • Triloborchidiplostomum Dubois & Angel, 1972
  • Trionchus MacCallum, 1916
  • Tumaclinostomum Van der Kuyp, 1953
  • Turbinilla Bory de St.Vincent, 1823
  • Turquinia Coy Otero, 1989
  • Ugandalubens Goodman, 1988
  • Ultragenotrema Skrjabin & Baskakow, 1925
  • Unistriata Filimonova, 1985
  • Vaginaespina Tripathi, 1956
  • Velasquezotrema Eduardo & Javellana, 1987
  • Verodunia Lahille & Joan, 1918
  • Vesperugidendrium Pande, 1937
  • Virgulina Bory de St.-Vincent, 1823
  • Vitellariotrema Gupta & Krishna, 1977
  • Voelkeria Travassos & Kohn, 1966
  • Watsonius Stiles & Goldberger, 1910
  • Wellmanius Stiles & Goldberger, 1910
  • Wrightrenicola Odening, 1962
  • Xenopharynx Nicoll, 1912
  • Xenopodistomum Macnae, Rock & Makowski, 1973
  • Xiphidiocercaria Diesing, 1855
  • Zeferinella Artigas & Paulino, 1988
  • Zeylanurotrema Crusz & Sanmugasunderam, 1974
  • Zoosperma Bory de St Vincent, 1823
  • Zygodendrium Salem, 1976
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構造

體壁組織可分為體被與肌肉;可自我更新;具有吸收營養、感覺等功能[6]

參考文獻

參見

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