坚齿螺科(学名:Camaenidae),又名南亚蜗牛科[4][5][6][7],是旋蜗牛总科(Helicoidea)之下的一个科,全为陆生[2]会呼吸空气的有肺类腹足纲软体动物。一般常见的蜗牛大都属于本科,而本科亦是柄眼类支序中物种多样性最高的一科。
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本科物种的栖息地都在东亚及澳大拉西亚的热带地区[8]。旧有文献认为肋齿螺科(Pleurodontidae)是本科的异名[4],但现时分子系统发生学已认为这两个科实属不同物种,而且地域分布亦不相同。
各科物种广泛分布于印度洋及西太平洋,包括下列各个地区:
以记录海洋物种为主的WoRMS在本科纪录有超过100个属[2];而在GBIF更是有198个属、590个物种[9]。然而本科物种的分类近三十年来的变化也颇大。
截至2018年6月5日 (2018-06-05)[update],WoRMS纪录本科包括下列各亚科[2][10]:
- 扁蜗牛亚科 Bradybaeninae Pilsbry, 1934 (1898)[11]:旧属扁蜗牛科。
- 盾蜗牛族 Aegistini Kuroda & Habe, 1949
- 扁蜗牛族 Bradybaenini Pilsbry, 1934 (1898):异名有 Eulotidae Möllendorff, 1898; Fruticicolinae Kobelt, 1904; Buliminopsinae Hoffmann, 1928
- 真厚螺族 Euhadrini Habe, Okutani & Nishiwaki, 1994:只有真厚螺属(Euhadra Pilsbry, 1890)一个属[12]。
- 坚齿螺亚科 Camaeninae Pilsbry, 1895[13]
- 厚螺亚科 Hadrinae Iredale, 1937[14]:包括有160个属。
- 光泽蜗牛亚科 Helicostylinae Ihering, 1909[15]:旧属扁蜗牛科,有23个属。异名有 Pfeifferiinae Gray, 1855; Cochlostylidae Möllendorff, 1890
- 亚科地位未定
- 两坑螺属 Amphicoelina Zilch, 1960
- † Archaeoxesta Kobelt in Neuenhaus, 1909
- 榼子螺属 Entadella Páll-Gergely & Hunyadi, 2016
- † Hodopoeus Pilsbry & Cockerell, 1945
- † Kanabohelix Pilsbry, 1927
- Kenyirus Clements & Tan, 2012
- Ogeramua Christensen, 2017
2017年,原巴蜗牛科被合并进本科去,原来巴蜗牛科之下的两个亚科成为了本科的亚科,令本科之下的分类单元超过400个[2][16]。现时本科包括下列各个属[17][18]:
- 盾蜗牛族 Aegistini Kuroda & Habe, 1949
- 盾蜗牛属 Aegista :又名大脐蜗牛属 ,亲缘关系上为多系群,其中亦包含Plectotropis。[19]
- 长蜗牛属 Dolicheulota Pilsbry, 1901
- 岚都螺属 Landouria Godwin-Austen, 1918
- 鞭巴蜗牛属 Mastigeulota Pilsbry, 1895
- 伪盾蜗牛属 Pseudaspasita Möllendorff, 1901
- 岛蜗牛属 Nesiohelix Kuroda & Emura, 1943
- 环肋螺属 Plectotropis
- 萨摩蜗牛属 Satsuma A. Adams, 1868[6][7][20],菱蜗牛属
- 高砂厚螺属 Takasagohadra Kuroda, 1941
- 林氏高砂厚螺 Takasagohadra lini S.-P. Wu & C.-C. Hwang, 2013
- Takasagohadra multifasciata Kuroda, 1941
- 巴蜗牛族 Bradybaenini Pilsbry, 1934 (1898):有30个属[21]。
- 尖巴蜗牛属 Acusta E. von Martens, 1860
- 阿玛螺属 Armandiella Ancey, 1901
- 巴蜗牛属 Bradybaena Pilsbry, 1934 (1898)
- 华蜗牛属 Cathaica Möllendorff, 1884
- 朝鲜蜗牛属 Chosenelix Pilsbry, 1927
- 粒雕螺属 Coccoglypta Pilsbry, 1895
- 宁馨螺属 Eueuhadra M. Wu, 2004
- 灌丛蜗牛属 Fruticicola Held, 1838
- Fruticoconus Möllendorff, 1882
- 小索螺属 Funiculus Heude, 1888
- 葛氏螺属 Grabauia Yen, 1935
- 胶辽螺属 Jiaoliaous Zhang, 2024,胶辽蜗牛属[22]。
- 库页蜗牛属 Karaftohelix Pilsbry, 1927
- 射带蜗牛属 Laeocathaica Möllendorff, 1899
- 间齿螺属 Metodontia Möllendorff, 1886
- 平瓣蜗牛属 Platypetasus Pilsbry, 1895[23]
- 上华蜗牛属 Pliocathaica Andreae, 1900
- 桥齿蜗牛属 Ponsadenia Schileyko, 1978,桥齿螺属
- 拟蛇蜗牛属 Pseudiberus Ancey, 1887
- 假拟锥螺属 Pseudobuliminus Gredler, 1886
- 索状螺属 Rudens Heude, 1890
- Rudens rudens (Heude, 1888)
- 宜昌螺属 Secusana Gredler, 1894
- Secusana cerasina (Gredler, 1892)
- †Secusana ravia W. Yü, 1982
- Secusana superba (Möllendorff, 1888)
- Semibuliminus Möllendorff, 1899
- 华砾螺属 Sinochloritis M. Wu & Z.-Y. Chen, 2019
- 李氏华砾螺 Sinochloritis lii M. Wu & Z.-Y. Chen, 2019
