Les Scarabaeidae , en français Scarabéidés , sont une famille d'insectes coléoptères de la super-famille des Scarabaeoidea . Ils sont connus sous le nom vernaculaire de Scarabée . C'est notamment la famille des dynastes , du scarabée rhinocéros européen , des bousiers et des hannetons .
Faits en bref Règne, Sous-règne ...
Accouplement de Neorrhina punctatum .
Famille
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Les limites de cette famille sont encore débattues et controversées. Les chercheurs européens en ont une définition plutôt restrictive, alors qu'un certain nombre de chercheurs américains y incorporent bien plus de taxons[1] .
Les antennes des insectes de cette famille sont en massue dont l'extrémité peut s'ouvrir en un éventail de feuillets. L'extrémité de l'abdomen est découverte en raison d'élytres normalement tronqués. La plupart des espèces volent correctement. Plusieurs de ces espèces stridulent en frottant l'extrémité de leurs élytres contre la face dorsale de leur abdomen. Cette vaste famille comprend des coprophages et des saproxylophages nuisibles aux végétaux[2] .
Le terme de Scarabée provient, via le vieux français « escarbot », du latin scarabaeus , par altération du grec ancien κάραβος / kárabos , « escarbot, scarabée, insecte »[3] , également à l'origine du terme carabe [4] . Le terme « scarabée» désigne de façon générale en zoologie l'ensemble de la super-famille des Scarabéoïdes , mais ce terme peut désigner chez certains auteurs, d'une façon plus restrictive, la famille des Scarabaeidae, et même pour certains uniquement les coprophages de cette famille[ réf. nécessaire] .
Plusieurs groupes, qui peuvent être paraphylétiques , de ces insectes ont des noms vernaculaires tels que les Dynastes ou scarabées rhinocéros , les bousiers , les hannetons ... Certaines espèces ont un nom vernaculaire particulier comme le Pique-prune (Osmoderma eremita ).
La famille a été décrite par l'entomologiste français Pierre André Latreille en 1802[5] .
Liste des sous-familles
Selon le Catalogue of Life (23 juin 2023) [6] , cette famille se divise en 28 sous-familles :
Aclopinae Blanchard, 1850
Aegialiinae Castelnau, 1840
Allidiostomatinae Arrow, 1940
Aphodiinae Leach, 1815
Aulonocneminae Janssens, 1946
Cetoniinae Leach, 1815
Chironinae Blanchard, 1845
† Cretoscarabaeinae Nikolajev, 1995
Dynamopodinae Arrow, 1911
Dynastinae MacLeay, 1819
Eremazinae Stebnicka, 1977
Hopliinae Latreille, 1829
Lichniinae Burmeister, 1844
† Lithoscarabaeinae Nikolajev, 1992
Melolonthinae MacLeay, 1819
Oncerinae LeConte, 1861
Orphninae Erichson, 1847
Pachydeminae Reitter, 1902
Phaenomeridinae Erichson, 1847
Podolasiinae Howden, 1997
† Prototroginae Nikolajev, 2000
Rutelinae MacLeay, 1819
Scarabaeinae Latreille, 1802
Sericinae Kirby, 1837
Sericoidinae Erichson, 1847
Termitotroginae Wasmann, 1918
Trichiinae Kolbe, 1897
Valginae Schenkling, 1922
Liste des genres
Selon le Catalogue of Life (23 juin 2023) [6] , cette famille regroupe 2 590 genres :
Ablabera Dejean, 1833
Ablaberoides Blanchard, 1850
Acanthaphodius Schmidt, 1909
Acanthobodilus Dellacasa, 1983
Acanthonitis Janssens, 1937
Acanthovalgus Kraatz, 1895
Aceratus Prell, 1936
Acheilo Britton, 1988
Achelyna Erichson, 1848
Achloa Erichson, 1840
Achranoxia Kraatz, 1888
Aclopus Erichson, 1835
Acoma Casey, 1889
Acraspedon Arrow, 1899
Acrobolbia Ohaus, 1912
Acrossidius Schmidt, 1914
Acrossoides Schmidt, 1914
Acrossus Mulsant, 1842
Acrothyrea Kraatz, 1882
Actinobolus Westwood, 1841
Acylochilus Ohaus, 1909
Adebrattia Bordat & Howden, 1995
Adelaeus Ratcliffe, 2014
Adeloparius Schmidt, 1914
Adoretops Kraatz, 1883
Adoretosoma Blanchard, 1851
Adoretus Dejean, 1833
Adorodocia Brenske, 1893
Adoroleptus Brenske, 1893
Adorrhinyptia Arrow, 1917
Adoryphorus Blackburn, 1889
Adossa Britton, 1995
Adraria Villiers, 1956
Aegialia Latreille, 1807
Aegidiellus Paulian, 1984
Aegidinus Arrow, 1904
Aegidium Westwood, 1845
Aegopsis Burmeister, 1847
Aegostheta Dejean, 1833
Aequatoria Arrow, 1899
Aethiaratrogus Decelle, 1982
Aethiessa Burmeister, 1842
Afroataenius Petrovitz, 1969
Afrodiapterna Dellacasa, 1984
Afrodiastictus Pittino & Mariani, 1986
Afrodrepanus Krikken, 2009
Afroemadiellus Bordat, 2019
Afrolepis Decelle, 1968
Afrotrichonotulus Bordat, 2016
Agamopus Bates, 1887
Aganocrossus Reitter, 1895
Agaocephala Le Peletier & Serville, 1828
Agaocnemis Moser, 1918
Agestrata Eschscholtz, 1829
Aglaphyra Brenske, 1896
Agnorimus Miyake & Iwase, 1991
Agoliinus Schmidt, 1914
Agolius Mulsant & Rey, 1869
Agrilinellus Dellacasa, Dellacasa & Gordon, 2008
Agrilinus Mulsant & Rey, 1869
Ahermodontus Báguena, 1930
Aidophus (Balthasar, 1963)
Airapus Stebnicka & Howden, 1996
Ajmeraphodius Král, Mencl & Rakovič, 2021
Alaia Petrovitz, 1980
Alepida Allsopp, 2018
Aliaclitopa Lacroix, 2003
Alissonotum Arrow, 1908
Aliuscanthoniola Deschodt & Scholtz, 2008
Allara Britton, 1955
Alleucosma Schenkling, 1921
Allidiostoma Arrow, 1940
Alloblackburneus Bordat, 2009
Allobodilus Petrovitz, 1963
Allogymnopleurus Janssens, 1940
Allokotarsa Péringuey, 1904
Allomyrina Arrow, 1911
Allonitis Janssens, 1936
Allophileurinus Dupuis & Dechambre, 2001
Allophyllus Fauvel, 1903
Allorrhina Burmeister, 1842
Alloscelus Boucomont, 1923
Allothnonius Britton, 1978
Alocoderus Schmidt, 1914
Alogistotarsa Péringuey, 1904
Alvarengius Frey, 1975
Alvarinus Blanchard, 1850
Amadotrogus Reitter, 1902
Amaurina Kolbe, 1895
Amazula Kraatz, 1882
Amblochilus Blanchard, 1851
Amblonoxia Reitter, 1902
Amblymelanoplia Dombrow, 2002
Amblyodus Westwood, 1878
Amblyoproctus Kolbe, 1910
Amblyterus MacLeay, 1819
Amerilochus Skelley, 2007
Amerisaprus Stebnicka & Skelley, 2003
Amidorus Mulsant, 1870
Amietina Cambefort, 1981
Amiserica Nomura, 1974
Amithao Thomson, 1878
Ammoecioides Bordat, 1999
Ammoecius Mulsant, 1842
Amorphochelus Fairmaire, 1898
Amphiceratodon Huchet, 2000
Amphimallon Latreille, 1825
Amphistomus Lansberge, 1874
Amphitrichia Brenske, 1894
Ampliodactylus Smith, 2008
Ampotis Dechambre & Madge, 1979
Anacamptorrhina Blanchard, 1842
Anacanthodes Britton, 1990
Anaetius Petrovitz, 1969
Anagnathocera Arrow, 1922
Anahi Martinez, 1958
Analleucosma Antoine, 1989
Anamatochoranus Lacroix, 1995
Anartioschiza Kolbe, 1894
Anatista Brême, 1844
Anatona Burmeister, 1842
Anatonochilus Péringuey, 1907
Ancistrosoma Curtis, 1835
Ancognatha Erichson, 1847
Ancylonyx Britton, 1957
Anelaphinis Kolbe, 1892
Anenaria Lacroix, 1993
Aneucomides Blackburn, 1898
Anisocanthon Martinez & Pereira, 1956
Anisochelus Burmeister, 1844
Anisonyx Latreille, 1807
Anisopholis Moser, 1913
Anisoplia Fischer Von Waldheim, 1824
Anisorrhina Westwood, 1842
Ankaratrotrox Paulian, 1954
Annegialia Howden, 1971
Anochilia Burmeister, 1842
Anocoela Moser, 1914
Anoctus Sharp, 1875
Anodontopopillia Ohaus, 1901
Anomacaulus Fairmaire, 1878
Anomala Samouelle, 1819
† Anomalites Fric, 1885
Anomalochela Moser, 1913
Anomalomorpha Arrow, 1908
Anomalophylla Reitter, 1887
Anomalorhina Jameson, Paucar-Cabrera & Solis, 2003
Anomalurobius Paulian, 1952
Anomiopsoides Blackwelder, 1944
Anomiopus Westwood, 1842
Anomioserica Arrow, 1946
Anomius Mulsant & Rey, 1869
Anomolyna Fairmaire, 1897
Anomonyx Saylor, 1940
Anonychonitis Janssens, 1950
Anoplanomala Arrow, 1917
Anoplocheilus MacLeay, 1838
Anoplodrepanus Simonis, 1981
Anoplognatho Rivers, 1889
Anoplognathus Leach, 1815
Anoplostethus Brullé, 1837
Anoronotum Arrow, 1939
Anoxia Laporte, 1832
Anoxoides Petrovitz, 1971
Antandroyanus Lacroix, 1993
Antecessorirhyparus Minkina, 2020
† Antemnacrassa Gomez Pallerola, 1979
Anthoplia Medvedev, 1949
Anthotocus Britton, 1957
Anthracophora Burmeister, 1842
Anthracophorides Moser, 1918
Anthrencya Lacroix, 1993
Anticheira Eschscholtz, 1818
Anticheiroides Soula, 1998
† Antiqusolidus Lu, Ahrens, Bai, Shih & Ren, 2023
Antitrochalus Brenske, 1900
Antitrogus Burmeister, 1855
Antodon Brême, 1844
Aparammoecius Petrovitz, 1958
Apeltastes Howden, 1968
Aphaenoserica Brenske, 1900
Aphanesia Britton, 1990
Aphanesthes Kraatz, 1880
Aphelinis Antoine, 1987
Aphengium Harold, 1868
Aphengoecus Péringuey, 1901
Aphodaulacus Koshantschikov, 1911
Aphodiellus Schmidt, 1914
Aphodiopsis Paulian, 1942
Aphodius Hellwig, 1798
Aphodobius Péringuey, 1901
Aphodoharmogaster Endrödi, 1983
Aphodopsammobius Endrödi, 1964
Aphonides Rivers, 1889
Aphonodelus Kolbe, 1905
Aphonus LeConte, 1856
Aphotaenius Cartwright, 1952
Apicencya Lacroix, 1993
Aplopsis Blanchard, 1850
Apocnosoides Antoine, 2001
Apogonia Kirby, 1818
Aporecolpa Lansberge, 1886
Aposchiza Brenske, 1903
Apotolamprus Olsoufieff, 1947
Apotriodonta Baraud, 1962
Apsteiniella Schmidt, 1917
Aptenocanthon Matthews, 1974
Apterepilissus Montreuil, 2011
Apteroathlia Smith & Evans, 2018
Apterodemidea Gutiérrez, 1952
Aptychonitis Janssens, 1937
Araboplia Uliana & Sabatinelli, 2017
Arachnodes Westwood, 1842
Archedinus Morón & Krikken, 1990
Archeohomaloplia Nikolajev, 1982
† Archeopsammoporus Minkina, 2020
Archocamenta Brenske, 1895
Archophanes Kolbe, 1905
Archophileurus Kolbe, 1910
Archoserica Brenske, 1897
Arctodium Burmeister, 1844
Ardella Paulsen, 2021
Areoda MacLeay, 1819
Argentophileurus Penco & Zubarán, 2013
Argeremazus Stebnicka & Dellacasa, 2003
Argoplia Dombrow, 2003
Argyripa Thomson, 1878
Arielina Rossi, 1958
Arraphytarsa Péringuey, 1904
Arriguttia Martinez, 1960
Arupaia Stebnicka, 1999
Asactopholis Brenske, 1894
Asaphomorpha Brancsik, 1893
Aschnarhyparus Makhan, 2006
Asiactenius Nikolajev, 2000
Asiopertha (Medvedev, 1949)
Asophrops Petrovitz, 1961
Aspidolea Bates, 1888
Aspilochilus Rojkoff, 2013
Aspilus Westwood, 1874
Astaena Erichson, 1847
