University of Copenhagen Zoological Museum
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1852年,他辭去了丹麥陸軍中尉的職務,並於次年獲得科學碩士學位。之後,他在哥本哈根大學動物博物館(英语:University of Copenhagen Zoological Museum)擔任喬珀托斯·史汀史翠普的助手。史汀史翠普於1885年退休後,他成為動物學教授和動物博物館館長。隨著年齡的
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forces of annelids and chilopods and its bearing upon the evolution of the arthropodan haemocoel. Zoological Journal of the Linnean Society(英语:Zoological Journal
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and the late Miocene Dinocrocuta gigantea (Carnivora, Mammalia). Zoological Journal of the Linnean Society. March 2010, 158 (3): 683–696. doi:10.1111/j
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Evolving Eden. An Illustrated Guide to the Evolution of the African Large-Mammal Fauna. Columbia University Press, New York, 2004. ISBN 0-231-11944-5 Jordi
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2004, 154: 8 [2009-12-11]. ISSN 0044-5088. (原始内容存档于2009-06-26). Scharff, N & Enghoff, H. Arachnida. Zoological Museum, University of Copenhagen. 2005.