- 华脊纳螺属 Sinorachis M. Wu & Z.-Y. Chen, 2019
- Sinorachis aureus (Heude, 1890)
- Sinorachis baihu M. Wu & Z.-Y. Chen, 2019
- Sinorachis onychinus (Heude, 1885)
- 拟窄圆螺属 Stenogyropsis Möllendorff, 1899
- Stenogyropsis chorismenostoma Z.-Y. Chen, B. Huang & Páll-Gergely, 2022
- Stenogyropsis cocoa Páll-Gergely & Hunyadi, 2017
- Stenogyropsis potanini (Möllendorff, 1899)
- 亮盘螺属 Stilpnodiscus Möllendorff, 1900,光盘螺属
- 毛巴蜗牛属 Trichobradybaena M. Wu & J.-Y. Guo, 2003
- Trichobradybaena chagangensis M. Wu & J.-Y. Guo, 2003
- Trichobradybaena submissa (Deshayes, 1874)
- Trichobradybaena tuberculata M. Wu & J.-Y. Guo, 2003
- 毛华蜗牛属 Trichocathaica Gude, 1919
- 室女螺属 Virginihelix Kuroda & Habe, 1949
- Virginihelix arcasiana (Crosse & Debeaux, 1863) ,接受名为:Karaftohelix arcasiana (Crosse & Debeaux, 1863)
- Virginihelix virgo (Pilsbry, 1927) ,接受名为:Bradybaena virgo (Pilsbry, 1927)
- 坚螺亚科 Camaeninae Pilsbry, 1895
- 缅砾螺属 Burmochloritis Godwin-Austen, 1920
- 小坚螺属 Camaenalla
- 坚螺属 Camaena Albers, 1850
- Caracolus Montfort, 1810
- Chloritis H.Beck, 1837
- 小丽螺属 Ganesella W. T. Blanford, 1863[6]
- 皱口蜗牛属 Moellendorffia Ancey, 1887[6]:又名多粒螺属。
- 小多粒螺属 Moellendorffiella Pilsbry, 1905
- Moellendorffiella erdmanni (Schmacker & O.Boettger, 1894)[24],见于重庆武隆县
- Moellendorffiella faberiana (Möllendorff, 1888)
- Moellendorffiella yifengi [24],见于云南文山开化
- 新洋葱螺属 Neocepolis Pilsbry, 1891
- 拟盖螺属 Pseudostegodera M. Wu & Z.-Y. Chen, 2021
- Pseudostegodera qiului Z.-Y. Chen, 2021
- Rhagada (Cox, 1870)
- 中华坚螺属 Sinocamaena M. Wu, 2024
- 陈氏中华坚螺 Sinocamaena cheni M. Wu, 2024
- 盖螺属 Stegodera E. von Martens, 1876
- 伤口螺属 Traumatophora Ancey, 1887
- 绒蜗牛属 Trichelix Ancey, 1887
- Trichelix biscalpta (Heude, 1885)
- Trichelix diminuta (Pilsbry & Y. Hirase, 1905)
- Trichelix eucharista (Pilsbry, 1901)
- Trichelix hiraseana (Pilsbry, 1905)
- Trichelix horrida (L. Pfeiffer, 1863)
- Trichelix tokunoensis (Pilsbry & Y. Hirase, 1905)
- Trichelix xiaoxiang R.-X. Lin & L.-W. Lin, 2022
- 毛蜗牛属 Trichochloritis Pilsbry, 1891,毛砾螺属[6]
- 白口绒蜗牛属 Yakuchloritis Habe, 1955。屋久砾螺属
- 光泽蜗牛亚科 Helicostylinae Ihering, 1909
- 光泽蜗牛属 Helicostyla Férussac, 1821,尖旋螺属
- 其他:
- Figuladra Köhler & Bouchet, 2020
- Figuladra appendiculata (Reeve, 1854)
- Figuladra barneyae J.Stanisic, 2010
- Figuladra bayensis (Brazier, 1876)
- Figuladra challisi (J.C.Cox, 1873)
- Figuladra incei (L.Pfeiffer, 1846)
- Figuladra muirorum J.Stanisic, 2010
- Figuladra narelleae J.Stanisic & Potter, 2010
- Figuladra pallida (Hedley & Musson, 1892)
- Figuladra reducta (Iredale, 1937)
- Figuladra robertirwini L. Stanisic, J. Stanisic & Köhler, 2024[25]
- Figuladra vidulus L. Stanisic, J. Stanisic & Köhler, 2024[25]
Köhler, F. Three new species and two new genera of land snails from the Bonaparte Archipelago in the Kimberley, Western Australia (Pulmonata, Camaenidae). Molluscan Research. 2010, 30 (1): 1–16.