Astaenoplia Martinez, 1957
Asthenopholis Brenske, 1898
Asthenorhella Westwood, 1874
Asthenorhina Westwood, 1843
Astibicola Britton, 1990
Astoxenus Péringuey, 1907
Astroscara Schürhoff, 1937
Ataeniopsis Petrovitz, 1973
Ataenius Harold, 1867
Atanyproctus Petrovitz, 1954
Ateuchetus Bedel, 1892
† Ateuchites Meunier, 1898
Ateuchus Weber, 1801
Athesphatoplia Lacroix, 1998
Athlia Erichson, 1835
Atlantemolanum González-Alvarado, Molano-Rendón & Vaz-de-Mello, 2019
Atrichelaphinis Kraatz, 1898
Atrichiana Distant, 1911
Attavicinus Philips & Bell, 2008
Atysilla Strand, 1942
Augoderia Burmeister, 1847
Augosoma Burmeister, 1841
Aulacopalpus Guérin-Méneville, 1838
Aulacopris White, 1859
Aulacoschiza Decelle, 1973
Aulacoserica Brenske, 1900
Aulonocnemis Klug, 1838
Aurum Hutchinson & Moeseneder, 2019
Ausmontins Deschodt & Davis, 2018
Australagolius Mencl, 2008
Australammoecius Petrovitz, 1958
Australaphodius (Balthasar, 1942)
Australoxenella Howden & Storey, 1992
Automolius Britton, 1978
Autoserica Brenske, 1897
Axillonia Krikken, 2018
Bacchusia Mikšić, 1976
Badelina Thomson, 1880
Badiasis Machatschke, 1970
Badizoblax Thomson, 1877
† Baisarabaeus Nikolajev, 2005
Balanogonia Paucar-cabrera, 2003
Balbera Fairmaire, 1898
Ballucus Gordon & Skelley, 2007
Baloghonthobium Paulian, 1986
Balsameda Thomson, 1880
Barryfilius Allsopp, 2022
Barutus Ratcliffe, 1981
Barybas Blanchard, 1850
Basilewskyanus Endrödi, 1956
Basilewskynia Schein, 1957
Batesiana Chalumeau, 1983
Bdelyropsis Pereira & Vulcano & Martinez, 1960
Bdelyrus Harold, 1869
Beckhoplia Dombrow, 2005
Beriqua Péringuey, 1904
Besourenga Vaz-de-Mello, 2008
Bietia Fairmaire, 1898
Bilga Fairmaire, 1893
Bilobatus Machatschke, 1970
Biphyllocera White, 1841
Biralus Mulsant & Rey, 1869
Bisallardiana Antoine, 2003
Bisencya Lacroix, 1993
Bivalgus Paulian, 1961
Bizanus Péringuey, 1902
Blabephorus Fairmaire, 1898
Blackburneus Schmidt, 1914
Blaesia Burmeister, 1842
Blanchardoplia Lacroix, 1998
Blebea Lacroix, 1994
Blepharotoma Blanchard, 1850
Blikana Péringuey, 1902
Blitopertha Reitter, 1903
Bobiricola Endrödi, 1973
Bodiloides Dellacasa & Dellacasa, 2005
Bodilopsis Adam, 1994
Bodilus Mulsant & Rey, 1869
Bohepilissus Paulian, 1975
Bolax Fisher Von Waldheim, 1829
Bolbites Harold, 1868
Boletoscapter Matthews, 1974
Bombodes Westwood, 1848
Bonoraella Ruter, 1978
Bordatius Pittino & Mariani, 1986
Boreocanthon Halffter, 1958
Bosphorus Şenyüz, 2017
Bothrorrhina Burmeister, 1842
Bothynus Hope, 1837
Boucomontiellus Balthasar, 1932
Brachagenius Kraatz, 1890
Brachiaphodius Koshantschikov, 1913
Brachylepis Kolbe, 1894
Brachyllus Brenske, 1896
Brachymacroma Kraatz, 1896
Brachymis Thomson, 1858
Brachypholis Brenske, 1898
Brachysiderus Waterhouse, 1881
Brachysternus Guérin-Méneville, 1831
Bradypodidium Vaz-de-Mello, 2008
Brahmina Blanchard, 1851
Brancoplia Baraud, 1986
Brenskeiella Berg, 1898
Breviclypeus Ricchiardi, 2017
Bricoptis Burmeister, 1842
Brindalus Landin, 1960
Brittonius Özdikmen & Demir, 2008
Bruchaphodius Martinez, 1952
Bruneixenus Howden & Storey, 1992
Bubas Dejan, 1833
Buettikeria Sabatinelli & Pontuale, 1998
Bunbunius Nomura, 1970
Burmeisteriellus Berg, 1898
Burmeistoplia Dombrow, 2003
Butozania Mikšić, 1955
Byrasba Harold, 1869
Byrrhidium Harold, 1869
Byrrhomorpha Blackburn, 1892
Byrsalepis Brenske, 1898
Byrsopolis Burmeister, 1844
Caccobiomorphus Balthasar, 1964
Caccobius Thomson, 1859
Caeconthobium Paulian, 1984
Caelius Lewis, 1895
Caelorrhina Hope, 1841
Caesariatoplia Lacroix, 1998
Calamosternus Motschulsky, 1859
† Calcitoryctes Krell, 2011
Calicnemis Laporte, 1832
Caligodorus Gordon & Skelley, 2007
Callabonica Blackburn, 1895
Calliferoplia Lacroix, 1998
Callipechis Burmeister, 1842
Callirhinus Blanchard, 1851
Callistemonus Péringuey, 1901
Callistethus Blanchard, 1851
Callistopopillia Ohaus, 1903
Calloodes White, 1845
Callophylla Moser, 1916
Calloserica Brenske, 1894
Callynomes Mohnike, 1873
Calocolobopterus Dellacasa, 1984
Calodactylus Blanchard, 1850
Calomacraspis Bates, 1888
Calometopidius Bourgoin, 1917
Calometopus Blanchard, 1850
Calypsoryctes Howden, 1970
Camapterus Ricchiardi, 2000
Cambefortantus Paulian, 1986
Cambefortius Branco, 1989
Camenta Erichson, 1847
Camentoserica Brenske, 1900
Camerounophylla Lacroix, 2005
Camerunopholis Lacroix, 2002
Campsiura Hope, 1831
Campulipus Kirby, 1827
Campylophyllus Decelle, 1979
Campylotrochalus Brenske, 1900
Candezeollus Stebnicka & Howden, 1995
Canestera Saylor, 1938
Canthidium Erichson, 1847
Canthochilum Chapin, 1934
Canthodimorpha Davis, Scholtz & Harrison, 1999
Canthon Hoffmannsegg, 1817
Canthonella Chapin, 1930
Canthonidia Paulian, 1939
Canthonosoma MacLeay, 1871
Canthotrypes Paulian, 1939
Cantotius Bordat, 2022
Canudema Lacroix, 1994
Canuschiza Lacroix, 1999
Carinaulus Tesař, 1945
Carlschoenherria Bezděk, 2016
Carneiola Endrödi, 1974
Carneluttia Mikšić, 1976
Carneodon Özdikmen, 2009
Carneoryctes Özdikmen, 2009
Cartwrightia Islas, 1958
Castanochilus Ohaus, 1909
Castelnauphylla Lacroix, 1999
Catharsiocopris Balthasar, 1967
Catharsius Hope, 1837
Catoclastus Solier, 1851
Catrachia Coca-Abia & Robbins, 2006
Cavonus Sharp, 1875
Caymania Ratcliffe & Cave, 2010
† Ceafornotensis Woolley, 2016
Celidota Burmeister, 1842
Centrantyx Fairmaire, 1884
Centrochilus Krikken, 1976
Centrognathus Guérin-Méneville, 1840
Cephalocyclus Dellacasa & Gordon & Dellacasa, 1998
Cephalodesmius Westwood, 1842
Cephaloncheres Brenske, 1898
Cephaloschiza Moser, 1920
Ceramida Baraud, 1987
Ceraspis Le Peletier & Serville, 1828
Ceratochelus Dombrow, 2002
Ceratogonia Kolbe, 1899
Ceratolontha Arrow, 1948
Ceratophileurus Ohaus, 1911
Ceratoryctoderus Arrow, 1908
Cerhomalus Quedenfeldt, 1884
Ceroplophana Gestro, 1893
Cesamexico Koçak & Kemal, 2008
Cestradoretus Ohaus, 1912
Cetonia Fabricius, 1775
Chaetocosmetes Moser, 1917
Chaetodermina Heller, 1921
Chaetonyx Schaum, 1862
Chaetopisthes Westwood, 1847
Chaetopteroplia Medvedev, 1949
Chaetovalgus Moser, 1914
Chalcasthenes Arrow, 1937
Chalcentis Burmeister, 1844
Chalcochlamys Ohaus, 1905
Chalcocopris Burmeister, 1846
Chalcocrates Heller, 1903
Chalconotus Dejean, 1833
Chalcopharis Heller, 1902
Chalcoplethis Burmeister, 1844
Chalcosoma Hope, 1837
Chalcothea Burmeister, 1842
Chalcotheomima Mikšić, 1970
Chalepides Casey, 1915
Charadronota Burmeister, 1842
Chariochilus Brenske, 1898
Chariodactylus Moser, 1919
Chariodema Blanchard, 1850
Charioserica Brenske, 1899
Charitovalgus Kolbe, 1904
Chasmatopterus Dejean, 1821
Chasme Le Peletier & Serville, 1828
Chaunocolus Saylor, 1937
Cheilo Britton, 1990
Cheiragra MacLeay, 1864
Cheirodontus Britton, 1957
Cheirolasia Westwood, 1842
Cheironitis Lansberge, 1875
Cheiroplatys Hope, 1837
Cheirora Britton, 1957
Cheirotonus Hope, 1840
Cheleion Vardal & Forshage, 2010
Chelilabia Morón & Nogueira, 1998
Cherbezatina Lacroix, 1993
Cheronia Croissandeau, 1893
Chewia Legrand, 2004
Chilenopilus Smith & Mondaca, 2016
Chilodiplus Sharp, 1877
Chiloloba Burmeister, 1842
Chiloschiza Moser, 1924
Chilothorax Motschulsky, 1859
Chilotrogus Reitter, 1905
Chioneosoma Kraatz, 1891
Chipita Soula, 2008
Chiriquibia Bates, 1889
Chiron MacLeay, 1819
Chittius Tarasov, 2008
Chlaenobia Blanchard, 1850
Chloresthia Fairmaire, 1905
Chlorixanthe Bates, 1889
Chlorobapta Kraatz, 1880
Chlorocala Kirby, 1828
Chlorota Burmeister, 1844
Chnaunanthus Burmeister, 1844
Chondropyga Kraatz, 1880
Chondrorrhina Kraatz, 1880
Chromoptilia Westwood, 1842
Chromovalgus Kolbe, 1897
Chrysina Kirby, 1828
Chrysophora Le Peletier & Serville, 1828
Chrysoserica Brenske, 1897
Chthonobius Burmeister, 1847
Chuchina Soula, 2005
Cinacanthus Schmidt, 1914
Circellium Latreille, 1829
Clania Schein, 1956
Clastocnemis Burmeister, 1840
Clavipalpus Laporte, 1832
Cleptocaccobius Cambefort, 1984
Clerota Burmeister, 1842
Clilopocha Lea, 1914
Clinteria Burmeister, 1842
Clinterocera Motschulsky, 1857
Clinteroides Schoch, 1898
Clipadoretus Ohaus, 1941
Clithria Burmeister, 1842
Clitopa Erichson, 1847
Clomecyra Lacroix, 1999
Clypeasta Fairmaire, 1904
Clypeodrepanus Krikken, 2009
Clypeolontha Li & Yang, 1999
Clypeoserica Frey, 1970
Clyster Arrow, 1908
Cnemargulus Semenov, 1903
Cnematoplatys Schmidt, 1914
Cnemida Kirby, 1827
Cnemidophileurus Kolbe, 1910
Cnemisus Motschulsky, 1868
Cnemoschiza Decelle, 1979
Cocherellus Paulian, 1991
Cochinchidema Lacroix, 1996
Cochliotis Kolbe, 1894
Cochliotodes Burgeon, 1946
Coega Péringuey, 1902
Coelocorynus Kolbe, 1895
Coelocratus Burmeister & Schaum, 1841
Coelogenia Moser, 1913
Coelosis Hope, 1837
Coelotrachelus Schmidt, 1914
Coenochilus Schaum, 1841
Coenonycha Horn, 1876
Coenoryctoderus Prell, 1933
Coilodera Hope, 1831
Colacus Ohaus, 1910
Collagenus Ratcliffe & Hardy, 2005
Colobopteridius Clément, 1958
Colobopteroides Paulian, 1942
Colobopterus Mulsant, 1842
Colobostoma Blanchard, 1850
Colpochelyne Britton, 1987
Colpochila Erichson, 1843
Colpochilodes Blackburn, 1898
Colymbomorpha Blanchard, 1850
Comaserica Brenske, 1897
Comatapogonia Lacroix, 2008
Comencya Lacroix, 1993
Comophorina Strand, 1928
Comoramorphochelus Lacroix, 1997
Compsocephalus White, 1845
Compsodactylus Fuhrmann, 2012
Comythovalgus Kolbe, 1884
Conebius Fauvel, 1903
Coniopholis Erichson, 1847
Conioserica Brenske, 1900
Conradtia Kolbe, 1892
Copridaspidus Boucomont, 1920
Coprimorphus Mulsant, 1842
Copris Geoffroy, 1762
Coproecus Reiche, 1841
Coprophanaeus Olsoufieff, 1924
Coptochiroides Balthasar, 1938
Coptochirus Harold, 1859
Coptodactyla Burmeister, 1846
Coptognathus Burmeister, 1847
Coptomia Burmeister, 1842
Coptomiopsis Pouillaude, 1919
Coptorhina Hope, 1833
Corynophyllus Hope, 1845
