P2A、E2A、F2A、T2A。其中F2A源於手足口病毒(英语:foot-and-mouthdiseasevirus)(Foot-and-mouthdiseasevirus)、E2A源於馬甲型鼻炎病毒(Equine rhinitis A virus)、P2A源於豬捷申病毒(英语:Porcine teschovirus)(Porcine
Clercq K. 2'-C-methylcytidine as a potent and selective inhibitor of the replication of foot-and-mouthdiseasevirus. Antiviral Research. March 2007, 73 (3):