林朝棨. 蘭嶼之貝類及其動物地理. 中国贝志 (中华民国贝类学会). 1974, 1: 42–63 [2018-10-10] (中文(繁体)).
Cuezzo, M. G. Phylogenetic analysis of the Camaenidae (Mollusca: Stylommatophora) with special emphasis on the American taxa. Zoological Journal of the Linnean Society. 2003, 138 (4): 449–476. doi:10.1046/j.1096-3642.2003.00061.x (英语).
Camaenidae. GBIF. Copenhagen, Denmark: GBIF Secretariat. [2017-02-15].
Bouchet, P.; Rocroi, J.-P.; Hausdorf, B.; Kaim, A.; Kano, Y.; Nützel, A.; Parkhaev, P.; Schrödl, M.; Strong, E.E. Revised Classification, Nomenclator and Typification of Gastropod and Monoplacophoran Families. Malacologia. 2017, 61 (1–2): 1–526. OCLC 7296100651. S2CID 91051256. doi:10.4002/040.061.0201.
Bieler R, Bouchet P, Gofas S, Marshall B, Rosenberg G, La Perna R, Neubauer TA, Sartori AF, Schneider S, Vos C, ter Poorten JJ, Taylor J, Dijkstra H, Finn J, Bank R, Neubert E, Moretzsohn F, Faber M, Houart R, Picton B, Garcia-Alvarez O (编). Euhadrini Habe, Okutani & Nishiwaki, 1994. MolluscaBase. 2018-02-06 [2018-07-16] –通过WoRMS.
Bieler R, Bouchet P, Gofas S, Marshall B, Rosenberg G, La Perna R, Neubauer TA, Sartori AF, Schneider S, Vos C, ter Poorten JJ, Taylor J, Dijkstra H, Finn J, Bank R, Neubert E, Moretzsohn F, Faber M, Houart R, Picton B, Garcia-Alvarez O (编). Bradybaenidae Pilsbry, 1934 (1898). MolluscaBase. 2017-07-13 [2018-05-13] –通过WoRMS.
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Bank R. Bieler R, Bouchet P, Gofas S, Marshall B, Rosenberg G, La Perna R, Neubauer TA, Sartori AF, Schneider S, Vos C, ter Poorten JJ, Taylor J, Dijkstra H, Finn J, Bank R, Neubert E, Moretzsohn F, Faber M, Houart R, Picton B, Garcia-Alvarez O , 编. Bradybaenini Pilsbry, 1934 (1898). MolluscaBase. 2017-06-13 [2018-06-05] –通过WoRMS.
Zhang, Guoyi; Naggs, Fred; Peter, Andrus; Christopher M, Wade. Phylogenetic insights into the terrestrial snails Helicoidei (Gastropoda: Stylommatophora) with special emphasis on the Camaenidae. Zoological Journal of the Linnean Society. 2024. doi:10.1093/zoolinnean/zlae027.
Zhang, Guoyi; Liu, Zhongshu; Qi, Feng; Qaunyu, Zhang. Systematics and biogeography of the genera Pseudiberus Ancey, 1887 and Platypetasus Pilsbry, 1895 (Stylommatophora: Camaenidae), with the description of Pseudiberus shanheicus n. sp.. Journal of Conchology. 2024. doi:10.61733/jconch/4510.
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Lorelle Stanisic, John Stanisic and Frank Köhler. 2024. Systematic Revision of the eastern Queensland Land Snail Genus Figuladra Köhler and Bouchet, 2020 based on mitochondrial phylogenetics and comparative morpho-anatomy (Eupulmonata, Camaenidae). Molluscan Research. 44(2); 160-208. DOI: 10.1080/13235818.2024.2321650
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