Corynoserica Moser, 1918
Corynotrichius Kolbe, 1892
Corythoderus Klug, 1845
Coscinocephalus Prell, 1936
Cosmesthes Kraatz, 1880
Cosmiomorpha Saunders, 1852
Cosmiophaena Kraatz, 1898
Cosmovalgus Kolbe, 1897
Costelytra Given, 1952
Costinota Schürhoff, 1933
Cotalpa Burmeister, 1844
Cotinis Burmeister, 1842
Craniorphnus Kolbe, 1895
Craterocyphus Schmidt, 1914
Crathoplus Blanchard, 1851
Cremastocheilus Knoch, 1801
Crepischiza Brenske, 1903
† Cretaclopus Nikolajev, 2004
† Cretaegialia Nikolajev, 1993
† Cretomelolontha Nikolajev, 1998
† Cretonitis Nikolajev, 2007
† Cretorabaeus Nikolajev, 1995
† Cretoscarabaeus Nikolajev, 1995
† Cretoserica Nikolajev, 1998
Cryptaphodius Bordat, 2009
Cryptocanthon Balthasar, 1942
Cryptodontes Burmeister, 1847
Cryptodus MacLeay, 1819
Cryptoscatomaseter Gordon & Skelley, 2007
Cryptotrogus Kraatz, 1888
Cubidens Britton, 1992
Cunderdinia Lea, 1916
Cyclidiellus Krikken, 1976
Cyclidinus Westwood, 1874
Cyclidius MacLeay, 1838
Cyclocephala Dejean, 1821
Cyclomera Klug, 1855
Cylichnus Burmeister, 1844
Cylindrocrates Schein, 1958
Cymboptera Ohaus, 1923
Cymophorus Kirby, 1827
Cyobius Sharp, 1875
Cyphochilus Waterhouse, 1867
Cyphonistes Burmeister, 1847
Cyphonocephalus Westwood, 1842
Cyphonotus Fischer Von Waldheim, 1823
Cyphoserica Brenske, 1900
Cyptochirus Lesne, 1900
Cyriodera Burmeister, 1842
Cyriopertha Reitter, 1903
Cyrtocamenta Brenske, 1897
Cyrtolophus Fairmaire, 1895
Cyrtothyrea Kolbe, 1895
Cyrtotrochalus Brenske, 1900
Dactylopopillia Arrow, 1917
Daintreeola Howden & Storey, 2000
Dalatamala Prokofiev, 2013
Dalgopus Dechambre, 2001
Danielssonia Bordat & Howden, 1995
Dasygnathus MacLeay, 1819
Dasylepida Moser, 1913
Dasytrogus Reitter, 1902
Dasyus Le Peletier & Serville, 1828
Dasyvalgoides Endrödi, 1952
Dasyvalgus Kolbe, 1904
Debeckius Allsopp, 2022
Debutina Lacroix, 1993
Decellophylla Lacroix, 2005
Dechambrophylla Lacroix, 1999
Dedalopterus Sabatinelli & Pontuale, 1998
Degallieridium Vaz-de-Mello, 2008
Dellacasiellus Gordon & Skelley, 2007
Delopleurus Erichson, 1848
Delphinobius Fairmaire, 1900
Deltepilissus Pereira, 1949
Deltochilum Eschscholtz, 1822
Deltorhinum Harold, 1867
Demarziella Balthasar, 1961
Democrates Burmeister, 1847
Dendropaemon Perty, 1830
Denhezia Dechambre, 2006
Dentatoplia Lacroix, 1997
Denticnema Schein, 1959
Dentiheterochelus Dombrow, 2002
Dermolepida Arrow, 1941
Deroserica Moser, 1915
Descarpentriesia Ruter, 1964
Desertaclopus Ocampo & Mondaca, 2012
Desfontainesia Alexis & Delpont, 1999
Desicasta Thomson, 1878
Desmonyx Arrow, 1907
Diabroctis Gistel, 1857
Diaclaspus Brenske, 1896
Diacucephalus De Wailly, 1950
Dialithus Parry, 1849
Dialytellus Brown, 1929
Dialytes Harold, 1869
Dialytodius Gordon & Skelley, 2007
Diaphonia Newman, 1840
Diaphoroserica Péringuey, 1904
Diaphylla Erichson, 1847
Diaplochelus Burmeister, 1844
Diapterna Horn, 1887
Diastellopalpus Lansberge, 1886
Diastictus Mulsant, 1842
Dicaulocephalus Gestro, 1888
Dicellachilus Waterhouse, 1905
Dicentrines Burmeister, 1844
Dichecephala Brenske, 1895
Dichelhoplia Blanchard, 1850
Dichelomorpha Burmeister, 1855
Dichelonyx Harris, 1826
Dicheloschema Moser, 1924
Dichelus Le Peletier & Serville, 1828
Dicheros Gory & Percheron, 1833
Dichodontus Burmeister, 1847
Dichotomius Hope, 1838
Dichrosoma Kraatz, 1885
Dicrania Le Peletier & Serville, 1828
Dicranocara Frolov & Scholtz, 2003
Dicranocnemus Burmeister, 1844
Dicranoplia Reitter, 1903
Dicronocephalus Hope, 1831
Dicronorhina Hope, 1837
Didactylia Orbigny, 1896
Didrepanephorus Wood-mason, 1878
Digenethle Thomson, 1877
Digitonthophagus Balthasar, 1959
Dihymenonyx Gutiérrez, 1949
Dikellites Britton, 1990
Dilatatoplia Lacroix, 1998
Diloboderus Reiche, 1859
Dilochrosis Thomson, 1878
Dilophochila Bates, 1888
Dilortomaeus Bordat, 2009
Dinacoma Casey, 1889
Dinamoraza Lacroix, 1989
Dinarobina Lacroix, 1989
Dinoryctes Felsche, 1906
Diorygopyx Matthews, 1974
Dipelicus Hope, 1842
Diphrontis Gerstaecker, 1882
Diphucephala Le Peletier & Serville, 1828
Diphycerus Fairmaire, 1878
Diphydactylus Thomson, 1858
Diphyllomorpha Hope, 1843
Diploa Kolbe, 1892
Diplognatha Gory & Percheron, 1833
Diplotaxis Kirby, 1837
Diplotropis Boheman, 1857
Dirrhina Burmeister, 1842
Dischista Burmeister, 1842
Discopeltis Burmeister, 1842
Disphysema Harold, 1873
Djadjoua Lacroix, 1993
Djafouna Lacroix, 1993
Doaphius Balthasar, 1941
Doleroserica Péringuey, 1904
Dolerotarsa Péringuey, 1904
Dolichiomicroscelis Dombrow, 2003
Dolichoplia Lacroix, 1998
Dolichostethus Kolbe, 1892
Dorbignyolus Branco, 1989
Doryscelis Burmeister, 1842
Dorysthetus Blanchard, 1845
Doxocalia Brenske, 1897
Drepanocanthoides Schmidt, 1914
Drepanocanthus Péringuey, 1901
Drepanocerus Kirby, 1828
Drepanoplatynus Boucomont, 1921
Drogo Deschodt, Davis & Scholtz, 2016
Dudleyellus Bordat, 2009
Dungoorus Carne, 1958
Dwesasilvasedis Deschodt & Scholtz, 2008
Dymusia Burmeister, 1842
Dynastes MacLeay, 1819
Dyscinetus Harold, 1869
Dysphanochila Blackburn, 1898
Ebolowanius Dechambre, 1979
Eccoptomia Kraatz, 1880
Echyra Erichson, 1847
Ectinohoplia Redtenbacher, 1868
Ectinoplectron Ohaus, 1915
Eideria Neita & Ocampo, 2012
Elaphinis Burmeister, 1842
Elaphocera Gene, 1836
Elaphocerella Moser, 1919
Elcarmeniella Franz, 1955
Elpidus Péringuey, 1907
Emadiellus Schmidt, 1914
Embrithoplia Lacroix, 1998
Empecamenta Brenske, 1895
Empecta Erichson, 1847
Empectoides De Wailly, 1950
Emphania Erichson, 1847
Empycastes Gerstaecker, 1871
Enamillus Sharp, 1877
Enaria Erichson, 1847
Enarotadius Endrödi, 1971
Encya Dejean, 1833
Endebius Lansberge, 1880
Endoxazus Kolbe, 1892
Endroedianibe Chalumeau, 1981
Endroedyolus Scholtz & Howden, 1987
Enema Hope, 1837
Engertia Dalle Torre, 1912
Engistanoxia Lacroix, 2002
Engyopsina Britton, 1990
Ennearabdus Lansberge, 1874
Enoplotarsus Lucas, 1859
Enracius Dechambre, 1999
Enthora Erichson, 1847
Entypophana Moser, 1913
Entyposis Kolbe, 1894
Eocorythoderus Marumaya, 2012
Eodrepanus Barbero, Palestrini & Roggero, 2009
Eophileurus Arrow, 1908
Eosaulostomus Carne, 1956
Eotrichia Medvedev, 1951
Epactoides Olsoufieff, 1947
Epadoretus Reitter, 1903
Epectinaspis Blanchard, 1851
Epicaulis Dejean, 1833
Epichalcoplethis Bates, 1904
Epichrysus White, 1841
Epidrepanus Roggero, Barbero & Palestrini, 2015
Epilissus Reiche, 1841
Epipholis Moser, 1917
Epirinus Dejean, 1833
Epironastes Carne, 1957
Epistalagma Fairmaire, 1880
Epitrichius Tagawa, 1941
Epixanthis Burmeister, 1842
Erbmahcedius Hutchinson & Allsopp, 2021
Ercomoana Lacroix, 1994
Eremazus Mulsant, 1851
Eremobothynus Ohaus, 1910
Eremophygus Ohaus, 1910
Eremoplia Kolbe, 1914
Eremotrogus Kolbe, 1914
Eriesthis Burmeister, 1844
Erigavo Lacroix, 2006
Eriopeltastes Burmeister & Schaum, 1840
Erioscelis Burmeister, 1847
Eriphoserica Brenske, 1899
Eriulis Burmeister, 1842
Erlangeria Preiss, 1902
Erytodes Schmidt, 1917
Erytus Mulsant & Rey, 1870
Escarabaeus Zídek & Pokorný, 2011
Esymus Mulsant & Rey, 1869
Etiserica Péringuey, 1904
Eucamenta Péringuey, 1904
Euchilia Burmeister, 1842
Euchirus Burmeister & Schaum, 1840
Euchloropus Arrow, 1907
Euchroea Burmeister, 1842
Eucopidocaulus Prell, 1912
Eucranium Brullé, 1837
Eucyclophylla Waterhouse, 1875
Eudicella White, 1839
Eudinopus Burmeister, 1840
Eudolus Mulsant & Rey, 1869
Euetheola Bates, 1888
Euhemicyclium Bordat, 2003
Euheptaulacus Dellacasa, 1983
Euknemoplia Lacroix, 1997
Eulaiades Fairmaire, 1899
Eulepida Kolbe, 1894
Eulepidopsis Burgeon, 1946
Euligyrus Casey, 1915
Eumacronota Mikšić, 1976
Eunanus Ohaus, 1909
Euoniticellus Janssens, 1953
Euonthophagus Balthasar, 1959
Euorodalus Dellacasa, 1983
Euotophorus Dellacasa, Gordon & Dellacasa, 2016
Euparia Le Peletier & Serville, 1828
Euparixia Brown, 1927
Euparixoides Hinton, 1936
Euparotrix Stebnicka & Howden, 1996
Eupatorus Burmeister, 1847
Eupegylis Duvivier, 1892
Euphoresia Brenske, 1897
Euphoria Burmeister, 1842
Eupleurus Mulsant, 1842
Eupoecila Burmeister, 1842
Euranoxia Semenov, 1890
Euremina Westwood, 1867
Europtron Marseul, 1867
Euryaspis Blanchard, 1851
Eurychelus Blanchard, 1850
Euryomia Burmeister, 1842
Eurypeza Lacroix, 2006
Euryschiza Brenske, 1898
Eurysternus Dalman, 1824
Euryvalgus Moser, 1908
Eusaproecius Branco, 1989
Euselates Thomson, 1880
Euserica Reitter, 1896
Eutelesmus Waterhouse, 1880
Eutrichesis Waterhouse, 1882
Eutrichillum Martinez, 1967
Eutyctus Semenov, 1890
Evanos Castelnau, 1840
Evbrittonia Szito, 1993
Exanticheira Soula, 1998
Exaphodius Endrödi, 1960
Eximidema Keith & Reichenbach, 2018
Exochlorota Soula, 2002
Exochogenys Carne, 1958
Exolontha Reitter, 1902
Exomala Reitter, 1903
Exopholis Motschulsky, 1859
Exoptenomela Soula, 2002
Exostethus Britton, 1979
Exothyridium Soula, 2002
Exoxyomus Bordat, Mencl & Rakovič, 2012
Extenuoptyophis Smith & Mondaca, 2015
Faargia Martinez, 1982
Fairmairoplia Lacroix, 1997
Falsignambia Paulian, 1987
Falsotoclinius Keith & Montreuil, 2004
Faula Blanchard, 1850
Feeridium Vaz-de-Mello, 2008
Ferrerianus Dellacasa, Dellacasa & Gordon, 2007
Firminus Coca-Abia, 2003
Flaviellus Gordon & Skelley, 2007
Flechtmanniella Stebnicka, 1999
Fornasinius Bertoloni, 1852
Fossocarus Howden, 1961
Frankenbergerius Balthasar, 1938
Frenchella Blackburn, 1892
Fruhstorferia Kolbe, 1894
Fulvaphodius Bordat, 2022
Gametis Burmeister, 1842
Gametoides Antoine, 2005
Gamka Péringuey, 1904
Ganganomala Ratcliffe, Jameson & Zorn, 2018
Gariep Péringuey, 1907
Garreta Janssens, 1940
Gastromaladera Nomura, 1973
Gastroserica Brenske, 1897
Geniates Kirby, 1818
Geniatosoma Costa Lima, 1940
Genieridium Vaz-de-Mello, 2008
Genuchinus Westwood, 1874
Genuchus Kirby, 1825
Geomyphilus Gordon & Skelley, 2007
