^ Hong, Won Shic. 1987. "The Distribution of Western North American Hepaticae. Endemic Taxa and Taxa with a North Pacific Arc Distribution". The Bryologist
絶滅。石炭紀の地層から発見されている。 ^ Schuster, R. M. (1984). “Diagnoses of some new taxa of Hepaticae”. Phytologia 56: 65–74. ^ Crandall-Stotler, Barbara; Stotler, Raymond