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, and Larsson, H.C.E. Kemp, Tom , 编. Evidence for Avian Intrathoracic Air Sacs in a New Predatory Dinosaur from Argentina. PLoS ONE. 2008, 3 (9): e3303
Barrett, Paul M.; Gower, David J. Postcranial skeletal pneumaticity and air-sacs in the earliest pterosaurs. Biology Letters. 2009-08-23, 5 (4) [2022-07-03]
4公尺，三指的手上长有大而钝的爪子。腿相对较短、脚爪较钝，其脊椎带有很高的神经棘，沿背部形成一个“帆”。大多数椎骨和部分其它骨骼被嵌入的气腔（英语：air sacs）所填充（英语：Skeletal pneumaticity）。尾巴末端有一节类似尾综骨的椎骨，表明此处存在一个由羽毛构成的扇状物。
恐龙生理学
pneumaticity, air sacs, and the physiology of sauropod dinosaurs. [2008-02-27]. （原始内容存档于2011-04-11）. and Vertebral pneumaticity, air sacs, and the physiology
Barrett, Paul M.; Gower, David J. Postcranial skeletal pneumaticity and air-sacs in the earliest pterosaurs. Biology Letters. 2009-08-23, 5 (4) [2022-08-07]