Geopsammodius Gordon & Pittino, 1992
Geotrogus Guérin-Méneville, 1842
Gibboryctes Endrödi, 1974
Giesbertiolus Howden, 1988
Gillaspytes Howden, 1980
Gilletellus Janssens, 1937
Gilletianus Balthasar, 1933
Glacilitarsus Matsumoto, 2014
Glaphyronyx Moser, 1924
Glaphyserica Brenske, 1899
Glenopopillia Lin, 1980
Globencya Lacroix, 1993
Globulischiza Frey, 1974
Glossocheilifer Blackburn, 1898
Glycosia Schoch, 1896
Glycyphana Burmeister, 1842
Glycyserica Brenske, 1899
Glyphoderus Westwood, 1838
Glyptoglossa Brenske, 1895
Glyptothea Bates, 1889
Glyptotheomima Mikšić, 1976
Gnaphalopoda Reiche, 1860
Gnaphalostetha Reiche, 1856
Gnathocera Kirby, 1825
Gnathogolofa Arrow, 1914
Gnatholabis Erichson, 1847
Gnorimella Casey, 1915
Gnorimidia Lansberge, 1887
Gnorimimelus Kraatz, 1880
Gnorimotrichius Krajčik, 2009
Gnorimus Le Peletier & Serville, 1828
Goiginus Endrödi, 1982
Goliathopsis Janson, 1881
Goliathus Lamarck, 1801
Golinca Thomson, 1878
Golofa Hope, 1837
Gonaphodiellus Schmidt, 1914
Gonaphodioides Dellacasa, Dellacasa & Gordon, 2012
Gonaphodiopsis Dellacasa, Dellacasa & Gordon, 2012
Gonaphodius Reitter, 1892
Gongrolophus Stebnicka & Howden, 1996
Goniaspidius Burmeister, 1844
Goniophileurus Kolbe, 1910
Goniorphnus Arrow, 1911
Goniorrhina Quedenfeldt, 1888
Gorditus Ratcliffe, 2010
Gordonius Skelley, Dellacasa & Dellacasa, 2009
Gouna Péringuey, 1902
Gracilaclopus Ocampo & Mondaca, 2012
Grammodoplia Lacroix, 1997
Grammopyga Kolbe, 1895
Grandaustralis Hutchinson & Moeseneder, 2013
Grandinaphodius Ziani, 2002
Granulopsammodius Rakovič, 1981
Graviengertia Prokofiev, 2016
Grebennikovius Mlambo, Scholtz & Deschodt, 2019
Griveaudella Lacroix, 1993
Gromphas Brullé, 1837
Gronocarus Schaeffer, 1927
Gryphonycha Péringuey, 1904
Guanyinaphodius Masumoto & Kiuchi, 2001
Guatemalica Neervoort Van De Poll, 1886
Gymnephoria Ratcliffe, 2019
Gymnetina Casey, 1915
Gymnetis MacLeay, 1819
Gymnogaster Blanchard, 1851
Gymnoloma Dejean, 1833
Gymnophana Arrow, 1910
Gymnopleurus Illiger, 1803
Gymnopyge Linell, 1896
Gymnoschiza Moser, 1914
Gynaecoserica Brenske, 1896
Gyronotus Lansberge, 1874
Gyroplia Brenske, 1893
Hadrodiplognatha Kraatz, 1898
Hadropechys Britton, 1990
Hadrops Britton, 1987
Hadrorhyparus Howden, 1995
Hadrosticta Kraatz, 1892
Halffterinetis Morón & Nogueira, 2007
Hamatoplectris Frey, 1967
Hammondantus Cambefort, 1978
Hamonthophagus Roggero, Dierkens, Barbero & Palestrini, 2017
Hansreia Halffter & Martinez, 1977
Haplidia Hope, 1837
Haplophileurus Kolbe, 1910
Haploscapanes Arrow, 1908
Haplosoma Semenov, 1890
Harmodactylus Péringuey, 1901
Harmogaster Harold, 1861
Haroldaphodius Bordat, 2003
Haroldiataenius Chalumeau, 1981
Haroldiellus Gordon & Skelley, 2007
Haroldius Boucomont, 1914
Harpechys Britton, 1957
Harpina Burmeister, 1844
Harposceles Burmeister, 1847
Hatamus Sharp, 1877
Hecistopsilus Kolbe, 1894
Hegemus Thomson, 1881
Helictopleurus Orbigny, 1915
Heliocopris Hope, 1837
Hellaserica Baraud & Nicolas, 1966
Hemiaspidius Krikken, 1982
Hemicamenta Brenske, 1897
Hemichaetoplia Baraud, 1986
Hemichalcothea Mikšić, 1970
Hemichnoodes Kraatz, 1880
Hemicorythoderus Tangelder & Krikken, 1982
Hemictenius Reitter, 1897
Hemicyclium Schmidt, 1914
Hemicyrthus Reiche, 1860
Hemiheterorrhina Mikšić, 1974
Hemilia Kraatz, 1880
Hemiphaedimus Mikšić, 1972
Hemipharis Burmeister, 1842
Hemiphileurus Kolbe, 1910
Hemiprotaetia Mikšić, 1963
Heptaulacus Mulsant, 1842
Heptelia Brenske, 1898
Heptophylla Motschulsky, 1857
Herculaisia Seilliere, 1910
Heteralleucosma Antoine, 1989
Heterochelus Burmeister, 1844
Heterochlorota Soula, 2002
Heteroclita Burmeister, 1842
Heteroclitopus Péringuey, 1901
Heterocnemis Albers, 1852
Heteroconus Kolbe, 1900
Heterocotinis Martinez, 1948
Heterocranus Bourgoin, 1919
Heterogeniates Ohaus, 1909
Heterogenius Moser, 1911
Heteroglobus Dupuis & Dechambre, 2008
Heterogomphus Burmeister, 1847
Heteroligus Kolbe, 1900
Heteronitis Gillet, 1911
Heteronychus Dejean, 1833
Heteronyx Guérin-Méneville, 1830
Heterophana Burmeister, 1842
Heterophthalmus Blanchard, 1851
Heteropseudinca Valck Lucassen, 1933
Heterorhina Westwood, 1842
Heteroschiza Moser, 1917
Heteroserica Brenske, 1899
Heterosoma Schaum, 1844
Heterosternus Dupont, 1832
Heterostriatus Dupuis, 2006
Heterotephraea Antoine, 2002
Heterotrochalus Moser, 1918
Heterovalgus Krikken, 1978
Hexataenius Fairmaire, 1891
Hexodon Olivier, 1789
Hiberasta Fairmaire, 1901
Hiekeianus Endrödi, 1978
Hieritis Burmeister, 1855
Hilyotrogus Fairmaire, 1886
Himalhoplia Sabatinelli, 1983
Hispanioryctes Howden & Endrodi, 1978
† Holcorobeus Nikritin, 1977
Holisonycha Péringuey, 1904
Holocanthon Martinez & Pereira, 1956
Holocephalus Hope, 1838
Holochelus Reitter, 1889
Holocnemus Brenske, 1894
Hologymnetis Martinez, 1949
Holomelia Brenske, 1891
Holopycnia Brenske, 1896
Holorhopaea Britton, 1978
Holotrichia Hope, 1837
Homalochilus Blanchard, 1851
Homaloserica Brenske, 1897
Homalotarsus Janssens, 1932
Homeochlorota Soula, 2006
Homocopris Burmeister, 1842
Homoeomorphus Burmeister, 1847
Homoeoschiza Kolbe, 1894
Homoiosternus Ohaus, 1901
Homolotropus MacLeay, 1871
Homonyx Guérin-Méneville, 1839
Homophileurus Kolbe, 1910
Homopliopsis Lacroix, 1998
Homothermon Ohaus, 1898
Homothyrea Kolbe, 1895
Hoplebaea Brenske, 1899
Hopleidos Künckel D'Herculais, 1887
Hoplia Illiger, 1803
Hopliopsis Blanchard, 1850
Hoplitovalgus Kolbe, 1904
Hoplochelus Blanchard, 1851
Hoplocnemis Harold, 1869
Hoplolontha Fairmaire, 1904
Hoplomaladera Nomura, 1974
Hoplopelidnota Bates, 1904
Hoplopus Laporte, 1832
Hoplopyga Thomson, 1880
Hoplopygothrix Schürhoff, 1933
Hoplorida Moser, 1918
Hoploryctoderus Prell, 1933
Hoplosternodes Burgeon, 1946
Hornietus Stebnicka, 2000
Hornosus Dellacasa, Dellacasa & Gordon, 2015
Horridocalia Endrödi, 1974
Hovachelus Fairmaire, 1897
Hovophileurus Arrow, 1911
Howdenocarus Hardy, 1978
Howdenypa Arnaud, 1993
Humblotania Lacroix, 1993
Hyalonthophagus Palestrini & Giacone, 1988
Hybaloides Quedenfeldt, 1884
Hybalus Dejean, 1833
Hybocamenta Brenske, 1898
Hyboschema Péringuey, 1901
Hyboserica Péringuey, 1904
† Hybosorites Nikolajev, 1996
Hybovalgus Kolbe, 1904
Hylamorpha Arrow, 1899
Hylobothynus Ohaus, 1910
Hymenochelus Reitter, 1890
Hymenoplia Eschscholz, 1830
Hypaspidius Arrow, 1899
Hyperius Fairmaire, 1878
Hyphelithia Kraatz, 1880
Hyphoryctes Blackburn, 1895
Hypocanthidium Balthasar, 1938
Hypolepida Britton, 1978
Hyposerica Brenske, 1897
Hypothyce Howden, 1968
Hypotrichia LeConte, 1861
Hypselogenia Burmeister, 1840
Iarupea Martinez, 1953
Iberoaphodius Rössner, 2018
Ichnestoma Gory & Percheron, 1833
Ictigaster Britton, 1986
Idaecamenta Péringuey, 1904
Idaeserica Péringuey, 1904
Idanastes Britton, 1987
Idiapogonia Arrow, 1916
Idiochelyna Arrow, 1916
Idionycha Arrow, 1932
Idiovalgus Arrow, 1910
Idutywa Péringuey, 1902
Ignambia Heller, 1916
Iguazua Stebnicka, 1997
Ihosyus Montreuil & Lacroix, 2020
Inanda Péringuey, 1902
Inca Le Peletier & Serville, 1828
Incala Thomson, 1857
Indieraligus Dechambre, 1979
Indoblackburneus Mencl, Král & Rakovič, 2019
Indognorimus Krikken, 2009
Indotrichius Krikken, 2009
Indovalgus Ricchiardi, 2012
Ingogius Endrödi, 1976
Ingrisma Fairmaire, 1893
Insimuloissacaris Smith & Mondaca, 2015
Iranotrogus Nikolajev, 2006
Irazua Ratcliffe, 2003
Iridisoma Delgado & Morón, 1991
Irrasinus Gordon & Skelley, 2007
Isacanthon Pacheco & Vaz-de-Mello, 2019
Ischiopsopha Gestro, 1874
Ischnopopillia Kraatz, 1892
Ischnoscelis Burmeister, 1842
Ischnostomiella Krikken, 1978
Ischnotarsia Kraatz, 1880
Ischnovalgus Kolbe, 1897
Isocopris Pereira & Martinez, 1960
Isonychus Mannerheim, 1828
Isoplia Burmeister, 1855
Issacaris Fairmaire, 1889
Ixodina Roth, 1851
Ixorida Thomson, 1880
Jalalabadia Balthasar, 1967
Jalisco Dellacasa, Gordon & Dellacasa, 2003
Jansonia Schürhoff, 1937
Janssensantus Paulian, 1976
Janssensellus Cambefort, 1975
Jossonthophagus Dierkens, Moretto & Génier, 2017
Joziratia Lacroix, 1993
Jumnos Saunders, 1839
Junkia Dalla Torre, 1913
† Juraclopus Nikolajev, 2005
Kabindeknomiosoma Lacroix, 2005
Kadlecia Frey, 1965
Kareiga Péringuey, 1902
Karolinella Minkina, 2017
Karooida Perissinotto, 2020
Kenyalepis Lacroix, 2011
Kenyatopadema Lacroix, 2009
Kerowagia Delpont, 1996
Kheper Janssens, 1940
Khoina Péringuey, 1902
Kibakoganea Nagai, 1984
Kirprellius Allsopp, 2022
Knysna Péringuey, 1902
Kolbeellus Jacobson, 1906
Komrina Lacroix, 1993
Koreoxyomus Kim, 1996
Korisaba Péringuey, 1902
Koshantschikovius Schmidt, 1914
Kraseophylla Péringuey, 1904
Krikkenius Branco, 1991
Kryzhanovskia Nikolajev & Kabakov, 1977
Kubousa Péringuey, 1902
Kuenckeliana Lacroix, 1989
Kurtops Roggero, Barbero & Palestrini, 2017
Labarrus Mulsant & Rey, 1869
Labroma Sharp, 1873
Laceratoplia Lacroix, 1997
Lachnodera Erichson, 1847
Lachnota Reitter, 1889
Lacroixidema Keith, 2000
Lacroixilepis Keith, 2007
Lagochile Hoffmannsegg, 1817
Lamellothyrea Krikken, 1980
Lamproserica Brenske, 1897
Lansbergia Ritsema, 1888
Laotrichia Keith, 2007
Larhodius Balthasar, 1963
Lasiexis Semenov-Tian-Shanskii & Medvedev, 1936
Lasiocala Blanchard, 1851
Lasiopsis Erichson, 1847
Lasioserica Brenske, 1897
Lasiotrichius Reitter, 1899
Lasiotropus Reitter, 1913
Latipalpus Moser, 1921
Latodrepanus Krikken, 2009
Lebbea Lacroix, 1993
Lecanoderus Kolbe, 1907
Lecanotrogus Kolbe, 1894
Lechorodius Gordon & Skelley, 2007
Leiopsammodius Rakovič, 1981
Lenosoma MacLeay, 1863
Leonotus Britton, 1986
Leoserica Fabrizi, Eberle & Ahrens, 2019
Leotrichillum Vaz-de-Mello, 2008
Lepanus Balthasar, 1966
Lepidiota Kirby, 1828
Lepidomela Kolbe, 1894
Lepidoserica Nikolajev, 1979
Lepidotrogus Kolbe, 1894
