Ochrophyte

Ochrophyte
	Gusta šuma morskih algi sa podlogom
Naučna klasifikacija
Domen:	Eukaryota
Kladus:	Diaphoretickes
Kladus:	SAR
Infracarstvo:	Heterokonta
Tip:	Gyrista
Podtip:	Ochrophytina; Cavalier-Smith, 1995
Klase
	Olisthodiscophyceae; Chrysista Limnistia Chrysophyceae; Eustigmatophyceae; Synchromophyceae; ; Raphidoistia Raphidomonadea; ; Fucistia Aurophyceae; Chrysomerophyceae; Chrysoparadoxophyceae; Phaeosacciophyceae; Phaeophyceae; Schizocladiophyceae; Xanthophyceae; ; ; Diatomista Hypogyrista Dictyochophyceae; Pinguiophyceae; ; Khakista Bolidophyceae; Bacillariophyceae; ; ;
Sinonimi
	Heterokontophyta van den Hoek et al., 1978; Ochrista Cavalier-Smith, 1986; Stramenochromes Leipe et al., 1994;

Ochrophytes su grupa uglavnom fotosintetskih stramenopila (heterokonta), smeštenih ili u razdeo Ochrophyta ili podrazdeo Ochrophytina.^[5]^[6]^[7] Njihov plastid je poreklom od crvenih algi.^[8]^[7]

Кратке чињенице Naučna klasifikacija, Klase ...

Затвори

Klasifikacija grupe se još razvija. Prvobitno, okrofiti su smatrani razdelom koji se naziva Ochrophyta. Neki autori su ga podelili na dva podrayreda, Phaeista Cavalier-Smith 1995 (koji obuhvata Hypogyristea i Chrysista u nekim klasifikacijama, ili Limnista i Marista u drugima) i Khakista Cavalier-Smith, 2000 (sadrži Bolidophyceae i silikatne alge).^[9]

Kladogram ispod prikazuje evolutivne odnose između svih klasa okrofita. Zasnovan je na najnovijim filogenetskim analizama:^[10]^[7]^[11]^[12]

Ochrophytina

Chrysista

"SI klada"

Fucistia

	Xanthophyceae – žutozelene alge Allorge ex Fritsch 1935

	Chrysoparadoxophyceae Wetherbee et al. 2019

Phaeosacciophyceae
R.A.Andersen, L.Graf & H.S.Yoon 2020

	Schizocladiophyceae Henry, Okuda & Kawai 2003

	Phaeophyceae – smeđe alge Kjellman 1891

Aurophyceae
Cavalier-Smith 2013

"PX klada"

Raphidoistia

Raphidomonadea
Chadefaud ex Silva 1980

"SII klada"

Olisthodiscophyceae
Barcytė, Eikrem & M. Eliáš 2021

Limnistia

	Chrysophyceae – zlatne alge Pascher 1914

	Synchromophyceae Horn et al. 2007

Eustigmatophyceae
Hibberd & Leedale 1971

Diatomista

Hypogyrista

	Dictyochophyceae Silva 1980

	Pinguiophyceae Kawachi et al. 2002

Khakista

	Bolidophyceae Guillou & Chretiennot-Dinet 1999

	Bacillariophyceae – Silikatne alge Dangeard 1933

"SIII klada"

Verovatna sestrinska klada Ochrophyta/Ochrophytina je Actinophryidae,^[13] prvobitno klasifikovana kao heliozoanski protisti.^[14]

