卷緣樁菇

卷緣樁菇（學名：Paxillus involutus）又稱卷邊網褶菌或卷邊樁菇，最早於1785年由皮埃爾·比利亞爾描述發表，當時被歸在傘菌屬^[3]，但翌年由奧古斯特·巴奇發表的學名Agaricus involutus才被視作有效描述^[1][4]；1838年埃利亞斯·芒努斯·弗里斯定立新屬樁菇屬，並將本種作為模式種^[5]。本種外型如棕色、木質的陀螺，子實體可高約6公分^[6]，蕈傘直徑為4—12公分，邊緣向下捲曲^[7]；菌髓黃色且多汁，並具特殊氣味^[7][8]；孢子印為棕色，擔孢子呈橢球形，長7.5—9微米，寬5—6微米^[9]。許多乳菇屬物種、黑毛樁菇與耳狀網褶菌（英語：Tapinella panuoides）等蕈類的外形皆與本種相似^[10][11]，另外同屬物種Paxillus obscurisporus（英語：Paxillus obscurisporus）為自本種分出，其子實體較本種為大^[12]。

卷緣樁菇
科學分類
界：	真菌界 Fungi
門：	擔子菌門 Basidiomycota
綱：	傘菌綱 Agaricomycetes
目：	牛肝菌目 Boletales
科：	網褶菌科 Paxillaceae
屬：	網褶菌屬 Paxillus
種：	卷緣樁菇 P. involutus
二名法
Paxillus involutus (Batsch) Fr. (1838)
異名[1][2]
Agaricus contiguus Bull. (1785) Agaricus involutus Batsch (1786) Agaricus adscendibus Bolton（英語：James Bolton） (1788) Omphalia involuta (Batsch) Gray (1821) Rhymovis involuta (Batsch) Rabenh.（英語：Gottlob Ludwig Rabenhorst） (1844)

卷緣樁菇
查看產生下列表格的真菌學模板 真菌形態特徵
	子實層上有菌褶
	蕈傘為扁平
	子實層自基部沿蕈柄向下生長
	裸露蕈柄
	孢印褐色
	菌根真菌
	致命

皮埃爾·比利亞爾於1785年繪製的卷緣樁菇

卷緣樁菇可與許多針葉樹或落葉樹形成外菌根（英語：Ectomycorrhiza）^[13]，常在外菌根相關研究中被用作模式生物，其基因組已被完整定序^[14]。有研究顯示本種與樹木共生時可增加樹木根部對土壤真菌鐮刀菌感染的抗性^[15][16]，並抑制植株吸收土壤中的有害重金屬^[17][18]。本種為北半球溫帶地區常見的蕈類，分布範圍包括歐洲、亞洲與北美洲^[10][19]，在印度^[20]、伊朗^[21]、中國^[22]、日本也有分布，最北可至阿拉斯加與格陵蘭^[23][24]，並已作為外來種被引入澳洲^[25][26]、紐西蘭^[27]與南美洲的智利^[28]。二戰以前卷緣樁菇在中歐與東歐都被認為在烹煮過可食^[7][19]，僅知生食後可能引起腸胃症狀^[9]，但1944年德國真菌學家尤利烏斯·舍費爾（英語：Julius Schäffer）食用此菌後中毒身亡^[19][29]，後來研究發現卷緣樁菇的一個抗原可激活自體免疫反應，使人體的免疫系統攻擊自身的紅血球，以致自體免疫性溶血，多發生在長期食用此菌者^[9][30]。1990年代起本種逐漸被列為不可食用的蕈類^[31]，但東歐每年仍每年有人食用此菌後中毒而死^{[32][33][34][35]}。

參考文獻

• 真菌主題

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24. Knudson, Henning. Arctic and Alpine Mycology 6. Copenhagen, Denmark: Museum Tusculanum Press. 2006: 12. ISBN 87-635-1277-7.
25. Cleland, John Burton. Toadstools and Mushrooms and Other Larger Fungi of South Australia. Adelaide, Australia: South Australian Government Printer. 1976: 177.
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