Lepischiza Moser, 1914
Lepisia Le Peletier & Serville, 1828
Lepithrix Le Peletier & Serville, 1828
Lepitrichula Schein, 1959
Lepivalgus Moser, 1914
Leptochristina Baraud & Branco, 1991
Leptocnemis Dombrow, 2001
Leptohoplia Saylor, 1935
Leptorhyparus Howden, 2003
Leribe Péringuey, 1904
Leucocelis Burmeister, 1842
Leucopholis Dejean, 1833
Leucophorus Brenske, 1892
Leucothyreus MacLeay, 1819
Leuretra Erichson, 1847
Liatongus Reitter, 1892
Lichnia Erichson, 1835
Lichniops Gutiérrez, 1946
Lichniopsoides Martinez, 1953
Licnostrategus Prell, 1933
Ligyrus Burmeister, 1847
Limarus Mulsant & Rey, 1869
Limepomera Lacroix, 1999
Linargius Rakovič & Mencl, 2012
Lindbergianus Petrovitz, 1961
Liogenys Guérin-Méneville, 1830
Lioglyptoxenus Pittino, 2006
Liomenochilus Hutchinson & Allsopp, 2022
Liostraca Burmeister, 1842
Liothorax Motschulsky, 1859
Liotrichius Kolbe, 1892
Liparetrus Guérin-Méneville, 1830
Lissadoretus Arrow, 1917
Lissogenius Schaum, 1844
Lissomelas Bates, 1889
Listrochelus Blanchard, 1851
† Lithanomala Nikolajev, 1992
† Lithoscarabaeus Nikolajev, 1992
Litocopris Waterhouse, 1891
† Lobateuchus Montreuil, Génier & Nel, 2010
Lobogeniates Ohaus, 1917
Loboparius Schmidt, 1914
Loboserica Kolbe, 1914
Lobovalgus Kolbe, 1897
Lomanoxia Martinez, 1951
Lomanoxoides Stebnicka, 1999
Lomaptera Gory & Percheron, 1833
Lonchothyrea Kolbe, 1895
Lonchotus Burmeister, 1847
Longaphodius Endrödi, 1980
Longicrura Frey, 1974
Lophodonitis Janssens, 1938
Lophorrhina Westwood, 1842
Lophorrhinides Perissinotto, Clennell & Beinhundner, 2019
Loraphodius Reitter, 1892
Loraspis Mulsant & Rey, 1869
Lorditomaeus Péringuey, 1901
Loricatarsa Frey, 1969
Lorkovitschia Mikšić, 1968
Lucassenia Olsoufieff, 1940
Luispenaia Martinez, 1972
Lunaphodius Balthasar, 1964
Lutera Westwood, 1875
Lutfius Özdikmen, 2009
Luxolinus Gordon & Skelley, 2007
Lycomedes Brême, 1844
Lyraphora Kraatz, 1880
Machala Lacroix, 1993
Macleayella Britton, 1990
Macraspis MacLeay, 1819
Macrocranius Schürhoff, 1935
Macrocyphonistes Ohaus, 1910
Macrodactylus Latreille, 1825
Macroderes Westwood, 1842
Macromina Westwood, 1874
Macronota Hoffmannsegg, 1817
Macronotops Krikken, 1977
Macroplia Brenske, 1898
Macropliopsis Lacroix, 1997
Macropoidelimus Morón, 1983
Macropoides Guérin-Méneville, 1844
Macropopillia Ohaus, 1905
Macroretroides Bordat, Dellacasa & Dellacasa, 2000
Macroretrus Péringuey, 1908
Macrotina Strand, 1934
Maculaphodius Gordon & Skelley, 2007
Madahoplia Lacroix, 1998
Madecorphnus Paulian, 1992
Madiana Ratcliffe & Romé, 2019
Madiniella Chalumeau & Gruner, 1976
Maechidinus Lea, 1919
Maechidius MacLeay, 1819
Maghrebotrogus Montreuil & Keith, 2022
Mahafalyanus Lacroix, 1993
Makalaka Péringuey, 1908
Makoanus Lacroix, 1993
Maladera Mulsant, 1871
Malagoniella Martinez, 1961
Malaia Heller, 1891
Malaisius Arrow, 1941
Malawinophylla Lacroix, 2013
Mallotarsus Blanchard, 1850
Manjarivolo Paulian, 1974
Manodactylus Moser, 1919
Manonychus Moser, 1919
Manopus Castelnau, 1840
Manoserica Moser, 1924
Marmarina Kirby, 1827
Marmylida Thomson, 1880
Marronus Coquerel, 1866
Martineziana Chalumeau & Özdikmen, 2006
Martinezidium Vaz-de-Mello, 2008
Mascarena Arrow, 1919
Masumotokoganea Hirasawa, 1992
Matthewsius Gunter & Weir, 2017
Mauromecistoplia Dombrow, 2002
Mausoleopsis Lansberge, 1882
Mawenzhena Alexis & Delpont, 2001
Mayataia Lacroix, 1993
Mazahuapertha Morón & Nogueira, 1998
Mecaspidiellus Antoine, 1997
Mecopelidnota Bates, 1904
Mecynodes Mulsant, 1870
Mecynorhina Hope, 1837
Medeotrogus Keith, 2001
Megaceras Hope, 1837
Megaceropsis Dechambre, 1976
Megacoryne Britton, 1987
Megalonitis Janssens, 1937
Megapertha (Reitter, 1903)
Megaphonia Schürhoff, 1933
Megarhopaea Britton, 1978
Megasoma Kirby, 1825
Megasybacodes Kakizoe, Maruyama & Masumoto, 2019
Megateloides Landin, 1974
Megatelus Reitter, 1892
Megatharsis Waterhouse, 1891
Megathopa Eschscholtz, 1822
Megathoposoma Balthasar, 1939
Megistophylla Burmeister, 1855
Megistoplia Lacroix, 1997
Melanchroea Kraatz, 1900
Melanhyphus Fairmaire, 1881
Melanocamenta Brenske, 1899
Melanocanthon Halffter, 1958
Melanopopillia Lin, 1980
Melinopterus Mulsant, 1842
Mellissius Wollaston, 1869
Melolontha Fabricius, 1775
† Melolonthites Heer, 1847
Melolonthoides De Wailly, 1950
Mendidaphodius Reitter, 1901
Mendidius Harold, 1868
Menigionyx Arrow, 1941
Mentophilus Castelnau, 1840
Mericserica Brenske, 1897
Meridioclita Krikken, 1982
Merinanus Lacroix, 1993
Merogyrus Gordon & Skelley, 2007
Meroloba Thomson, 1880
Mesomerodon Ohaus, 1905
Mesontoplatys Motschulsky, 1863
Mesoserica Brenske, 1897
Mesosternus Morón, 1987
Mesotrochalus Kolbe, 1914
Messyrhus Rakovič & Král, 1997
† Mesydra Ponomarenko, 1977
Mesystoechus Waterhouse, 1878
Metacatharsius Paulian, 1939
Metadorodocia Machatschke, 1957
Metallesthes Kraatz, 1880
Metallopseudinca Valck Lucassen, 1933
Metanastes Arrow, 1911
Metapachylus Bates, 1889
Metaphileurus Kolbe, 1910
Metapogonia Lacroix, 2008
Metascelis Westwood, 1842
Metatrogus Britton, 1978
Metoporaphodius Frolov & Akhmetova, 2017
Meurguesia Ruter, 1969
Michaeloplia Lacroix, 1997
Micraegialia Brown, 1931
Micranomala Arrow, 1911
Micreuchilia Pouillaude, 1917
Microchalcothea Moser, 1910
Microchilus Blanchard, 1851
Microcoenus Britton, 1957
Microcopris Balthasar, 1958
Microdilochrosis Jákl, 2009
Microdoris Burmeister, 1844
Microlomaptera Kraatz, 1885
Microlontha Petrovitz, 1967
Microogenius Gutiérrez, 1951
Micropeltus Blanchard, 1842
Micropertha Baraud, 1991
Microphileurus Kolbe, 1910
Microphylla Kraatz, 1890
Microplidus Péringuey, 1902
Microplus Burmeister, 1844
Micropoecila Kraatz, 1880
Microrhopaea Lea, 1920
Microrutela Bates, 1904
Microryctes Arrow, 1908
Microserica Brenske, 1894
Microsericaria Nikolajev, 1979
Microtermitodius Pittino, 2006
Microteuchestes Landin, 1974
Microthopus Burmeister, 1855
Microtrochalus Brenske, 1900
Microvalgus Kraatz, 1883
Mictophileurus Ohaus, 1911
Milatrogus Montreuil & Keith, 2022
Milichus Péringuey, 1901
Mimadoretus Arrow, 1901
Mimaegialia Nikolajev, 2007
Mimela Kirby, 1823
Mimeoma Casey, 1915
Mimogeniates Martinez, 1964
Mimonthophagus Balthasar, 1963
Mimovalgus Arrow, 1944
Minidorysthetus Soula, 1998
Minisiderus Arrow, 1902
Minutoplia Lacroix, 1998
† Miolachnosterna Wickham, 1914
Miotemna Lacordaire, 1855
Mireia Ruter, 1953
Miridiba Reitter, 1902
Mitracamenta Brenske, 1903
Mitracephala Thomson, 1859
Mitrophorus Burmeister, 1844
Mnematidium Ritsema, 1888
Mnematium MacLeay, 1821
Modialis Fairmaire & Germain, 1860
Molynoptera Kraatz, 1897
Molynopteroides Antoine, 1989
† Mongolrobeus Nikolajev, 2004
Monochelus Le Peletier & Serville, 1828
Monoplistes Lansberge, 1874
Monotropus Erichson, 1847
Monteitheolus Howden & Storey, 2000
Monteithocanthon Gunter & Weir, 2017
Morettius Roggero, Dierkens, Barbero & Palestrini, 2017
Moriaphila Kraatz, 1880
Morokia Janson, 1905
Moroniella Ramírez-Ponce, 2015
Moronius Grossi & Vaz-de-Mello, 2015
Moronoserica Ahrens, Lukic & Liu, 2023
Moseriana Ruter, 1965
Mothon Semenov & Medvedev, 1927
Motuotrichius Huang & Chen, 2016
Mozartius Nomura & Nakane, 1951
Mucama Soula, 2002
Mucrencya Lacroix, 1993
Musurgus Vauloger, 1898
Mycernus Broun, 1904
Mycterophallus Neervoort Van De Poll, 1886
Myloxena Berg, 1881
Myloxenoides Martinez, 1975
Myodermides Ruter, 1964
Myodermidius Bourgoin, 1920
Myodermum Burmeister, 1840
Myrhessus Balthasar, 1955
Myrmecaphodius Martinez, 1952
Myrmecochilus Wasmann, 1900
Mysarus Petrovitz, 1962
Mystroceros Burmeister, 1842
Mzansica Perissinotto, 2020
Nagainokoganea Hirasawa, 1992
Namakwanus Scholtz & Howden, 1987
Namaphilus Deschodt & Davis, 2017
Nanaga Péringuey, 1902
Nanniscus Burmeister, 1844
Nannopopillia Kolbe, 1894
Nanorhopaea Britton, 1978
Nanos Westwood, 1842
Nanotermitodius Howden, 2003
Napoa Stebnicka, 1999
Narycius Dupont, 1835
Navigator Moeseneder & Hutchinson, 2016
Nayarita Morón & Nogueira, 1998
Neagolius Koshantschikov, 1912
Nebulasilvius Deschodt & Scholtz, 2008
Nedymoserica Brenske, 1900
Nefoncerus Saylor, 1938
Nematophylla Arrow, 1906
Nematoserica Arrow, 1917
Neocalaphodius Bordat, 1990
Neocamenta Burgeon, 1945
Neoclita Perissinotto, 2017
Neoclithria Neervoort Van De Poll, 1886
Neoclitopa Lacroix, 1997
Neocnecus Sharp, 1878
Neocolobopterus Landin, 1974
Neocorvicoana Ratcliffe & Mico, 2001
Neocorynophyllus Carne, 1985
Neodasygnathus Carne, 1957
Neodiapterna Dellacasa, 1984
Neoemadiellus Clément, 1986
Neogutierrezia Martinez, 1953
Neoheptaulacus Paulian & Villiers, 1941
Neoheteronyx Blackburn, 1890
Neohyphus Heller, 1896
Neomaladera Baraud, 1965
Neometanastes Yamaya, 2010
Neomyrhessus Minkina, 2023
Neomystroceros Alexis & Delpont, 1998
Neonastes Carne, 1957
Neonitis Péringuey, 1901
Neopegylis Lacroix, 2015
Neophaedimus Lucas, 1870
Neophaenognatha Allsopp, 1983
Neophyllotocus Blackburn, 1898
Neopsammodius Rakovič, 1986
Neorhyssemus Gordon & Pittino, 1992
Neorrhina Thomson, 1878
Neoryctes Arrow, 1908
Neosaproecius Branco, 1990
Neoscelis Schoch, 1897
Neoserica Brenske, 1894
Neosericania Miyake & Yamaya, 1994
Neotrichaphodioides Dellacasa, Dellacasa & Skelley, 2010
Neotrichiorhyssemus Rakovič & Král, 1997
Neotrichonotulus Dellacasa, Gordon & Dellacasa, 2004
Neovalgus Miyake, 1985
Nepaloserica Frey, 1965
Nephrodopus Sharp, 1873
Nepytis Erichson, 1842
Nesadmiralitia Prokofiev, 2016
Neso Blackburn, 1898
Nesohoplia Scott, 1912
Nesosisyphus Vinson, 1946
Nesovinsonia Martinez & Pereira, 1958
Nettelislasia Martinez, 1952
Neuquenodactylus Smith & Mondaca, 2015
Neuroserica Brenske, 1900
Nialaphodius Kolbe, 1908
Nialosternus Hollande & Therond, 1998
Nialus Mulsant & Rey, 1869