[1]
Cavalier-Smith T (1995). „Zooflagellate phylogeny and classification”. Cytology. 37 (11): 1010—1029. PMID 8868448.
[2]
Cavalier-Smith, T. (1986). The kingdom Chromista, origin and systematics. In: Round, F.E. and Chapman, D.J. (eds.). Progress in Phycological Research. 4: 309–347.
[3]
Reviers, B. de. (2006). Biologia e Filogenia das Algas. Editora Artmed, Porto Alegre, p. 157.
[4]
Reviers, B. de. (2006). Biologia e Filogenia das Algas. Editora Artmed, Porto Alegre, pp. 15-16.
[5]
Thomas Cavalier-Smith; Ema E.-Y. Chao (2006). „Phylogeny and megasystematics of phagotrophic heterokonts (kingdom Chromista)”. Journal of Molecular Evolution. 62 (4): 388—420. Bibcode:2006JMolE..62..388C. PMID 16557340. S2CID 29567514. doi:10.1007/s00239-004-0353-8.
[6]
Ingvild Riisberga; Russell J. S. Orr; Ragnhild Kluge; Kamran Shalchian-Tabrizi; Holly A. Bowers; Vishwanath Patil; Bente Edvardsen; Kjetill S. Jakobsen (2009). „Seven gene phylogeny of heterokonts”. Protist. 160 (2): 191—204. PMID 19213601. doi:10.1016/j.protis.2008.11.004.
[7]
Cavalier-Smith, Thomas (2018). „Kingdom Chromista and its eight phyla: a new synthesis emphasising periplastid protein targeting, cytoskeletal and periplastid evolution, and ancient divergences”. Protoplasma. 255 (1): 297—357. PMC 5756292 . PMID 28875267. doi:10.1007/s00709-017-1147-3 .
[8]
Ševčíková T, Horák A, Klimeš V, . (2015). „Updating algal evolutionary relationships through plastid genome sequencing: did alveolate plastids emerge through endosymbiosis of an ochrophyte?”. Sci Rep. 5: 10134. Bibcode:2015NatSR...510134S. PMC 4603697 . PMID 26017773. doi:10.1038/srep10134 .
[9]
„Heterokontophyta”. SHIGEN. National Institute of Genetics. Приступљено 18. 6. 2009.
[10]
Derelle R, López-García P, Timpano H, Moreira D (новембар 2016). „A Phylogenomic Framework to Study the Diversity and Evolution of Stramenopiles (=Heterokonts)”. Mol Biol Evol. 33 (11): 2890—2898. PMC 5482393 . PMID 27512113. doi:10.1093/molbev/msw168 .
[11]
Graf L, Yang EC, Han KY, Küpper FC, Benes KM, Oyadomari JK, Herbert RJH, Verbruggen H, Wetherbee R, Andersen RA, Yoon HS (децембар 2020). „Multigene Phylogeny, Morphological Observation and Re-examination of the Literature Lead to the Description of the Phaeosacciophyceae Classis Nova and Four New Species of the Heterokontophyta SI Clade”. Protist. 171 (6): 125781. PMID 33278705. S2CID 227315556. doi:10.1016/j.protis.2020.125781 .
[12]
Dovilė Barcytė; Wenche Eikrem; Anette Engesmo; Sergio Seoane; Jens Wohlmann; Aleš Horák; Tatiana Yurchenko; Marek Eliáš (2. 3. 2021). „Olisthodiscus represents a new class of Ochrophyta”. Journal of Phycology. 57 (4): 1094—1118. PMID 33655496. doi:10.1111/jpy.13155 . hdl:10852/86515 .
[13]
Azuma, Tomonori; Pánek, Tomáš; Tice, Alexander K.; Kayama, Motoki; Kobayashi, Mayumi; Miyashita, Hideaki; Suzaki, Toshinobu; Yabuki, Akinori; Brown, Matthew W.; Kamikawa, Ryoma (10. 4. 2022). „An Enigmatic Stramenopile Sheds Light on Early Evolution in Ochrophyta Plastid Organellogenesis”. Molecular Biology and Evolution. 39 (4). PMC 9004409 . PMID 35348760. doi:10.1093/molbev/msac065.
[14]
Mikrjukov, Kirill A.; Patterson, David J. (2001). „Taxonomy and phylogeny of Heliozoa. III. Actinophryids” (PDF). Acta Protozoologica. 40: 3—25.

[1] [1]
Cavalier-Smith T (1995). „Zooflagellate phylogeny and classification”. Cytology. 37 (11): 1010—1029. PMID 8868448.

[2] [2]
Cavalier-Smith, T. (1986). The kingdom Chromista, origin and systematics. In: Round, F.E. and Chapman, D.J. (eds.). Progress in Phycological Research. 4: 309–347.

[3] [3]
Reviers, B. de. (2006). Biologia e Filogenia das Algas. Editora Artmed, Porto Alegre, p. 157.

