Wallace DC (2000). „mtDNA variation in the South African Kung and Khwe-and their genetic relationships to other African populations”. Am. J. Hum. Genet. 66
Haplogroup R-M269, cunoscut și sub numele de R1b1a1a2, este o sub-cladă a haplogrupului R1b al cromozomului uman uman . Este de interes deosebit pentru
Tyler-Smith C (iunie 2019). „A Rare Deep-Rooting D0 African Y-chromosomal Haplogroup and its Implications for the Expansion of Modern Humans Out of Africa”
Palanichamy et al. (2015), "The presence of mtDNA haplogroups (HV14 and U1a) and Y-chromosome haplogroup (L1) in Dravidian populations indicates the spread