Nigroplia Dombrow, 2002
Nigrotrichia Matsumoto, 2016
Nimbacola Paulian, 1970
Nimbus Mulsant & Rey, 1869
Niphetophora Kraatz, 1883
Niphobleta Kraatz, 1880
Nipponaphodius Nakane, 1959
Nipponoagoliinus Ochi & Kawahara, 2001
Nipponoserica Nomura, 1973
Nipponovalgus Sawada, 1941
Nitiocellus Branco, 2010
Nitorellus Britton, 1957
Niuailan Allsopp & Hutchinson, 2020
Nobiellus Dellacasa & Dellacasa, 2005
Nobius Mulsant & Rey, 1869
Nolicus Petrovitz, 1962
Nosphisthis Blackburn, 1898
Nothophanus Heller, 1896
Notocaulus Quedenfeldt, 1884
Novapus Sharp, 1875
Nselenius (Holm & Perissinotto, 2004)
Nuiba Prokofiev, 2016
Nunoidium Vaz-de-Mello, 2008
Nyassinus Westwood, 1879
Obaphodius Bordat, 1988
Oberthuroplia Lacroix, 1998
Ochicanthon Vaz-de-mello, 2003
Octocollis Moeseneder & Hutchinson, 2012
Octoplasia Brenske, 1892
Odochilus Harold, 1877
Odontacrossus Dellacasa, Král, Dellacasa & Bordat, 2014
Odontolochus Schmidt, 1917
Odontoloma Boheman, 1857
Odontolytes Koshantschikov, 1916
Odontoplia Fairmaire, 1897
Odontopsammodius Gordon & Pittino, 1992
Odontorrhina Burmeister, 1842
Odontotonyx MacLeay, 1871
Odontoxyomus Král, Rakovič & Mencl, 2015
Odontria White, 1846
Oedanomerus Waterhouse, 1875
Oedichira Burmeister, 1855
Oedipovalgus Kolbe, 1897
Oligophylla Kraatz, 1894
Omaloplia Schönherr, 1817
Omocnemus Schein, 1958
Omocrates Burmeister, 1844
Oncerus LeConte, 1856
Oncochirus Kolbe, 1891
Oniticellus Dejean, 1821
Onitis Fabricius, 1798
Onochaeta Erichson, 1847
Onoreidium Vaz-de-Mello, 2008
Onorius Frolov & Vaz-de-Mello, 2015
Ontherus Erichson, 1847
Onthobium Reiche, 1860
Onthophagus Latreille, 1802
Onychionyx Arrow, 1914
Onychoserica Moser, 1916
Onychosophrops Frey, 1972
Onychothecus Boucomont, 1912
Oocamenta Péringuey, 1904
Oogenius Solier, 1851
Ophropyx Britton, 1987
Oplognathus MacLeay, 1819
Oplostomus MacLeay, 1838
Opsitocus Britton, 1957
Orammoecius Král, Rakovič & Mencl, 2016
Oreoderus Burmeister, 1842
Oreogenoplia Lacroix, 1998
Oreotrogus Kolbe, 1897
Oreovalgus Kolbe, 1904
Orizabus Fairmaire, 1878
Orocanthus Endrödi, 1983
Oroclita Krikken, 1982
Orodaliscoides Schmidt, 1914
Orodaliscus Reitter, 1900
Oromus Mulsant & Rey, 1869
Orphnus MacLeay, 1819
Orrodoretus Ohaus, 1912
Orsilochus Burmeister, 1847
Orthocavonus Carne, 1957
Orthoserica Brenske, 1900
Orubesa Reitter, 1895
Oruscatus Bates, 1870
Oryctes Hellwig, 1798
† Oryctoantiquus Ratcliffe, Smith & Erwin, 2005
Oryctoderinus Endrödi, 1978
Oryctoderus Boisduval, 1835
Oryctomorphus Guérin-Méneville, 1830
Oryctophileurus Kolbe, 1910
Oscarinus Gordon & Skelley, 2007
Osmanius Branco & Baraud, 1988
Osmoderma Le Peletier & Serville, 1828
Otoclinius Brenske, 1896
Otophorus Mulsant, 1842
Ottokelleria D'Andretta & Martinez, 1957
Outenikwanus Scholtz & Howden, 1987
Outeniqua Péringuey, 1902
Ovomanonychus Costa, Cherman & Iannuzzi, 2020
Ovoserica Frey, 1968
Oxyadoretus Arrow, 1917
Oxyataenius Dellacasa & Stebnicka, 2001
Oxychirus Quedenfeldt, 1888
Oxycorythus Solsky, 1876
Oxyligyrus Arrow, 1908
Oxyomoides Dellacasa, Dellacasa & Gordon, 2016
Oxyomus Dejean, 1833
Oxypelta Pouillaude, 1920
Oxyserica Brenske, 1899
Oxysternon Castelnau, 1840
Oxythyrea Mulsant, 1842
Ozodius Skelley, 2007
Pachacama Soula, 2006
Pachnessa Brenske, 1894
Pachnoda Burmeister, 1842
Pachnodoides Alexis & Delpont, 2002
Pachrodema Blanchard, 1851
Pachycamenta Brenske, 1895
Pachychilecamenta Brenske, 1903
Pachycnema Le Peletier & Serville, 1828
Pachycolus Erichson, 1847
Pachydema Laporte, 1832
Pachydemocera Reitter, 1902
Pachyderoserica Moser, 1920
Pachygastra Germar, 1848
Pachylomera Griffith & Pidgeon, 1831
Pachyoryctes Arrow, 1908
Pachypoides Fairmaire, 1884
Pachypus Dejean, 1821
Pachyrhinadoretus Ohaus, 1912
Pachyserica Brenske, 1897
Pachysoma MacLeay, 1821
Pachystethus Blanchard, 1851
Pachytephraea De Palma & Malec, 2020
Pachytricha Hope, 1841
Pacuvia Curtis, 1845
Palacephala Lacroix, 1999
Palaeophileurus Kolbe, 1910
Paleira Reiche, 1871
Paleopragma Thomson, 1880
† Paleotrichius Poinar, 2011
Palnia Stebnicka, 1985
Panelus Lewis, 1895
Panglaphyra Kraatz, 1880
Panotrogus Reitter, 1902
Pantodinus Burmeister, 1847
Pantolia Burmeister, 1842
Papuana Arrow, 1911
Parabodilus Hollande & Thérond, 1998
Parabyrsopolis Ohaus, 1915
Paracamenta Péringuey, 1904
Paracanthon Balthasar, 1938
Paraceratochelus Dombrow, 2002
Parachilia Burmeister, 1842
Parachlorota Soula, 2002
Parachorius Harold, 1873
Parachrysina Bates, 1888
Paraclitopa Waterhouse, 1875
Paracoptochirus Balthasar, 1963
Paracorythoderus Wasmann, 1918
Paracotalpa Ohaus, 1915
Paracrossidius Balthasar, 1932
Paracryptocanthon Howden & Cook, 2002
Paracyclidius Howden, 1971
Paradeloparius Landin, 1959
Paradidactylia Balthasar, 1937
Paradoaphius Bordat, 2022
Paradorodocia Machatschke, 1957
Paradorysthetus Soula, 1998
Paraegidium Vulcano, Pereira & Martinez, 1966
Paragnorimus Becker, 1910
Paragymnopleurus Shipp, 1897
Paraheteronyx Moser, 1924
Paraheterosternus Morón, 1983
Paraholotrichia Lacroix & Coache, 2017
Paraixodina Roggero, Barbero & Palestrini, 2015
Paralleucosma Antoine, 1989
Parallidiostoma Ocampo & Colby, 2009
Paramacraspis Ohaus, 1915
Paramaladera Nomura, 1974
Paramicroplus Lacroix, 1998
Parammoecius Seidlitz, 1888
Paramorphochelus Lacroix, 1997
Paranelaphinis Antoine, 1988
Paranimbus Schmidt, 1914
Paranodon Cockerell, 1905
Paranomala Casey, 1915
Parapetiia Martinez, 1961
Paraphacosomoides Balthasar, 1969
Paraphileurus Endrödi, 1978
Paraphytus Harold, 1877
Parapilinurgus Arrow, 1910
Paraplectrone Mikšić, 1985
Paraplesiataenius Chalumeau, 1992
Parapoecilophila Hauser, 1904
Paraprotaetia Moser, 1907
Parapsammodius Verdu, Stebnicka & Galante, 2006
Paraptenomela Soula, 2002
Parapucaya Prell, 1934
Pararhabdotis Kraatz, 1899
Pararhopaea Blackburn, 1911
Pararhynchocephala Paulian, 1991
Pararhyssemus Balthasar, 1955
Parascatonomus Paulian, 1932
Paraschizognathus Ohaus, 1904
Parasciton Britton, 1990
Paraserica Reitter, 1896
Parastasia Westwood, 1842
Parastraella Antoine, 2005
Parasymmela Pacheco, Monné, Vaz-de-Mello & Ahrens, 2022
Parataenius Balthasar, 1961
Paratelaugis Ohaus, 1915
Parateuchus Shipp, 1895
Parathyridium Ohaus, 1915
Paratrichaphodius Bordat, 2013
Paratrichius Janson, 1881
Paratriodonta Baraud, 1962
Parautoserica Lacroix, Coache & Filippi, 2022
Paraxeloma Holm, 1988
Pardalosus Gordon & Skelley, 2007
Parelaphinis Marais & Holm, 1989
Parepixanthis Kraatz, 1893
Pareriesthis Moser, 1918
Pareteronychus Prell, 1934
Pareuchilia Kraatz, 1880
Parhomonyx Ohaus, 1915
Parhoplognathus Ohaus, 1915
Parisolea Bates, 1888
Parisomorphus Schaufuss, 1890
Paroniticellus Balthasar, 1963
Paronthobium Paulian, 1984
Paronyx Britton, 1990
Paroplia Fairmaire, 1903
Paroryctoderus Dechambre, 1994
Parthenoserica Brenske, 1899
Parvuhowdenius Deschodt & Scholtz, 2008
Pasaphylla Péringuey, 1904
Passaliolla Balthasar, 1945
Patatra Soula, 2008
Paulianellus Balthasar, 1938
Paulosawaya Martinez & D'andretta, 1956
Peckolus Scholtz & Howden, 1987
Pectinichelus Ballion, 1871
Pectinosoma Arrow, 1913
Pedaria Laporte, 1832
Pedaridium Harold, 1868
Pedinorrhina Kraatz, 1880
Pedinotrichia Matsumoto, 2016
Pegylis Erichson, 1847
Pelidnota MacLeay, 1819
† Pelidnotites Cockerell, 1920
Peltonotus Burmeister, 1847
Penalus Paulian, 1985
Pentacoryna Moser, 1926
Pentanomala Ohaus, 1919
Pentaphylla De Wailly, 1950
Pentecamenta Brenske, 1896
Pentelia Brenske, 1891
Penthima Kraatz, 1892
Pentodina Endrödi, 1968
Pentodon Hope, 1837
Pentodontoschema Péringuey, 1901
Peotoxus Krikken, 1983
Peperonota Westwood, 1847
Pereiraidium Vaz-de-Mello, 2008
Pericamenta Péringuey, 1904
Periclitopa Brenske, 1896
Pericoptus Burmeister, 1847
Periphanesthes Kraatz, 1880
Periproctoides Lacroix, 2006
Periproctus Kolbe, 1910
Periserica Brenske, 1898
Perissosoma Waterhouse, 1875
Peritrichia Burmeister, 1844
Perrindema Lacroix, 1997
Peruquime Mondaca & Valencia, 2016
Petinopus Blackburn, 1898
Petrovitzia Mikšić, 1965
Petrovitzius Rakovič, 1979
Phaeadoretus Reitter, 1903
Phaeaphodius Reitter, 1892
Phaedimus Waterhouse, 1841
Phaedotrogus Paulian, 1985
Phaenognatha Hope, 1841
Phaenomeris Hope, 1833
Phalacronothus Motschulsky, 1859
Phalangogonia Burmeister, 1844
Phalangonyx Reitter, 1889
Phalangosoma Quedenfeldt, 1884
Phalops Erichson, 1848
Phanaeus MacLeay, 1819
Phaneresthes Kraatz, 1894
Pharaonus Blanchard, 1851
Pharaphodius Reitter, 1892
Phiara Brenske, 1897
Philacelota Heller, 1900
Phileucourtus Dechambre, 2008
Phileurus Latreille, 1807
Philgertia Zídek, 2021
Philistina MacLeay, 1838
Philochloenia Dejean, 1833
Philoscaptus Brèthes, 1919
Philoserica Brenske, 1897
Phobetus LeConte, 1856
Pholeoaphodius Bordat, 2009
Pholidochris Kolbe, 1894
Phonopleurus Moser, 1919
Phonotaenia Kraatz, 1880
Phorine Britton, 1987
Photyna Brenske, 1897
Phoxomela Schaum, 1844
Phoxomeloides Schoch, 1898
Phycochus Broun, 1886
Phylliocephala Blackburn, 1890
Phyllocamenta Moser, 1918
Phyllochlaenia Blanchard, 1846
Phyllococerus Waterhouse, 1876
Phyllognathus Eschscholtz, 1830
Phyllopertha Stephens, 1830
Phyllophaga Harris, 1826
Phylloserica Brenske, 1899
Phyllotocidium Blackburn, 1898
Phyllotocus Fischer Von Waldheim, 1823
Phyllotrochalus Brenske, 1900
Phytholaema Blanchard, 1851
Pichica Soula, 2002
Pileotrichius Bourgoin, 1921
Pilinopyga Kraatz, 1888
Pilinurgus Burmeister, 1842
Pimelomera Moser, 1924
Pimelopus Erichson, 1842
Pinacopodius Branco, 1991
Pinacotarsus Harold, 1875
Piscoperus Ratcliffe & Giraldo, 2014
Pittinius Rakovič & Král, 1997
Placodidus Péringuey, 1900
Plaesioserica Brenske, 1899
Plagiogonus Mulsant, 1842
Planolinellus Dellacasa & Dellacasa, 2005
Planolinoides Dellacasa & Dellacasa, 2005