[4] [4]
Reviers, B. de. (2006). Biologia e Filogenia das Algas. Editora Artmed, Porto Alegre, pp. 15-16.

[5] [5]
Thomas Cavalier-Smith; Ema E.-Y. Chao (2006). „Phylogeny and megasystematics of phagotrophic heterokonts (kingdom Chromista)”. Journal of Molecular Evolution. 62 (4): 388—420. Bibcode:2006JMolE..62..388C. PMID 16557340. S2CID 29567514. doi:10.1007/s00239-004-0353-8.

[6] [6]
Ingvild Riisberga; Russell J. S. Orr; Ragnhild Kluge; Kamran Shalchian-Tabrizi; Holly A. Bowers; Vishwanath Patil; Bente Edvardsen; Kjetill S. Jakobsen (2009). „Seven gene phylogeny of heterokonts”. Protist. 160 (2): 191—204. PMID 19213601. doi:10.1016/j.protis.2008.11.004.

[8phyla-7] [7]
Cavalier-Smith, Thomas (2018). „Kingdom Chromista and its eight phyla: a new synthesis emphasising periplastid protein targeting, cytoskeletal and periplastid evolution, and ancient divergences”. Protoplasma. 255 (1): 297—357. PMC 5756292 . PMID 28875267. doi:10.1007/s00709-017-1147-3 .

[8] [8]
Ševčíková T, Horák A, Klimeš V, . (2015). „Updating algal evolutionary relationships through plastid genome sequencing: did alveolate plastids emerge through endosymbiosis of an ochrophyte?”. Sci Rep. 5: 10134. Bibcode:2015NatSR...510134S. PMC 4603697 . PMID 26017773. doi:10.1038/srep10134 .

[9] [9]
„Heterokontophyta”. SHIGEN. National Institute of Genetics. Приступљено 18. 6. 2009.

[StramenoFramework-10] [10]
Derelle R, López-García P, Timpano H, Moreira D (новембар 2016). „A Phylogenomic Framework to Study the Diversity and Evolution of Stramenopiles (=Heterokonts)”. Mol Biol Evol. 33 (11): 2890—2898. PMC 5482393 . PMID 27512113. doi:10.1093/molbev/msw168 .

[Phaeosacciophyceae-11] [11]
Graf L, Yang EC, Han KY, Küpper FC, Benes KM, Oyadomari JK, Herbert RJH, Verbruggen H, Wetherbee R, Andersen RA, Yoon HS (децембар 2020). „Multigene Phylogeny, Morphological Observation and Re-examination of the Literature Lead to the Description of the Phaeosacciophyceae Classis Nova and Four New Species of the Heterokontophyta SI Clade”. Protist. 171 (6): 125781. PMID 33278705. S2CID 227315556. doi:10.1016/j.protis.2020.125781 .

[Olisthodiscophyceae-12] [12]
Dovilė Barcytė; Wenche Eikrem; Anette Engesmo; Sergio Seoane; Jens Wohlmann; Aleš Horák; Tatiana Yurchenko; Marek Eliáš (2. 3. 2021). „Olisthodiscus represents a new class of Ochrophyta”. Journal of Phycology. 57 (4): 1094—1118. PMID 33655496. doi:10.1111/jpy.13155 . hdl:10852/86515 .

[13] [13]
Azuma, Tomonori; Pánek, Tomáš; Tice, Alexander K.; Kayama, Motoki; Kobayashi, Mayumi; Miyashita, Hideaki; Suzaki, Toshinobu; Yabuki, Akinori; Brown, Matthew W.; Kamikawa, Ryoma (10. 4. 2022). „An Enigmatic Stramenopile Sheds Light on Early Evolution in Ochrophyta Plastid Organellogenesis”. Molecular Biology and Evolution. 39 (4). PMC 9004409 . PMID 35348760. doi:10.1093/molbev/msac065.

[Heliozoa_III-14] [14]
Mikrjukov, Kirill A.; Patterson, David J. (2001). „Taxonomy and phylogeny of Heliozoa. III. Actinophryids” (PDF). Acta Protozoologica. 40: 3—25.

[5]

[6]

[7]

[8]

[1]

[2]

[3]

[4]

[9]

[10]

[11]

[12]

[13]

[14]

Ochrophyte

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Filogenija

Reference

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