Bembidion genus of beetles in the family Carabidae by number of species.[2] mm) and move very fast. Most of them live close to water.[2] biantitropical distribution , meaning they are found in both the Northern and Southern Hemispheres, but not in the tropics.[3] Tachys [2]
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Bembidion tetracolum
There have been many attempts to divide it into smaller genera, most notably by René Jeannel in 1941 and by G.G. Perrault in 1981, but none of them have been generally accepted.[2]
This genus is divided into numerus subgenera , some of which are elevated to full genus rank by various authors; as noted above, however, no universally accepted way of splitting the genus exists yet. Bembidion subgenera include:[4]
Actedium Motschulsky, 1864 Ananotaphus Netolitzky, 1931 Andrewesa Netolitzky, 1931 Antiperyphanes Jeannel, 1962 Antiperyphus Jeannel, 1962 Aptenidium Habu & Ueno, 1955 Apteromimus Wollaston, 1877 Armatocillenus Dupuis, 1912 Asioperyphus Vysoky, 1986 Aureoplataphus Netolitzky, 1943 Australoemphanes Toledano, 2005 Bembidion Latreille, 1802 Bembidionetolitzkya E.Strand, 1929 Bembidromus Toledano, 2000 Blepharoplataphus Netolitzky, 1920 Bracteon Bedel, 1879 Chilioperyphus Jeannel, 1962 Chinocillenus Netolitzky, 1942 Chlorodium Motschulsky, 1864 Cillenus Samouelle, 1819 Cyclolopha Casey, 1918 Desarmatocillenus Netolitzky, 1939 Diplocampa Bedel, 1896 Ecuadion Moret & Toledano, 2002 Emphanes Motschulsky, 1850 Endosomatium Wollaston, 1877 Euperyphus Jeannel, 1941 Eupetedromus Netolitzky, 1911 Furcacampa Netolitzky, 1931 Gnatholymnaeum Sharp, 1903 Gondwanabembidion Toledano, 2005 Hirmoplataphus Lindroth, 1963 Hoquedela Müller-Motzfeld, 1988 Hydriomicrus Casey, 1918 Hydrium LeConte, 1847 Hypsipezum Alluaud, 1917 Jammuphanes J.Schmidt & Marggi, 2014 Jedlickion Toledano, 2008 Josefia Toledano, 2000 Leuchydrium Casey, 1918 Limnaeoperyphus Nakane, 1963 Lindrochthus Maddison, 2012 Liocosmius Casey, 1918 Lymnaeum Stephens, 1828 Melomalus Casey, 1918 Metallina Motschulsky, 1850 Microserrullula Netolitzky, 1921 Microsinocys Toledano, 1998 Necpericompsus Netolitzky, 1935 Neja Motschulsky, 1864 Neobembidion Bousquet, 2006 Neoemphanes Habu, 1978 Nepha Motschulsky, 1864 Nesocidium Sharp, 1903 Nipponobembidion Habu & Baba, 1968 Notaphemphanes Netolitzky, 1920 Notaphocampa Netolitzky, 1914 Notaphus Dejean, 1821 Nothocys Jeannel, 1962 Notholopha Jeannel, 1962 Nothonepha Jeannel, 1962 Notoperyphus Bonniard de Saludo, 1970 Ochthedromus LeConte, 1847 Ocydromus Clairville, 1806 Ocyturanes Müller-Motzfeld, 1986 Odontium LeConte, 1847 Omoperyphus Netolitzky, 1931 Omotaphus Netolitzky, 1914 Pacmophena Jeannel, 1962 Pamirium Netolitzky, 1920 Paraprincidium Netolitzky, 1914 Pekinium Csiki, 1901 Peryphanes Jeannel, 1941 Peryphidium Tschitscherine, 1895 Peryphiolus Jeannel, 1941 Peryphodes Casey, 1918 Peryphophila Netolitzky, 1939 Peryphus Dejean, 1821 Philochthemphanes Netolitzky, 1943 Philochthus Stephens, 1828 Phyla Motschulsky, 1844 Plataphus Motschulsky, 1864 Plocamoperyphus Jeannel, 1962 Politophanes Müller-Motzfeld, 1998 Princidium Motschulsky, 1864 Pseudometallina Netolitzky, 1920 Pseudoperyphus Hatch, 1950 Pseudophilochthus Wollaston, 1877 Pseudosinocys Toledano, 2005 Pseudotrepanes Jeannel, 1962 Sakagutia Ueno, 1954 Semicampa Netolitzky, 1910 Sloanephila Netolitzky, 1931 Taiwanobembidion Habu, 1973 Talanes Motschulsky, 1864 Terminophanes Müller-Motzfeld, 1998 Testediolum Ganglbauer, 1891 Testedium Motschulsky, 1864 Thaumatoperyphus Netolitzky, 1935 Trechonepha Casey, 1918 Trepanedoris Netolitzky, 1918 Trepanes Motschulsky, 1864 Trichoplataphus Netolitzky, 1914 Zeactedium Netolitzky, 1931 Zecillenus Lindroth, 1980 Zemetallina Lindroth, 1976 Zeperyphodes Lindroth, 1976 Zeperyphus Lindroth, 1976 Zeplataphus Lindroth, 1976 † Archaeophilochthus Ortuño & Arillo, 2010
† Eodontium J.Schmidt & Michalik, 2017
Hist. Nat. Crust. Ins. 3: 82. Type species : Carabus quadrimaculatus Linnaeus , 1761 (= Bembidion quadrimaculatum ).Carl H. Lindroth. The Carabidae (Coleoptera) of Fennoscandia and Denmark . Leiden - Copenhagen: Brill - Scandinavian Science Press, 1985. ISBN 90-04-07727-8 . P. 129-199. Philip Jackson Darlington. Biogeography of the Southern End of the World . Cambridge: Harvard University Press, 1965. P. 22, 45.
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