Planolinus Mulsant & Rey, 1869
Planophileurus Chapin, 1932
Platychelus Burmeister, 1844
Platycoelia Dejean, 1833
Platyderides Schmidt, 1917
Platygenia MacLeay, 1819
Platygeniops Krikken, 1978
Platynocephalus Westwood, 1854
Platyonitis Janssens, 1942
Platyphileurus Ohaus, 1910
Platyrutela Bates, 1888
Platysodes Westwood, 1874
Platytomus Mulsant, 1842
Plectris Le Peletier & Serville, 1828
Plectrodes Horn, 1867
Plectrone Wallace, 1867
Pleistophylla Péringuey, 1904
Pleophylla Erichson, 1847
Plesiocamenta Lacroix & Montreuil, 2019
Plesiopalacephala Lacroix, 2006
Plesiorutela Jameson, 1997
Plesiosternus Morón, 1983
Pleuraphodius Schmidt, 1914
Pleuronitis Lansberge, 1875
Pleuronota Kraatz, 1892
Pleurophorus Mulsant, 1842
Plochilia Fairmaire, 1896
Plochiliana Ruter, 1978
Plotopuserica Brenske, 1899
Plusioserica Brenske, 1899
Podalgus Burmeister, 1847
Podischnus Burmeister, 1847
Podolasia Harold, 1869
Podopholis Moser, 1915
Podopogonus Moser, 1917
Podostena Howden, 1997
Podotenoides Paulian, 1942
Podotenus Schmidt, 1914
Podovalgus Arrow, 1910
Poecilopharis Kraatz, 1880
Pogoniotarsus Kraatz, 1880
Pogonopus Arrow, 1910
Pogonotarsus Burmeister, 1842
Pollaplonyx Waterhouse, 1875
Polyenaria De Wailly, 1950
Polyphylla Harris, 1841
Polyphyllum Blanchard, 1851
Polyplastus Janson, 1888
Polystalactica Kraatz, 1882
Popillia Dejean, 1821
Porphyronota Burmeister, 1842
Prionoryctes Arrow, 1911
Priscorrhina Krikken, 1984
† Priscyparus Skelley, 2021
Priska Jákl, 2018
Pristerophora Harold, 1869
Proagoderus Lansberge, 1883
Proagopertha Reitter, 1903
Proagosternus Blanchard, 1851
Problerhinus Deyrolle, 1864
Proborhinus Britton, 1988
Prochelyna Erichson, 1847
Proctophanes Harold, 1861
Proculigyrus López-García & Deloya, 2022
Prodontria Broun, 1904
Prodoretus Brenske, 1893
Promacropoides Sigwalt, 1987
† Prophaenognatha Bai, Ren & Yang, 2011
Propomacrus Newman, 1836
Prosphileurus Kolbe, 1905
Protaetia Burmeister, 1842
Protaetiomorpha Mikšić, 1968
Protelura Britton, 1987
Proteroschiza Brenske, 1896
Protochilus Krikken, 1976
Protoclita Krikken, 1978
Protoclitopa Decelle, 1979
† Protopalnia Nikolajev, 2007
† Prototrox Nikolajev, 2000
† Psammaegialia Nikolajev, Wang & Zhang, 2014
Psammodaphodius Endrödi, 1976
Psammodius Fallen, 1807
Psammoporus Thomson, 1859
Psammorpha Stebnicka, 1994
Psednoserica Brenske, 1899
Pseudachloa Péringuey, 1904
Pseudacrossus Reitter, 1892
Pseudadoretus Semenov, 1890
Pseudagenius Heller, 1923
Pseudagolius Schmidt, 1914
Pseudalleucosma Antoine, 1989
Pseudastoxenus Bourgoin, 1921
Pseudataenius Brown, 1927
Pseudenaria Fairmaire, 1901
Pseudencya Lacroix, 1991
Pseudepixanthis Kraatz, 1880
Pseuderytus Hollande & Thérond, 1998
Pseudesymus Orbigny, 1896
Pseudheterochelus Schein, 1959
Pseudholophylla Blackburn, 1911
Pseudignambia Paulian, 1984
Pseudinca Kraatz, 1880
Pseudisonychus Frey, 1971
Pseudoagoliinus Bordat, 1994
Pseudoahermodontus Dellacasa & Dellacasa, 1997
Pseudoanticheiroides Soula, 1998
Pseudoblackburneus Bordat, 2016
Pseudoblitopertha Keith, 2000
Pseudocanthon Bates, 1887
Pseudocavonus Blackburn, 1890
Pseudochalcothea Ritsema, 1882
Pseudochalcotheomima Mikšić, 1985
Pseudochironitis Ferreira, 1977
Pseudochlorota Ohaus, 1905
Pseudoclinteria Kraatz, 1882
Pseudoclithria Neervoort Van De Poll, 1886
Pseudocoelotrachelus Dellacasa, Dellacasa & Gordon, 2013
Pseudocongella Moser, 1924
Pseudocopris Ferreira, 1960
Pseudocotalpa Hardy, 1971
Pseudocyrtolophus Frolov & Montreuil, 2019
Pseudodiceros Mikšić, 1974
Pseudodicrania Gutiérrez, 1950
Pseudodiplotaxis Nonfried, 1894
Pseudodorysthetus Soula, 1998
Pseudodrepanocanthus (Bordat, 1993)
Pseudogeniates Ohaus, 1910
Pseudogonaphodiellus Dellacasa, Gordon & Dellacasa, 2007
Pseudoheptaulacus Dellacasa & Dellacasa, 2000
Pseudoheteronyx Blackburn, 1892
Pseudohomonyx Arrow, 1908
Pseudohypaspidius Soula, 1998
Pseudoliogenys Moser, 1919
Pseudomacraspis Ohaus, 1903
Pseudomaladera Nomura, 1974
Pseudomesystoechus Ohaus, 1912
Pseudomicroplus Lacroix, 1998
Pseudomothon Pittino, 1984
Pseudomyrhessus Minkina, 2023
Pseudonthobium Paulian, 1984
Pseudooxythyrea Baraud, 1985
Pseudopachydema Balthasar, 1930
Pseudopanotrogus Petrovitz, 1969
Pseudoparoplia Lacroix, 1997
Pseudopectinosoma Katovich, 2011
Pseudopedaria Felsche, 1904
Pseudoperiproctus Keith & Reichenbach, 2018
Pseudopharaphodius Bordat, 1990
Pseudopholis Duvivier, 1892
Pseudopilinurgus Moser, 1918
Pseudopodotenus Dellacasa, 1990
Pseudoptenomela Soula, 2002
Pseudorphnus Benderitter, 1913
Pseudoryctes Sharp, 1873
Pseudosaproecius Balthasar, 1941
Pseudoscaptobius Krikken, 1976
Pseudoschizognathus Ohaus, 1904
Pseudoschizonycha Keith, 2006
Pseudoserica Guérin-Méneville, 1830
Pseudosericania Kobayashi, 1980
Pseudostegopterus Ricchiardi, 2015
Pseudostereomera Bordat & Howden, 1995
Pseudosymmachia Dalle Torre, 1912
Pseudosyrichthus Péringuey, 1901
Pseudotephraea Kraatz, 1882
Pseudotetraclipeoides Dellacasa, Dellacasa & Gordon, 2015
Pseudoteuchestes Dellacasa, 1986
Pseudothyridium Soula, 2002
Pseudotocama Keith, 2008
Pseudotorynorrhina Mikšić, 1967
Pseudotrematodes Jacquelin Du Val, 1859
Pseudotrigonocnemis Keith, 2007
Pseudotrochalus Quedenfeldt, 1884
Pseudouteniqua Schein, 1959
Pseudoxyomus Petrovitz, 1962
Pseustophylla Péringuey, 1904
Psilocnemis Burmeister, 1842
Psilodontria Broun, 1895
Psilonychus Burmeister, 1855
Psilopholis Brenske, 1892
Ptenomela Bates, 1888
Pterobius Murayama, 2010
Ptychodesthes Kraatz, 1883
Ptyophis Redtenbacher, 1868
Pubinus Mulsant & Rey, 1869
Pucaya Ohaus, 1910
Puelchesia Ocampo & Smith, 2006
Pukupuku Muramoto, 2006
Pusiodactylus Smith, 2008
Pycnopanelus Arrow, 1931
Pycnoschema Thomson, 1859
Pygora Burmeister, 1842
Pygovalgus Kolbe, 1884
Pyronota Boisduval, 1835
Pyrrhopoda Kraatz, 1880
Qingaphodius Král, 1997
Rabula Péringuey, 1902
Raceloma Thomson, 1877
Rakovicius Pittino, 2006
Ramilia Kolbe, 1894
Ravautiana Lacroix, 1991
Raysymmela Saylor, 1947
Rectoscutaria Schein, 1958
Redotus Fairmaire, 1860
Reineria Mikšić, 1968
Renaudiana Lacroix, 1993
Renaudius Balthasar, 1951
Renorphnus Frolov & Montreuil, 2009
Repsimus MacLeay, 1819
Rhabdopholis Burmeister, 1855
Rhabdotis Burmeister, 1842
Rhabdotops Krikken, 1981
Rhachistoplia Lacroix, 1997
Rhadinolontha Arrow, 1930
Rhadinotaenia Kraatz, 1900
Rhagopteryx Burmeister, 1842
Rhamphadoretus Ohaus, 1912
Rhamphorrhina Klug, 1855
Rhinarion Ruter, 1965
Rhinaspis Perty, 1833
Rhinocerotopsis Maruyama, 2009
Rhinocoeta Burmeister, 1842
Rhinyptia Burmeister, 1844
Rhizogeniates Ohaus, 1909
Rhizoplatodes Péringuey, 1901
Rhizoplatys Westwood, 1841
Rhizoproctus Kolbe, 1894
Rhizotrogus Latreille, 1825
Rhodaphodius Adam, 1994
Rhomborhina Hope, 1837
Rhopaea Erichson, 1847
Rhynchadoretus Ohaus, 1913
Rhynchapogonia Lacroix, 2008
Rhynchocephala Fairmaire, 1883
Rhynchoserica Burgeon, 1942
Rhyparus Agassiz, 1846
Rhysothorax Bedel, 1911
Rhyssemodes Reitter, 1892
Rhyssemorphus Clouët, 1900
Rhyssemus Mulsant, 1842
Rhyxicephalus Moser, 1921
Rhyxiderma Moser, 1924
Rhyxiphloea Burmeister, 1842
Rigoutorum Hutchinson & Moeseneder, 2022
Rimuloplia Lacroix, 1997
Robinsonelliana Lacroix, 1996
Rubilepis De Wailly, 1950
Rufotrichia Matsumoto, 2016
Rugaphodius Gordon & Skelley, 2007
Rugopertha Machatschke, 1957
Rutela Latreille, 1802
Rutelarcha Waterhouse, 1874
Rutelisca Bates, 1888
Ruteloryctes Arrow, 1908
Ruteraetia Krikken, 1980
Sabatinellius Montreuil & Lacroix, 2020
Sabatinelloplia Lacroix, 1998
Saccharoscaptus Morón & Grossi, 2015
Sahlbergianus Dellacasa & Dellacasa, 2000
Saphobiamorpha Brookes, 1944
Saphobius Sharp, 1873
Saprolochus Stebnicka & Galante, 2007
Saprositellus Balthasar, 1967
Saprosites Redtenbacher, 1857
Saprovisca Stebnicka, 1993
Saprus Blackburn, 1904
Sariangus Rakovič & Mencl, 2011
Sarophorus Erichson, 1847
Sarothromerus Blackburn, 1907
Saulostomus Waterhouse, 1878
Sauvagesinella Paulian, 1934
Sayloria Frey, 1975
Scabrostomus Gordon & Skelley, 2007
Scapanes Burmeister, 1847
Scapanoclypeus Evans, 1987
Scaphorhina Quedenfeldt, 1884
Scaphorhinadoretus Ohaus, 1912
Scaptobius Schaum, 1841
Scaptocnemis Péringuey, 1901
Scarabaeolus Balthasar, 1965
Scarabaeus Linnaeus, 1758
Scatimus Erichson, 1847
Scatonomus Erichson, 1835
Scatrichus Génier & Kohlmann, 2003
Sceliages Westwood, 1837
Scelidothrix Dombrow, 2001
Scelophysa Burmeister, 1844
Schaefferellus Gordon & Skelley, 2007
Scheinia Ruter, 1957
Schismatocera Gautier Des Cottes, 1872
Schizadoretus Arrow, 1917
Schizochelus Blanchard, 1850
Schizognathus Fischer Von Waldheim, 1823
Schizonycha Dejean, 1833
Schizorhina Kirby, 1825
Schochidia Berg, 1898
Scitala Erichson, 1842
Scitaloides Britton, 1990
Sciton Blackburn, 1892
Scybalocanthon Martinez, 1948
Scybalophagus Martinez, 1953
Scythrodes Broun, 1886
Scythropesthes Kraatz, 1880
Sebakwe Péringuey, 1904
Sebaris Castelnau, 1840
Selaserica Brenske, 1897
Selomothus Fairmaire, 1891
Selviria Stebnicka, 1999
Semanopterus Hope, 1847
Semienaria De Wailly, 1950
Serica MacLeay, 1819
Sericania Motschulsky, 1860
Sericesthis Boisduval, 1835
Sericoides Guérin-Méneville, 1839
Sericospilus Sharp, 1882
Serraphodius Kabakov, 1996
Setiserica Miyake, 2002
Setodius Gordon & Skelley, 2007
Setylaides Stebnicka, 1994
Siamaphodius Masumoto, 1991
Siamophylla Keith, 2005
Sicardia Reitter, 1896
Sigorus Mulsant & Rey, 1869
Silluvia Landin, 1949
Silvaphilus Roets & Oberlander, 2010
Silvinha Vaz-de-Mello, 2008
Simogonius Harold, 1871
Simorrhina Kraatz, 1886
Sinaphodius Červenka, 1994
Sinapisoma Boucomont, 1928
Singhala Blanchard, 1851
Sinodiapterna Dellacasa, 1986
Sinodrepanus Simonis, 1985
Siralus Dechambre, 1998
Sisyphus Latreille, 1807
Sitiphus Fairmaire, 1894
Skelleyanus Dellacasa, Dellacasa, Gordon & Stebnicka, 2011
Smicorhina Westwood, 1847
Socotraproctus Král, Sehnal & Bezděk, 2012
Somalibia Lansberge, 1882
Somatoserica Brenske, 1899
Sophrops Fairmaire, 1887
Sorocha Soula, 2006
Spaniolepis Kolbe, 1894
Sparrmannia Castelnau, 1840
Spathoschiza Arrow, 1902
Spelaiorrhina Lansberge, 1886
Sphaeraphodius Kakizoe, Jiang & Wang, 2021
Sphaerorutela Jameson, 1997
Sphaeroscelis Burmeister, 1855
Sphaerotrochalus Brenske, 1900
Sphecoserica Brenske, 1899
Sphinctovalgus Kolbe, 1904
Sphodroxia Kraatz, 1890
Sphyrocallus Sharp, 1877
Spilophorus Westwood, 1848
Spilopopillia Kraatz, 1892
Spinanomala Ohaus, 1910
Spinochlamys Machatschke, 1970
Spodistes Burmeister, 1847
Spodochlamys Burmeister, 1855
Stalagmosoma Burmeister, 1842
Stebnickiella Skelley, 2007
Stegopterus Burmeister & Schaum, 1840
Stenochelyne Britton, 1990
Stenocrates Burmeister, 1847
Stenonota Fairmaire, 1889
Stenopegylis Arrow, 1943
Stenopisthes Moser, 1913
Stenoserica Brenske, 1900
Stenosophrops Nomura, 1977
Stenosternus Karsch, 1881
Stenotarsia Burmeister, 1842
Stenotothorax Schmidt, 1914
Stephanopholis Brenske, 1896
Stephanorrhina Burmeister, 1842
Stereomera Arrow, 1905
Stethaspis Hope, 1837
Stethodesma Bainbridge, 1840
Stethopseudinca Valck Lucassen, 1933
Stigmatoplia Dombrow, 2001
Stilbotrochalus Kolbe, 1914
Stiptocnemis Branco, 1989
Stiptopodius Harold, 1871
Stiptotarsus Branco, 1989
Stomanomala Kolbe, 1897
Storeyus Hasenpusch & Moeseneder, 2010
Straliga Fairmaire, 1901
Strategus Hope, 1837
Streblopus Lansberge, 1874
Strigoderma Burmeister, 1844
Strigodius Gordon & Skelley, 2007
Stripsipher Gory & Percheron, 1833
Strumadoretus Ohaus, 1912
Subrinus Mulsant & Rey, 1869
Sugrames Reitter, 1894
Sukelus Branco, 1992
Sulcophanaeus Olsoufieff, 1924
Suntemnonycha Péringuey, 1904
Supertermitoderus Mencl, 2011
Surutu Martinez, 1955
Sussorca Paulian, 1954
Sybacodes Fairmaire, 1896
Sybax Boheman, 1857
Sylvicanthon Halffter & Martinez, 1977
Symmela Erichson, 1835
Symphodon Schmidt, 1914
Synacta Fairmaire, 1899
Synapsis Bates, 1868
Synchilus Britton, 1956
Synclitopa Kolbe, 1897
Synenaria Lacroix, 1993
Syngeneschiza Brenske, 1898
Synistovalgus Kolbe, 1897
Syrichthodontus Péringuey, 1901
Syrichthomorphus Péringuey, 1901
Syrichthoschema Janssens, 1942
Systellopus Sharp, 1877
Systellorrhina Kraatz, 1895
Taeniodera Burmeister, 1842
Tafaia Valck Lucassens, 1939
Taiwanotrichia Kobayashi, 1990
Talautoclyster Yamaya, 2001
Tamnoserica Brenske, 1899
Tanyana Stebnicka, 2006
Tanyproctoides Petrovitz, 1971
Tanyproctus Ménétriés, 1832
Tanzanilepis Lacroix, 2008
Tanzanipholis Lacroix, 2002
Tanzanolus Scholtz & Howden, 1987
Taphrocephala Quedenfeldt, 1888
Tapinoschema Thomson, 1880
Tarsocamenta Moser, 1924
Tarsovalgus Miyake, 1993
Taurhina Burmeister, 1842
Tehuacania Endrödi, 1975
Teinogenys Sharp, 1873
Telaugis Burmeister, 1844
Telochilus Krikken, 1975
Telura Erichson, 1842
Teluroides Britton, 1990
Temnoplectron Westwood, 1841
Temnorhynchus Hope, 1837
Tephraea Burmeister, 1842
Tephraeoserica Brenske, 1900
Teraserica Brenske, 1897
Terebrogaster Lacroix, 1989
Termitaxis Krikken, 1970
Termitoderus Mateu, 1966
Termitodiellus Nakane, 1961
Termitodius Wasmann, 1894
Termitophilus Britton, 1957
Termitopisthes Wasmann, 1918
Termitotrox Reichensperger, 1915
Territonia Krikken, 2018
Tesarius Rakovič, 1981
Tesserodon Hope, 1837
Tesserodoniella Vaz-de-Mello & Halffter, 2006
Testaceitrichia Matsumoto, 2021
Tetraclipeoides Schmidt, 1914
Tetraechma Blanchard, 1845
Tetragonorrhina Kraatz, 1896
Tetragus Duarte & Grossi, 2022
Tetraodorhina Blanchard, 1842
Tetraserica Ahrens, 2004
Teuchestes Mulsant, 1842
Thabina Péringuey, 1902
Theodosia Thomson, 1880
Theotimius Huchet, 2000
Theuremaripa Soula, 2006
Thinorycter Semenov & Reichardt, 1925
Thoracoplia Prokofiev, 2015
Thoracotrichia Decelle, 1968
Thronistes Burmeister, 1847
Thrymoserica Brenske, 1897
Thyce LeConte, 1856
Thyregis Blackburn, 1904
Thyreogonia Reitter, 1898
Thyridium Burmeister, 1844
Thyriochlorota Ohaus, 1915
Tiamidema Prokofiev, 2014
Tiarocera Burmeister, 1842
Tiaronthophagus Roggero, Moretto, Barbero & Palestrini, 2019
Tibiodrepanus Krikken, 2009
Tibiotrichius Miyake, 1994
Tibiovalgus Krikken, 1978
Tiniocellus Péringuey, 1901
Tipicha Soula, 2002
Tlaocera Péringuey, 1904
Tmesorrhina Westwood, 1841
Tocama Reitter, 1902
Tomarus Erichson, 1847
Tomogonus Orbigny, 1904
Tonkinilla Prokofiev, 2021
Torynorrhina Arrow, 1907
Tosevskiana Pavicevic, 1985
Toxocnemis Dombrow, 2002
Toxospathius Fairmaire, 1878
Trachyserica Brenske, 1899
Tragiscus Klug, 1855
Trematodes Faldermann, 1835
Triacmoserica Kolbe, 1914
Tribialus Dupuis & Perrin, 2022
Tribopertha Reitter, 1903
Trichanomala Arrow, 1917
Trichaphodiellus Schmidt, 1914
Trichaphodioides Paulian, 1942
Trichaphodius Schmidt, 1914
Trichaulax Kraatz, 1880
Trichesthes Erichson, 1847
Trichillidium Vaz-de-Mello, 2008
Trichillum Harold, 1868
Trichinopus Waterhouse, 1875
Trichioaparammoecius Minkina, 2018
Trichiodera Burmeister, 1855
Trichiopsammobius Petrovitz, 1963
Trichiorhyssemus Clouët, 1901
Trichiotinus Casey, 1915
Trichius Fabricius, 1775
Trichocephala Moser, 1916
Trichogomphus Burmeister, 1847
Tricholepis Blanchard, 1851
Tricholontha Nomura, 1952
Trichomaladera Nomura, 1974
Trichoneptunides Legrand, 2001
Trichonotuloides Balthasar, 1945
Trichonotulus Bedel, 1911
Trichoparadidactylia Bordat, 2022
Trichoplus Burmeister, 1842
Trichopopillia Ohaus, 1897
Trichoschiza Moser, 1917
Trichostetha Burmeister, 1842
Trigonochilus Brenske, 1896
Trigonocnemis Kraatz, 1894
Trigonopeltastes Burmeister & Schaum, 1840
Trigonophorinus Pouillaude, 1913
Trigonophorus Hope, 1831
Trigonoscelus (Petrovitz, 1963)
Trigonostomus Brenske, 1893
Trinoxia Brenske, 1894
Triodontella Reitter, 1919
Triodontonyx Moser, 1921
Triodontus Westwood, 1845
Triodonyx Saylor, 1942
Trioplognathus Ohaus, 1904
Trioplus Burmeister, 1847
Trioserica Moser, 1922
Triplognatha Krikken, 1987
Trissodon Burmeister, 1847
Tristaphodius Balthasar, 1932
Trizogeniates Ohaus, 1917
Trochaloschema Reitter, 1896
Trochaloserica Brenske, 1900
Trochalus Laporte, 1832
Trogodes Boheman, 1857
Tropidonitis Janssens, 1937
Tropinota Mulsant, 1842
Tropiorhynchus Blanchard, 1851
Trymodera Gerstaecker, 1867
Trypoxylus Minck, 1920
Tsaratanoplia Lacroix, 1997
Tulbaghia Péringuey, 1904
Turanella Semenov, 1905
Uenoeus Ochi & Kon, 2017
Ulata Saylor, 1945
Uloptera Burmeister, 1842
Unidentis Josso & Prévost, 2008
Upsa Deschodt, Sole & Scholtz, 2020
Uroxys Westwood, 1842
Vadonaria De Wailly, 1950
Vadonidella Ruter, 1973
Valgoides Fairmaire, 1899
Valgus Scriba, 1790
Vansoniola Schein, 1958
Varencya Lacroix, 1993
Vayana Ohaus, 1915
Versicorpus Deschodt, Davis & Scholtz, 2011
Vezoanus Lacroix, 1993
Vieuella Ruter, 1964
Viridimicus Jameson, 1990
Vladimirellus Dellacasa, Dellacasa & Bordat, 2002
Volinus Mulsant & Rey, 1869
Wadaia Itoh, 1993
Walsternoplus Allsopp, Jákl & Rey, 2023
Walterantus Cambefort, 1977
Wambo Allsopp, 1988
Warwickia Smith & Evans, 2005
Watkinsia Britton, 1995
Webbella Britton, 1988
Wernoryctes Dechambre, 1999
Xanthotrogus Reitter, 1902
Xeloma Kraatz, 1881
Xenaclopus Arrow, 1915
Xenocanthon Martinez, 1952
Xenoceraspis Arrow, 1920
Xenochlorota Soula, 2002
Xenodorus Brême, 1844
Xenogeniates Villatoro & Jameson, 2001
Xenoheptaulacus Hinton, 1934
Xenoloba Bates, 1889
Xenopelidnota Bates, 1904
Xenoproctis Kolbe, 1896
Xenoreoderus Arrow, 1910
Xenoserica Ahrens, 2005
Xenotrochalus Moser, 1917
Xenotrogus Britton, 1978
Xeropsamobeus Saylor, 1937
Xinidium Harold, 1869
Xiphoscelis Burmeister, 1842
Xiphosceloides Holm, 1992
Xylonichus Boisduval, 1835
Xyloryctes Hope, 1837
Xyloscaptes Prell, 1934
Xylotrupes Hope, 1837
Xyridea Britton, 1990
Xyrine Britton, 1987
Xyroa Britton, 1987
Xyrodes Britton, 1987
Yaaxkumukia Morón & Nogueira, 1998
Yanovalgus Nomura, 1952
† Yixianscarabaeus Nikolajev, 2015
Youngaphodius Bordat, 1988
Yvescambefortius Ochi & Kon, 1996
Zaburina Saylor, 1945
Zagurya Fuhrmann, 2021
Zebinus Fairmaire, 1894
Zietzia Blackburn, 1894
Zimbabuephylla Lacroix, 1997
Zomba Lacroix, 2006
Zonocopris Arrow, 1932
Collection d'amulettes en forme de scarabées.
Le Scarabée sacré (Scarabaeus sacer ) est un animal vénéré dans l'Égypte antique . Symbole du soleil et de Rê , il était l'un des douze animaux sacrés associé aux douze heures du jour et de la nuit[ réf. nécessaire] .
Le Scarabée rhinocéros siamois ou scarabée de combat (Xylotrupes gideon ) est capturé dans la nature puis dressé pour des duels qui font l'objet de paris dans le nord de la Thaïlande . Ces duels de scarabées sont couramment organisés. Ces combats d’insectes du type xylotrupes appelés Kwang localement sont à bien des égards surprenants, notamment du fait qu’il est sans doute plus difficile d’attribuer à un scarabée qu’à un autre animal des caractéristiques anthropomorphiques et qu'il est plus compliqué encore de deviner ce qu’il perçoit. Le duel de scarabées met à l’épreuve une forme de zoomanité trouble fondée sur une forme d’attachement inédite entre l’homme et l’animal[7] , [8] .
Michael Chinery , Insectes de France et d'Europe occidentale, Flammarion, 2012 (1re éd. 1986), p. 264
Anatole Bailly ; 2020 : Hugo Chávez, Gérard Gréco, André Charbonnet, Mark De Wilde, Bernard Maréchal & contributeurs, « Le Bailly », 2020 (consulté le 24 octobre 2023 ) .
Latreille, 1802: Histoire naturelle, générale et particulière des Crustacés et des Insectes. Tome 3 . F. Dufart, Paris. Internet Archive BHL
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